treatments-xml/data/2D/72/DF/2D72DF6BE75FF661FF0F71D2159AFDC9.xml
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<document id="8DF284CD881BD7F4C0292EF86E7AF1CD" ID-CLB-Dataset="4462" ID-DOI="10.11646/phytotaxa.491.1.4" ID-GBIF-Dataset="f60d2dcc-f293-423c-9c89-1490c5a2537b" ID-ISSN="1179-3163" ID-Zenodo-Dep="5755359" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630386844379" checkinUser="marcus" docAuthor="Koldaeva, Marina N." docDate="2021" docId="2D72DF6BE75FF661FF0F71D2159AFDC9" docLanguage="en" docName="phytotaxa.491.1.4.pdf" docOrigin="Phytotaxa 491 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.491.1.4" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Chrysosplenium fallax Koldaeva 2021, sp. nov." docType="treatment" docVersion="7" lastPageNumber="43" masterDocId="D14BA713E75BF669FF8775061110FFA1" masterDocTitle="Chrysosplenium fallax (Saxifragaceae), a new species from the Russian Far East" masterLastPageNumber="46" masterPageNumber="35" pageNumber="39" updateTime="1699259338409" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="86CD7BD1AF8F45B9C5552973A55F2EFA">Chrysosplenium fallax (Saxifragaceae), a new species from the Russian Far East</mods:title>
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<treatment id="2D72DF6BE75FF661FF0F71D2159AFDC9" ID-DOI="http://doi.org/10.5281/zenodo.5755482" ID-GBIF-Taxon="185417804" ID-Zenodo-Dep="5755482" LSID="urn:lsid:plazi:treatment:2D72DF6BE75FF661FF0F71D2159AFDC9" httpUri="http://treatment.plazi.org/id/2D72DF6BE75FF661FF0F71D2159AFDC9" lastPageId="8" lastPageNumber="43" pageId="4" pageNumber="39">
<subSubSection id="EDC13DF6E75FF66DFF0F71D213F3FB4F" box="[136,739,1236,1263]" pageId="4" pageNumber="39" type="nomenclature">
<paragraph id="A5646E7DE75FF66DFF0F71D213F3FB4F" blockId="4.[136,739,1236,1263]" box="[136,739,1236,1263]" pageId="4" pageNumber="39">
<heading id="FE2CD911E75FF66DFF0F71D213F3FB4F" box="[136,739,1236,1263]" fontSize="11" level="2" pageId="4" pageNumber="39" reason="3">
<taxonomicName id="62DB15FEE75FF66DFF0F71D21313FB4F" ID-CoL="8H4WW" authority="Koldaeva" authorityName="Koldaeva" authorityYear="2021" box="[136,515,1236,1263]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="fallax" status="sp. nov.">
<emphasis id="97AFB26FE75FF66DFF0F71D2109EFB4E" bold="true" box="[136,398,1236,1263]" italics="true" pageId="4" pageNumber="39">Chrysosplenium fallax</emphasis>
Koldaeva
</taxonomicName>
,
<emphasis id="97AFB26FE75FF66DFD8871D21375FB4F" bold="true" box="[527,613,1236,1262]" italics="true" pageId="4" pageNumber="39">
<taxonomicNameLabel id="8C9C0F14E75FF66DFD8871D21375FB4F" box="[527,613,1236,1262]" pageId="4" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="3DE072F8E75FF66DFDF271D213C6FB4F" box="[629,726,1236,1262]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="5.[136,229,900,922]" captionStartId-1="5.[136,229,1927,1949]" captionStartId-2="6.[136,229,1899,1921]" captionStartId-3="7.[136,229,937,959]" captionTargetBox-0="[151,1436,187,873]" captionTargetBox-1="[151,1436,948,1903]" captionTargetBox-2="[151,1436,713,1874]" captionTargetBox-3="[151,1436,190,913]" captionTargetId-0="figure-15@5.[151,1436,187,873]" captionTargetId-1="figure-28@5.[151,1436,948,1903]" captionTargetId-2="figure-291@6.[151,1436,713,1874]" captionTargetId-3="figure-15@7.