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<document id="E506295E671E83FEBE8C86BBA31899DB" ID-CLB-Dataset="26368" ID-DOI="10.11646/zootaxa.4624.2.10" ID-GBIF-Dataset="7fc3c435-d3ed-45db-bd7d-48398d831f01" ID-ISSN="1175-5326" ID-Zenodo-Dep="3259222" ID-ZooBank="B54840C3-4AFE-475E-8B99-7FA8FD799645" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561701826928" checkinUser="plazi" docAuthor="Deschodt, Christian M. &amp; Davis, Adrian L. V." docDate="2019" docId="47328C10F127B336FF797B44FB0D80EA" docLanguage="en" docName="zootaxa.4624.2.10.pdf" docOrigin="Zootaxa 4624 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gyronotus ovalis Deschodt &amp; Davis 2019, new species" docType="treatment" docUuid="AA5B11ED-2B35-4A28-8453-19E5D618E07F" docUuidSource="ZooBank" docVersion="12" lastPageNumber="278" masterDocId="BB0BF468F125B335FFEE7C49FFDC8703" masterDocTitle="New southern African species and a revalidation in the dung beetle genus Gyronotus van Lansberge, 1874 (Scarabaeidae: Scarabaeinae) with an updated key" masterLastPageNumber="282" masterPageNumber="275" pageNumber="277" updateTime="1698737577708" updateUser="ExternalLinkService">
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<mods:title id="CC18E43CB1D7792F7AA81EDF7A2E7CE5">New southern African species and a revalidation in the dung beetle genus Gyronotus van Lansberge, 1874 (Scarabaeidae: Scarabaeinae) with an updated key</mods:title>
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<mods:namePart id="CFBB6F8ABDEA81EDB955565DC6FB9F91">Deschodt, Christian M.</mods:namePart>
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<mods:namePart id="E0D90F4DF7AD978FEC30D50DF2A43DBC">Davis, Adrian L. V.</mods:namePart>
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<mods:date id="FE33EC4B09DEE9A8B37724EACC1B2B11">2019</mods:date>
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<mods:number id="23E171ACE3FAC966D1329685C8799758">2019-06-28</mods:number>
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<mods:identifier id="BE66314FB78D78A62B3EAA48EECEC5F9" type="DOI">10.11646/zootaxa.4624.2.10</mods:identifier>
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<treatment id="47328C10F127B336FF797B44FB0D80EA" ID-DOI="http://doi.org/10.5281/zenodo.5923857" ID-GBIF-Taxon="158414873" ID-Zenodo-Dep="5923857" LSID="urn:lsid:zoobank.org:act:AA5B11ED-2B35-4A28-8453-19E5D618E07F" httpUri="http://treatment.plazi.org/id/47328C10F127B336FF797B44FB0D80EA" lastPageId="3" lastPageNumber="278" pageId="2" pageNumber="277">
<subSubSection id="87816E8DF127B337FF797B44FD368024" box="[151,746,1805,1832]" pageId="2" pageNumber="277" type="nomenclature">
<paragraph id="CF243D06F127B337FF797B44FD368024" blockId="2.[151,746,1805,1867]" box="[151,746,1805,1832]" pageId="2" pageNumber="277">
<heading id="946C8A6AF127B337FF797B44FD368024" bold="true" box="[151,746,1805,1832]" fontSize="11" level="1" pageId="2" pageNumber="277" reason="1">
<emphasis id="FDEFE114F127B337FF797B44FD368024" bold="true" box="[151,746,1805,1832]" pageId="2" pageNumber="277">
<taxonomicName id="089B4685F127B337FF797B44FD928024" ID-CoL="3HZSR" authority="Deschodt &amp; Davis" authorityName="Deschodt &amp; Davis" authorityYear="2019" box="[151,590,1805,1832]" class="Insecta" family="Scarabaeidae" genus="Gyronotus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="277" phylum="Arthropoda" rank="species" species="ovalis" status="sp. nov.">
<emphasis id="FDEFE114F127B337FF797B44FEB8802B" bold="true" box="[151,356,1805,1832]" italics="true" pageId="2" pageNumber="277">Gyronotus ovalis</emphasis>