[151,1436,190,913]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 1. Chrysosplenium fallax. A. Habitat. B. Flowering plant." captionText-1="FIGURE 2. Chrysosplenium fallax. AB. Branching of a generative plant. C. Structure of generative and vegetative plants. D. Structure of the shoot system. E. A vegetative plant with slanting axial shoot: (a) generative shoot; (b) lateral above-ground vegetative shoots; (c) axial above-ground vegetative shoot; (d) rhizome; (e) underground stolons; (f) above-ground repent stolon-like shoot." captionText-2="FIGURE 3. Axial vegetative shoot of Chrysosplenium fallax. A. Apical part of a shoot at the beginning of growth. B. Apical part of a growing shoot. C. Apical part of a shoot at the end of growth. D. Shoot in the late growing season. E. Prostrate shoot with rising apex." captionText-3="FIGURE 4. Chrysosplenium fallax.A. Flower. B. Inflorescence. C. Capsules (side view). D. Capsules (top view). E. Seed. F. Seed surface structure." figureDoi-0="http://doi.org/10.5281/zenodo.5755361" figureDoi-1="http://doi.org/10.5281/zenodo.5755363" figureDoi-2="http://doi.org/10.5281/zenodo.5755365" figureDoi-3="http://doi.org/10.5281/zenodo.5755367" httpUri-0="https://zenodo.org/record/5755361/files/figure.png" httpUri-1="https://zenodo.org/record/5755363/files/figure.png" httpUri-2="https://zenodo.org/record/5755365/files/figure.png" httpUri-3="https://zenodo.org/record/5755367/files/figure.png" pageId="4" pageNumber="39">Fig. 14</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="EDC13DF6E75FF66DFF0F7019136FFA03" pageId="4" pageNumber="39" type="diagnosis">
<paragraph id="A5646E7DE75FF66DFF0F7019136FFA03" blockId="4.[136,1452,1311,1550]" pageId="4" pageNumber="39">
<emphasis id="97AFB26FE75FF66DFF0F7019101BFA94" bold="true" box="[136,267,1311,1333]" pageId="4" pageNumber="39">Diagnosis:—</emphasis>
Unlike
<taxonomicName id="62DB15FEE75FF66DFED4702610D1FA94" authorityName="Franchet" authorityYear="1890" box="[339,449,1312,1333]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="villosum">
<emphasis id="97AFB26FE75FF66DFED4702610D1FA94" box="[339,449,1312,1333]" italics="true" pageId="4" pageNumber="39">C. villosum</emphasis>
</taxonomicName>
,
<taxonomicName id="62DB15FEE75FF66DFE4970261333FA94" authorityName="Koldaeva" authorityYear="2021" box="[462,547,1312,1333]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="fallax">
<emphasis id="97AFB26FE75FF66DFE4970261333FA94" box="[462,547,1312,1333]" italics="true" pageId="4" pageNumber="39">C. fallax</emphasis>
</taxonomicName>
forms a plagiotropic rhizome with long internodes and specialized underground stolons. The vegetative shoots do not take root by the end of the growing season, and are with acervate leaves on the top.
<taxonomicName id="62DB15FEE75FF66DFB54704214BCFAF8" authorityName="Koldaeva" authorityYear="2021" box="[1235,1452,1348,1369]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="fallax">
<emphasis id="97AFB26FE75FF66DFB54704214BCFAF8" box="[1235,1452,1348,1369]" italics="true" pageId="4" pageNumber="39">Chrysosplenium fallax</emphasis>
</taxonomicName>
differs from
<taxonomicName id="62DB15FEE75FF66DFEBF706E10B1FADC" authorityName="Maximowicz" authorityYear="1859" box="[312,417,1384,1405]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="4" pageNumber="39" phylum="Tracheophyta" rank="species" species="pilosum">
<emphasis id="97AFB26FE75FF66DFEBF706E10B1FADC" box="[312,417,1384,1405]" italics="true" pageId="4" pageNumber="39">C. pilosum</emphasis>
</taxonomicName>
by vegetative shoots lacking branching in the distal part, with larger acervate leaves on the top, which are pubescent on the abaxial side of the leaf blade.
</paragraph>
</subSubSection>
<subSubSection id="EDC13DF6E75FF66DFF0F70B61044F9AF" pageId="4" pageNumber="39" type="materials_examined">
<paragraph id="A5646E7DE75FF66DFF0F70B61044F9AF" blockId="4.[136,1452,1311,1550]" pageId="4" pageNumber="39">
<emphasis id="97AFB26FE75FF66DFF0F70B611D2FA64" bold="true" box="[136,194,1456,1477]" pageId="4" pageNumber="39">Type:</emphasis>
<materialsCitation id="15B36420E75FF66DFF5C70B6105FF9AF" ID-GBIF-Occurrence="3341011301" collectingDate="2017-05-12" collectionCode="VLA" collectorName="M. N. Koldaeva" country="Russia" county="Vladivostoksky Urban Okrug" location="Malaya Pionerskaya River valley" municipality="Muravyev-Amursky Peninsula" pageId="4" pageNumber="39" specimenCount="1" stateProvince="Primorskiy" typeStatus="holotype">
<collectingCountry id="DDCC2EEDE75FF66DFF5C70B610CFFA64" box="[219,479,1456,1477]" name="Russia" pageId="4" pageNumber="39">RUSSIAN FEDERATION</collectingCountry>
.
<collectingRegion id="671FA09FE75FF66DFE6F70B61390FA67" box="[488,640,1456,1478]" country="Russia" name="Primorskiy" pageId="4" pageNumber="39">Primorsky Krai</collectingRegion>
,
<collectingCounty id="4C0516F1E75FF66DFD0E70B612B9FA67" box="[649,937,1456,1478]" pageId="4" pageNumber="39">Vladivostoksky Urban Okrug</collectingCounty>
,
<collectingMunicipality id="4500F407E75FF66DFC3470B615CEFA67" box="[947,1246,1456,1478]" pageId="4" pageNumber="39">Muravyev-Amursky Peninsula</collectingMunicipality>
,
<location id="A00438A6E75FF66DFB6070B61024FA4B" LSID="urn:lsid:plazi:treatment:2D72DF6BE75FF661FF0F71D2159AFDC9:A00438A6E75FF66DFB6070B61024FA4B" country="Russia" county="Vladivostoksky Urban Okrug" municipality="Muravyev-Amursky Peninsula" name="Malaya Pionerskaya River valley" pageId="4" pageNumber="39" stateProvince="Primorskiy">Malaya Pionerskaya River valley</location>
, edge of a coniferous-broadleaved forest, moist habitat,
<date id="D16548BDE75FF66DFCCF70D212D8FA48" box="[840,968,1492,1513]" pageId="4" pageNumber="39" value="2017-05-12">
<collectingDate id="C121B155E75FF66DFCCF70D212D8FA48" box="[840,968,1492,1513]" pageId="4" pageNumber="39" value="2017-05-12">12 May 2017</collectingDate>
</date>
,
<emphasis id="97AFB26FE75FF66DFC5670D21585FA48" box="[977,1173,1492,1513]" italics="true" pageId="4" pageNumber="39">
<collectorName id="082E0BABE75FF66DFC5670D21572FA48" box="[977,1122,1492,1513]" pageId="4" pageNumber="39">M.N. Koldaeva</collectorName>
0115
</emphasis>
(
<typeStatus id="7A60D0DFE75FF66DFB2670D215E7FA4B" box="[1185,1271,1492,1514]" pageId="4" pageNumber="39" type="holotype">holotype</typeStatus>
LE!, isotypes LE!,
<collectionCode id="C3CAF6B8E75FF66DFF3A70FE11E5F9AC" box="[189,245,1528,1549]" country="United Kingdom" httpUri="http://grbio.org/cool/rz2a-nf0j" name="Veterinary Laboratory Agency" pageId="4" pageNumber="39">VLA</collectionCode>
!, VBGI!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="EDC13DF6E75FF66FFF0F733A10AEFD11" lastPageId="6" lastPageNumber="41" pageId="4" pageNumber="39" type="description">
<paragraph id="A5646E7DE75FF66FFF0F733A10AEFD11" blockId="4.[136,1452,1596,2091]" lastBlockId="6.[136,1452,158,689]" lastPageId="6" lastPageNumber="41" pageId="4" pageNumber="39">
<emphasis id="97AFB26FE75FF66DFF0F733A10FFF9F7" bold="true" box="[136,495,1596,1622]" pageId="4" pageNumber="39">Description of the new species</emphasis>