Deschodt &amp; Davis
</taxonomicName>
,
<taxonomicNameLabel id="E6DC5C6FF127B337FDB57B47FD368024" box="[603,746,1805,1832]" pageId="2" pageNumber="277" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="87816E8DF127B336FF797B78FB0D80EA" lastPageId="3" lastPageNumber="278" pageId="2" pageNumber="277" type="description">
<paragraph id="CF243D06F127B337FF797B78FEA48048" blockId="2.[151,746,1805,1867]" box="[151,376,1841,1867]" pageId="2" pageNumber="277">
(
<figureCitation id="57A02183F127B337FF717B78FEDF8048" box="[159,259,1841,1867]" captionStart="FIGURE 1" captionStartId="2.[151,250,1626,1651]" captionTargetBox="[161,1431,553,1571]" captionTargetId="figure@2.[151,1436,547,1581]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Gyronotus van Lansberge, 1874 species, dorsal view. A, Gyronotus ovalis Deschodt &amp; Davis, new species; B, Gyronotus kearneyorum Deschodt &amp; Davis, new species; C, Gyronotus dracomontanus Deschodt &amp; Davis, new species; D, Gyronotus pumilus (Boheman, 1857); E, Gyronotus marginatus Péringuey, 1888. Scale bar = 2.0 mm." httpUri="https://zenodo.org/record/3259224/files/figure.png" pageId="2" pageNumber="277">Figs. 1A</figureCitation>
,
<figureCitation id="57A02183F127B337FEE17B78FEC18048" box="[271,285,1841,1867]" captionStart="FIGURE 2" captionStartId="3.[151,250,1640,1665]" captionTargetBox="[161,1421,752,1611]" captionTargetId="figure@3.[151,1436,741,1621]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagus of Gyronotus ovalis Deschodt &amp; Davis, new species.A, left side; B, right side; C, ventral view; D, dorsal view. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/3259226/files/figure.png" pageId="2" pageNumber="277">2</figureCitation>
A–D, 3)
</paragraph>
<paragraph id="CF243D06F127B337FF797B33FD2780D8" blockId="2.[151,1437,1913,2012]" pageId="2" pageNumber="277">
Type material.
<materialsCitation id="7FF3375BF127B337FEA67B30FC0C80B4" ID-GBIF-Occurrence="2412500438" collectingDate="2014-12-10" collectingDateMax="2014-12-12" collectingDateMin="2014-12-10" elevation="1070" latitude="-28.744514" location="Grassland" longLatPrecision="1" longitude="31.140556" municipality="Nkandla Forest Reserve" pageId="2" pageNumber="277" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="102083A4F127B337FEA67B30FE6F8090" box="[328,435,1913,1939]" pageId="2" pageNumber="277" type="holotype">Holotype</typeStatus>
,
<specimenCount id="D99DF68FF127B337FE2C7B30FE218090" box="[450,509,1913,1939]" pageId="2" pageNumber="277" type="male">male</specimenCount>
: “
<collectingMunicipality id="2F40A77CF127B337FDFB7B30FCEC8097" box="[533,816,1913,1940]" pageId="2" pageNumber="277">Nkandla Forest Reserve</collectingMunicipality>
,
<location id="CA446BDDF127B337FCD17B30FC738090" LSID="urn:lsid:plazi:treatment:47328C10F127B336FF797B44FB0D80EA:CA446BDDF127B337FCD17B30FC738090" box="[831,943,1913,1939]" latitude="-28.744514" longLatPrecision="1" longitude="31.140556" municipality="Nkandla Forest Reserve" name="Grassland" pageId="2" pageNumber="277">Grassland</location>
|
<geoCoordinate id="AAAF5BC1F127B337FC257B30FB808097" box="[971,1116,1913,1940]" degrees="28.744514" direction="south" orientation="latitude" pageId="2" pageNumber="277" precision="1" value="-28.744514">S28.744514°</geoCoordinate>
<geoCoordinate id="AAAF5BC1F127B337FB867B30FB228097" box="[1128,1278,1913,1940]" degrees="31.140557" direction="east" orientation="longitude" pageId="2" pageNumber="277" precision="1" value="31.140556">E31.140557°</geoCoordinate>
,