:—Perennial herb, pulpy, pubescent with long, white hairs, reddish when dry. Leaves opposite. Stems and early leaves are reddish below. Rhizome
<quantity id="6223C398E75FF66DFCD1736612D1F9DA" box="[854,961,1632,1659]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" metricValueMax="5.0" metricValueMin="1.2" pageId="4" pageNumber="39" unit="cm" value="3.1" valueMax="5.0" valueMin="1.2">1.25 cm</quantity>
long, underground, plagiotropic, with 25 elongated internodes. Flowering stem single, erect, 1420(24) cm tall, middle part sparsely pubescent, and base and apical part almost glabrous. Cauline leaves in 24 pairs. Leaf blade subflabellate or flabellate-rounded, 0.31.3 ×
<quantity id="6223C398E75FF66DFF0F73CA101AF946" box="[136,266,1740,1767]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.9" metricValueMin="0.4" pageId="4" pageNumber="39" unit="cm" value="1.15" valueMax="1.9" valueMin="0.4">0.41.9 cm</quantity>
, base cuneate or broadly cuneate, adaxial surface glabrous, abaxial surface glabrous or with single hairs along veins at base, often with red specks; margin ciliate, with 579 smoothed-serrated crenates; petiole
<quantity id="6223C398E75FF66DFAD073F611BAF88E" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="9.0" metricValueMin="3.0" pageId="4" pageNumber="39" unit="cm" value="0.6" valueMax="0.9" valueMin="0.3">0.30.9 cm</quantity>
in length. Cymes compact, 616 flowered, surrounded by large, greenish-yellow bracts. Inflorescence branches and bracteal petioles with solitary long hairs. Flowers goblet-shaped, yellow, sometimes with red specks; pedicels short (
<quantity id="6223C398E75FF66DFF57725A1059F8D6" box="[208,329,1884,1911]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="3.0" metricValueMin="0.8" pageId="4" pageNumber="39" unit="mm" value="1.9" valueMax="3.0" valueMin="0.8">0.83 mm</quantity>
). Sepals 4, erect, convex outside, with the tip slightly bent outward; external almost rounded 2.2 ×
<quantity id="6223C398E75FF66DFF0F728611F3F83A" box="[136,227,1920,1947]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="4" pageNumber="39" unit="mm" value="2.1">2.1 mm</quantity>
; internal ovoid 1.8 ×
<quantity id="6223C398E75FF66DFE5B72861326F83A" box="[476,566,1920,1947]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="39" unit="mm" value="1.5">1.5 mm</quantity>
, shorter than external ones. Stamens 8, equal to calyx; anthers yellow. Ovary subsuperior. Styles erect, slightly divergent. Nectary not prominent, green. Capsule
<quantity id="6223C398E75FF66DFBC672A215B2F81E" box="[1089,1186,1956,1983]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="4" pageNumber="39" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">68 mm</quantity>
, ca. 22.5 times longer than calyx. Carpels 2, slightly unequal, divergent, arcuately curved. Seeds suborbicular, 0.70.8 ×
<quantity id="6223C398E75FF66DFB4D72CE1442F842" box="[1226,1362,1992,2019]" metricMagnitude="-4" metricUnit="m" metricValue="6.499999999999999" metricValueMax="7.0" metricValueMin="6.0" pageId="4" pageNumber="39" unit="mm" value="0.6499999999999999" valueMax="0.7" valueMin="0.6">0.60.7 mm</quantity>
, brown, minutely papillose, with 1823 longitudinal rows of large, densely located tubercles. Tubercles almost cylindrical, with one, rarely two papillae. Vegetative shoots above-ground (sterile) and underground (stolons). Above-ground shoots orthotropic (axial and lateral), prostrate and repent, stolon-like, pubescent with long hairs, most densely aggregated at apical part. Orthotropic shoots not evenly leafed; leaves highly unequal, with largest ones acervate to resemble a rosette at the apex but without forming a true rosette; apical part not rooted. Axial sterile shoots up to
<quantity id="6223C398E75DF66FFAA575E01479FEA0" box="[1314,1385,230,257]" metricMagnitude="-1" metricUnit="m" metricValue="2.1" pageId="6" pageNumber="41" unit="cm" value="21.0">21 cm</quantity>
long, with branches or non-branching. Leaves 49(11) pairs; distal 23 pairs largest, 2.15.6 × 2.0
<quantity id="6223C398E75DF66FFB52740C1433FE84" box="[1237,1315,266,293]" metricMagnitude="-2" metricUnit="m" metricValue="3.7" pageId="6" pageNumber="41" unit="cm" value="3.7">3.7 cm</quantity>
(except the uppermost one), with short internodes (
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); blade rounded, rounded-ovate or elliptical; base cuneate; margin undulate-crenate, crenate-sinuate or almost smooth, ciliate, with 57 crenates on each side. Lateral sterile shoots up to
<quantity id="6223C398E75DF66FFF4A74701004FE30" box="[205,276,374,401]" metricMagnitude="-1" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="41" unit="cm" value="16.0">16 cm</quantity>
long, with basal internode elongated; leaves (1)23 pairs, acervate to resemble a rosette at the apex. Pubescence of leaf abaxial side along main veins appears on 1
<superScript id="52AEC335E75DF66FFCBE749C1254FE09" attach="left" box="[825,836,410,424]" fontSize="6" pageId="6" pageNumber="41">st</superScript>
or 2
<superScript id="52AEC335E75DF66FFCF2749C1295FE09" attach="left" box="[885,901,410,424]" fontSize="6" pageId="6" pageNumber="41">nd</superScript>
pairs of lateral shoots or in 2
<superScript id="52AEC335E75DF66FFB40749C15C7FE09" attach="left" box="[1223,1239,410,424]" fontSize="6" pageId="6" pageNumber="41">nd</superScript>
to 5
<superScript id="52AEC335E75DF66FFA8F749C1405FE09" attach="left" box="[1288,1301,410,424]" fontSize="6" pageId="6" pageNumber="41">th</superScript>
pairs of axial shoots; in upper pairs, pubescence becomes denser. Blade adaxial side glabrous or with solitary hairs at the periphery, sometimes with silvery dots. Petioles sparsely pilose on both sides. Above-ground prostrate shoots with rising apex or flagelliform, evenly leafed; leaves highly unequal, with largest ones located in the third quarter from shoot base. Repent stolon-like shoots 12, evenly leafed; leaves 56 pairs, similar, small. Underground stolons 13(4), up to
<quantity id="6223C398E75DF66FFA08772C11BAFDC8" metricMagnitude="-1" metricUnit="m" metricValue="4.8" pageId="6" pageNumber="41" unit="cm" value="48.0">48 cm</quantity>
long,
<quantity id="6223C398E75DF66FFF7477481076FDC8" box="[243,358,590,617]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="6" pageNumber="41" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
in diameter, unbranched or weakly branched in distal part, white. An approximately 10-cm-long distal portion of stolon remains by the end of the growing season. In the population, most shoots are sterile axial; flowering shoots are single.