<quantity id="086390E3F127B337FAE37B30FA808090" box="[1293,1372,1913,1939]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="2" pageNumber="277" unit="m" value="1070.0">
<elevation id="44B6DA35F127B337FAE37B30FA808090" box="[1293,1372,1913,1939]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="2" pageNumber="277" unit="m" value="1070.0">1070m</elevation>
</quantity>
|
<date id="BB251BC6F127B337FA997B30FEC680B4" pageId="2" pageNumber="277" value="2014-12-10" valueMax="2014-12-12" valueMin="2014-12-10">
<collectingDate id="AB61E22EF127B337FA997B30FEC680B4" pageId="2" pageNumber="277" value="2014-12-10" valueMax="2014-12-12" valueMin="2014-12-10">10- 12.xii.2014</collectingDate>
</date>
,
<location id="CA446BDDF127B337FEC87BD4FDD580B4" LSID="urn:lsid:plazi:treatment:47328C10F127B336FF797B44FB0D80EA:CA446BDDF127B337FEC87BD4FDD580B4" box="[294,521,1949,1975]" latitude="-28.744514" longLatPrecision="1" longitude="31.140556" municipality="Nkandla Forest Reserve" name="Deschodt and Davis" pageId="2" pageNumber="277">Deschodt and Davis</location>
| pig dung baited pitfall traps” (TMSA)
</materialsCitation>
.
<materialsCitation id="7FF3375BF127B337FC357BD7FD2B80D8" ID-GBIF-Occurrence="2270855763" collectionCode="TMSA, UPSA, ISAM" pageId="2" pageNumber="277" specimenCount="10" specimenCount-female="8" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="102083A4F127B337FC357BD7FB9080BB" box="[987,1100,1950,1976]" pageId="2" pageNumber="277" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="D99DF68FF127B337FBB97BD4FB7280B4" box="[1111,1198,1949,1975]" pageId="2" pageNumber="277" type="male">2 males</specimenCount>
and
<specimenCount id="D99DF68FF127B337FB097BD4FA8B80B4" box="[1255,1367,1949,1975]" pageId="2" pageNumber="277" type="female">8 females</specimenCount>
, same data as holotype (
<quantity id="086390E3F127B337FEB37B88FE5480D8" box="[349,392,1985,2012]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="2" pageNumber="277" unit="in" value="5.0">5 in</quantity>
<collectionCode id="A98AA5C3F127B337FE617B88FE3C80D8" box="[399,480,1985,2011]" country="South Africa" httpUri="http://grbio.org/cool/s8gp-ktsi" name="Transvaal Museum" pageId="2" pageNumber="277">TMSA</collectionCode>
,
<quantity id="086390E3F127B337FE057B88FDCA80D8" box="[491,534,1985,2011]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="2" pageNumber="277" unit="in" value="3.0">3 in</quantity>
<collectionCode id="A98AA5C3F127B337FDF27B88FDBA80D8" box="[540,614,1985,2011]" country="South Africa" httpUri="http://grbio.org/cool/73qr-ciny" name="University of Pretoria" pageId="2" pageNumber="277">UPSA</collectionCode>
,
<quantity id="086390E3F127B337FD9C7B88FD4180D8" box="[626,669,1985,2011]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="277" unit="in" value="2.0">2 in</quantity>
<collectionCode id="A98AA5C3F127B337FD4A7B88FD3380D8" box="[676,751,1985,2011]" pageId="2" pageNumber="277">ISAM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="CF243D06F126B336FF297CD0FD9A87B0" blockId="3.[151,1437,153,719]" box="[199,582,153,179]" pageId="3" pageNumber="278">
Description of the
<typeStatus id="102083A4F126B336FE767CD0FE2187B0" box="[408,509,153,179]" pageId="3" pageNumber="278" type="holotype">holotype</typeStatus>
, male.
</paragraph>
<paragraph id="CF243D06F126B336FF297CF4FBC387DB" blockId="3.[151,1437,153,719]" box="[199,1055,189,216]" pageId="3" pageNumber="278">
Size. Length
<quantity id="086390E3F126B336FEB27CF4FE1C87D4" box="[348,448,189,215]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="3" pageNumber="278" unit="mm" value="12.3">12.3 mm</quantity>
and width 8.0 mm with head extended (not deflexed).