</paragraph>
</subSubSection>
<caption id="F1A43EF5E75EF66CFF0F76821202FC3B" ID-DOI="http://doi.org/10.5281/zenodo.5755361" ID-Zenodo-Dep="5755361" box="[136,786,900,922]" httpUri="https://zenodo.org/record/5755361/files/figure.png" pageId="5" pageNumber="40" startId="5.[136,229,900,922]" targetBox="[151,1436,187,873]" targetPageId="5">
<paragraph id="A5646E7DE75EF66CFF0F76821202FC3B" blockId="5.[136,786,900,922]" box="[136,786,900,922]" pageId="5" pageNumber="40">
<emphasis id="97AFB26FE75EF66CFF0F768211EEFC38" bold="true" box="[136,254,900,922]" pageId="5" pageNumber="40">FIGURE 1.</emphasis>
<taxonomicName id="62DB15FEE75EF66CFE84768210CCFC38" authorityName="Koldaeva" authorityYear="2021" box="[259,476,900,921]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="fallax">
<emphasis id="97AFB26FE75EF66CFE84768210CCFC38" box="[259,476,900,921]" italics="true" pageId="5" pageNumber="40">Chrysosplenium fallax</emphasis>
</taxonomicName>
. A. Habitat. B. Flowering plant.
</paragraph>
</caption>
<caption id="F1A43EF5E75EF66CFF0F728115E3F844" ID-DOI="http://doi.org/10.5281/zenodo.5755363" ID-Zenodo-Dep="5755363" httpUri="https://zenodo.org/record/5755363/files/figure.png" pageId="5" pageNumber="40" startId="5.[136,229,1927,1949]" targetBox="[151,1436,948,1903]" targetPageId="5">
<paragraph id="A5646E7DE75EF66CFF0F728115E3F844" blockId="5.[136,1452,1927,2021]" pageId="5" pageNumber="40">
<emphasis id="97AFB26FE75EF66CFF0F728111EEF83D" bold="true" box="[136,254,1927,1949]" pageId="5" pageNumber="40">FIGURE 2.</emphasis>
<taxonomicName id="62DB15FEE75EF66CFE83728110CEF83D" authorityName="Koldaeva" authorityYear="2021" box="[260,478,1927,1948]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="5" pageNumber="40" phylum="Tracheophyta" rank="species" species="fallax">
<emphasis id="97AFB26FE75EF66CFE83728110CEF83D" box="[260,478,1927,1948]" italics="true" pageId="5" pageNumber="40">Chrysosplenium fallax</emphasis>
</taxonomicName>
. AB. Branching of a generative plant. C. Structure of generative and vegetative plants. D. Structure of the shoot system. E. A vegetative plant with slanting axial shoot: (a) generative shoot; (b) lateral above-ground vegetative shoots; (c) axial above-ground vegetative shoot; (d) rhizome; (e) underground stolons; (f) above-ground repent stolon-like shoot.
</paragraph>
</caption>
<caption id="F1A43EF5E75DF66FFF0F726D1480F805" ID-DOI="http://doi.org/10.5281/zenodo.5755365" ID-Zenodo-Dep="5755365" httpUri="https://zenodo.org/record/5755365/files/figure.png" pageId="6" pageNumber="41" startId="6.[136,229,1899,1921]" targetBox="[151,1436,713,1874]" targetPageId="6">
<paragraph id="A5646E7DE75DF66FFF0F726D1480F805" blockId="6.[136,1452,1899,1957]" pageId="6" pageNumber="41">
<emphasis id="97AFB26FE75DF66FFF0F726D11EFF821" bold="true" box="[136,255,1899,1921]" pageId="6" pageNumber="41">FIGURE 3.</emphasis>
Axial vegetative shoot of
<taxonomicName id="62DB15FEE75DF66FFD84726D13CDF821" authorityName="Koldaeva" authorityYear="2021" box="[515,733,1899,1920]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="41" phylum="Tracheophyta" rank="species" species="fallax">
<emphasis id="97AFB26FE75DF66FFD84726D13CDF821" box="[515,733,1899,1920]" italics="true" pageId="6" pageNumber="41">Chrysosplenium fallax</emphasis>
</taxonomicName>
. A. Apical part of a shoot at the beginning of growth. B. Apical part of a growing shoot. C. Apical part of a shoot at the end of growth. D. Shoot in the late growing season. E. Prostrate shoot with rising apex.