</paragraph>
<paragraph id="CF243D06F126B336FF297CA8FCE887F8" blockId="3.[151,1437,153,719]" box="[199,820,225,251]" pageId="3" pageNumber="278">Habitus. Body shape ovoid. Dark brown in appearance.</paragraph>
<paragraph id="CF243D06F126B336FF297D4CFAB88664" blockId="3.[151,1437,153,719]" pageId="3" pageNumber="278">Head. Tip of clypeus bidentate, separated by a wide “U” shaped depression. Surface, densely punctate, punctures closer together than their diameter. Clypeogenal sutures fairly distinct and nearly straight. Frontogenal suture clearly visible only at outside tenth, middle four fifths not visible. No tubercle present. Antennae dark orange.</paragraph>
<paragraph id="CF243D06F126B336FF297D38FEB88518" blockId="3.[151,1437,153,719]" pageId="3" pageNumber="278">Thorax. Pronotal disc densely punctate, punctures medium sized and round to slightly ovoid, almost touching each other. Lateral margin rounded. Prosternum with large faint punctures, surface dull. Mesosternum with large almost contiguous punctures. Metasternum with small sized punctures up to two times further apart than their diameter. Surface of prosternum and metasternum otherwise shagreened. Meso-metasternal suture clearly visible and somewhat curved.</paragraph>
<paragraph id="CF243D06F126B336FF297E6CFADA8560" blockId="3.[151,1437,153,719]" pageId="3" pageNumber="278">Elytra. Striae faint and narrow with punctures spaced at intervals of two to three puncture diameters. Interstriae punctate, punctures small separated by one to two puncture diameters. Surface otherwise shagreened.</paragraph>
<paragraph id="CF243D06F126B336FF297E24FEC185A8" blockId="3.[151,1437,153,719]" pageId="3" pageNumber="278">Pygidium with large shallow punctures about two puncture diameters apart, otherwise shagreened giving a dull appearance.</paragraph>
<paragraph id="CF243D06F126B336FF297EFCFCEC85CC" blockId="3.[151,1437,153,719]" box="[199,816,693,719]" pageId="3" pageNumber="278">
Aedeagus. Parameres bilaterally asymmetrical (
<figureCitation id="57A02183F126B336FD307EFCFCFF85CC" box="[734,803,693,719]" captionStart="FIGURE 2" captionStartId="3.[151,250,1640,1665]" captionTargetBox="[161,1421,752,1611]" captionTargetId="figure@3.[151,1436,741,1621]" captionTargetPageId="3" captionText="FIGURE 2. Aedeagus of Gyronotus ovalis Deschodt &amp; Davis, new species.A, left side; B, right side; C, ventral view; D, dorsal view. Scale bar = 1.0 mm." httpUri="https://zenodo.org/record/3259226/files/figure.png" pageId="3" pageNumber="278">Fig. 2</figureCitation>
).
</paragraph>
<caption id="9BE46D8EF126B336FF797A21FE7D81A7" httpUri="https://zenodo.org/record/3259226/files/figure.png" pageId="3" pageNumber="278" startId="3.[151,250,1640,1665]" targetBox="[161,1421,752,1611]" targetPageId="3">
<paragraph id="CF243D06F126B336FF797A21FE7D81A7" blockId="3.[151,1436,1640,1701]" pageId="3" pageNumber="278">
<emphasis id="FDEFE114F126B336FF797A21FECE8182" bold="true" box="[151,274,1640,1665]" pageId="3" pageNumber="278">FIGURE 2.</emphasis>
Aedeagus of
<taxonomicName id="089B4685F126B336FE757A20FCD58182" authority="Deschodt &amp; Davis" authorityName="Deschodt &amp; Davis" authorityYear="2019" box="[411,777,1640,1665]" class="Insecta" family="Scarabaeidae" genus="Gyronotus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="ovalis" status="sp. nov.">
<emphasis id="FDEFE114F126B336FE757A20FD948182" box="[411,584,1641,1665]" italics="true" pageId="3" pageNumber="278">Gyronotus ovalis</emphasis>
Deschodt &amp; Davis
</taxonomicName>
,
<taxonomicNameLabel id="E6DC5C6FF126B336FCFD7A20FC528182" box="[787,910,1640,1665]" pageId="3" pageNumber="278" rank="species">new species</taxonomicNameLabel>
. A, left side; B, right side; C, ventral view; D, dorsal view. Scale bar = 1.0 mm.