</paragraph>
</caption>
<caption id="F1A43EF5E75CF66EFF0F76AF11F3FC43" ID-DOI="http://doi.org/10.5281/zenodo.5755367" ID-Zenodo-Dep="5755367" httpUri="https://zenodo.org/record/5755367/files/figure.png" pageId="7" pageNumber="42" startId="7.[136,229,937,959]" targetBox="[151,1436,190,913]" targetPageId="7">
<paragraph id="A5646E7DE75CF66EFF0F76AF11F3FC43" blockId="7.[136,1451,937,994]" pageId="7" pageNumber="42">
<emphasis id="97AFB26FE75CF66EFF0F76AF11EDFC1F" bold="true" box="[136,253,937,959]" pageId="7" pageNumber="42">FIGURE 4.</emphasis>
<taxonomicName id="62DB15FEE75CF66EFE8576AF10CAFC1F" authorityName="Koldaeva" authorityYear="2021" box="[258,474,937,958]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="species" species="fallax">
<emphasis id="97AFB26FE75CF66EFE8576AF10CAFC1F" box="[258,474,937,958]" italics="true" pageId="7" pageNumber="42">Chrysosplenium fallax</emphasis>
</taxonomicName>
. A. Flower. B. Inflorescence. C. Capsules (side view). D. Capsules (top view). E. Seed. F. Seed surface structure.
</paragraph>
</caption>
<caption id="F1A43EF5E75CF66EFF0F727A1057F817" ID-DOI="http://doi.org/10.5281/zenodo.5755369" ID-Zenodo-Dep="5755369" httpUri="https://zenodo.org/record/5755369/files/figure.png" pageId="7" pageNumber="42" startId="7.[136,229,1916,1938]" targetBox="[160,1430,1026,1885]" targetPageId="7">
<paragraph id="A5646E7DE75CF66EFF0F727A1057F817" blockId="7.[136,1451,1916,1974]" pageId="7" pageNumber="42">
<emphasis id="97AFB26FE75CF66EFF0F727A1011F830" bold="true" box="[136,257,1916,1938]" pageId="7" pageNumber="42">FIGURE 5.</emphasis>
Distribution of
<taxonomicName id="62DB15FEE75CF66EFE23727A1390F830" authorityName="Koldaeva" authorityYear="2021" box="[420,640,1916,1937]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="7" pageNumber="42" phylum="Tracheophyta" rank="species" species="fallax">
<emphasis id="97AFB26FE75CF66EFE23727A1390F830" box="[420,640,1916,1937]" italics="true" pageId="7" pageNumber="42">Chrysosplenium fallax</emphasis>
</taxonomicName>
. Red dots indicate localities of herbarium specimens; green dots are localities of authors specimens.
</paragraph>
</caption>
<subSubSection id="EDC13DF6E753F661FF3A759B10EDFE15" pageId="8" pageNumber="43" type="distribution">
<paragraph id="A5646E7DE753F661FF3A759B1066FE85" blockId="8.[136,1452,157,616]" pageId="8" pageNumber="43">
<emphasis id="97AFB26FE753F661FF3A759B105FFF16" bold="true" box="[189,335,157,183]" pageId="8" pageNumber="43">Distribution</emphasis>
:—
<taxonomicName id="62DB15FEE753F661FEF47598137FFF19" authorityName="Koldaeva" authorityYear="2021" box="[371,623,158,184]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="fallax">
<emphasis id="97AFB26FE753F661FEF47598137FFF19" box="[371,623,158,184]" italics="true" pageId="8" pageNumber="43">Chrysosplenium fallax</emphasis>
</taxonomicName>
is known only from southern
<collectingRegion id="671FA09FE753F661FC3C7598157BFF19" box="[955,1131,158,184]" country="Russia" name="Primorskiy" pageId="8" pageNumber="43">Primorsky Krai</collectingRegion>
,
<collectingCountry id="DDCC2EEDE753F661FBF27598145BFF19" box="[1141,1355,158,184]" name="Russia" pageId="8" pageNumber="43">Russian Federation</collectingCountry>
(
<figureCitation id="3DE072F8E753F661FADD7598148DFF18" box="[1370,1437,158,185]" captionStart="FIGURE 5" captionStartId="7.[136,229,1916,1938]" captionTargetBox="[160,1430,1026,1885]" captionTargetId="figure-41@7.[151,1436,1020,1891]" captionTargetPageId="7" captionText="FIGURE 5. Distribution of Chrysosplenium fallax. Red dots indicate localities of herbarium specimens; green dots are localities of authors specimens." figureDoi="http://doi.org/10.5281/zenodo.5755369" httpUri="https://zenodo.org/record/5755369/files/figure.png" pageId="8" pageNumber="43">Fig. 5</figureCitation>
). It is most frequently found in Shkotovsky District, as well as in Ussuriysky and Vladivostoksky (except islands) Urban Okrugs. In other areas, where plants of this species were collected, it is rare and has populations on small areas and with few individuals.