</paragraph>
</caption>
<paragraph id="CF243D06F126B336FF297A9AFC5A8036" blockId="3.[151,1437,1747,2026]" pageId="3" pageNumber="278">
Female. As with all
<taxonomicName id="089B4685F126B336FE447A9DFDC381ED" authorityName="van Lansberge" authorityYear="1874" box="[426,543,1748,1774]" class="Insecta" family="Scarabaeidae" genus="Gyronotus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="genus">
<emphasis id="FDEFE114F126B336FE447A9DFDC381ED" box="[426,543,1748,1774]" italics="true" pageId="3" pageNumber="278">Gyronotus</emphasis>
</taxonomicName>
species the tip of the front tibiae strongly curves downward with a shorter spur whereas the tip of the front tibiae of females is straight with a longer spur. Also, the mesotibiae of males strongly widen apically whereas the mesotibiae of females widen gradually.
</paragraph>
<paragraph id="CF243D06F126B336FF297B76FA4A805A" blockId="3.[151,1437,1747,2026]" box="[199,1430,1855,1882]" pageId="3" pageNumber="278">
Distribution.
<taxonomicName id="089B4685F126B336FEB37B09FD28805A" authority="Deschodt &amp; Davis" authorityName="Deschodt &amp; Davis" authorityYear="2019" box="[349,756,1855,1882]" class="Insecta" family="Scarabaeidae" genus="Gyronotus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="ovalis" status="sp. nov.">
<emphasis id="FDEFE114F126B336FEB37B09FDC0805A" box="[349,540,1855,1882]" italics="true" pageId="3" pageNumber="278">Gyronotus ovalis</emphasis>
Deschodt &amp; Davis
</taxonomicName>
,
<taxonomicNameLabel id="E6DC5C6FF126B336FD117B09FC5A805A" box="[767,902,1855,1882]" pageId="3" pageNumber="278" rank="species">new species</taxonomicNameLabel>
occurs in the grassland around Nkandla forest.
</paragraph>
<paragraph id="CF243D06F126B336FF297B2AFE9B80A2" blockId="3.[151,1437,1747,2026]" pageId="3" pageNumber="278">Etymology. The generic name and specific epithet is a pleonasm (Gyro=round and ovalis=oval) accentuating its ovoid shape.</paragraph>
<paragraph id="CF243D06F126B336FF297BE2FB0D80EA" blockId="3.[151,1437,1747,2026]" pageId="3" pageNumber="278">
Diagnosis. This species bears short setae on the elytra as opposed to almost glabrous elytra in the morphologically similar,
<taxonomicName id="089B4685F126B336FEC17B99FDC480EA" authorityName="Scholtz &amp; Howden" authorityYear="1987" box="[303,536,1999,2026]" class="Insecta" family="Scarabaeidae" genus="Gyronotus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="glabrosus">
<emphasis id="FDEFE114F126B336FEC17B99FDC480EA" box="[303,536,1999,2026]" italics="true" pageId="3" pageNumber="278">Gyronotus glabrosus</emphasis>
</taxonomicName>
and
<taxonomicName id="089B4685F126B336FDA17B99FBE480EA" authority="Deschodt &amp; Davis" authorityName="Deschodt &amp; Davis" authorityYear="2019" box="[591,1080,1999,2026]" class="Insecta" family="Scarabaeidae" genus="Gyronotus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="kearneyorum" status="sp. nov.">
<emphasis id="FDEFE114F126B336FDA17B99FC8380EA" box="[591,863,1999,2026]" italics="true" pageId="3" pageNumber="278">Gyronotus kearneyorum</emphasis>
Deschodt &amp; Davis
</taxonomicName>
,
<taxonomicNameLabel id="E6DC5C6FF126B336FBAD7B99FB1180EA" box="[1091,1229,1999,2026]" pageId="3" pageNumber="278" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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