</paragraph>
<paragraph id="A5646E7DE753F661FF3A742B13A2FECD" blockId="8.[136,1452,157,616]" pageId="8" pageNumber="43">
<emphasis id="97AFB26FE753F661FF3A742B1009FEE6" bold="true" box="[189,281,301,327]" pageId="8" pageNumber="43">Habitat</emphasis>
:—The new species grows in broad-leaved forests on moist and damp parts of gentle slopes, along river valleys and streams, at an elevation of
<quantity id="6223C398E753F661FDBD745413BEFECD" box="[570,686,338,364]" metricMagnitude="2" metricUnit="m" metricValue="2.0" metricValueMax="3.8" metricValueMin="0.2" pageId="8" pageNumber="43" unit="m" value="200.0" valueMax="380.0" valueMin="20.0">20380 m</quantity>
.
</paragraph>
<paragraph id="A5646E7DE753F661FF3A747010EDFE15" blockId="8.[136,1452,157,616]" pageId="8" pageNumber="43">
<emphasis id="97AFB26FE753F661FF3A74701029FE31" bold="true" box="[189,313,374,400]" pageId="8" pageNumber="43">Phenology</emphasis>
:—The flowering season of
<taxonomicName id="62DB15FEE753F661FDFF747013CAFE31" authorityName="Koldaeva" authorityYear="2021" box="[632,730,374,400]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="fallax">
<emphasis id="97AFB26FE753F661FDFF747013CAFE31" box="[632,730,374,400]" italics="true" pageId="8" pageNumber="43">C. fallax</emphasis>
</taxonomicName>
is the first and second ten days of May; the fruiting season, the first and second ten days of June.
</paragraph>
</subSubSection>
<subSubSection id="EDC13DF6E753F661FF3A74B8159AFDC9" pageId="8" pageNumber="43" type="etymology">
<paragraph id="A5646E7DE753F661FF3A74B81517FD81" blockId="8.[136,1452,157,616]" pageId="8" pageNumber="43">
<emphasis id="97AFB26FE753F661FF3A74B8102EFE79" bold="true" box="[189,318,446,472]" pageId="8" pageNumber="43">Etymology</emphasis>
:—The name of this species means that in its different phenological states
<taxonomicName id="62DB15FEE753F661FBF374B815C4FE79" authorityName="Koldaeva" authorityYear="2021" box="[1140,1236,446,472]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="fallax">
<emphasis id="97AFB26FE753F661FBF374B815C4FE79" box="[1140,1236,446,472]" italics="true" pageId="8" pageNumber="43">C. fallax</emphasis>
</taxonomicName>
becomes similar to one species of
<taxonomicName id="62DB15FEE753F661FEAA74E410F3FE5D" authorityName="Koldaeva" authorityYear="2021" box="[301,483,482,508]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="genus">
<emphasis id="97AFB26FE753F661FEAA74E410F3FE5D" box="[301,483,482,508]" italics="true" pageId="8" pageNumber="43">Chrysosplenium</emphasis>
</taxonomicName>
and then to another, which previously led to the misidentification of its specimens and confusion with other species, and, therefore, it has long remained undiscovered.
</paragraph>
<paragraph id="A5646E7DE753F661FF3A772F159AFDC9" blockId="8.[136,1452,157,616]" pageId="8" pageNumber="43">
<emphasis id="97AFB26FE753F661FF3A772F104FFDE2" bold="true" box="[189,351,553,579]" pageId="8" pageNumber="43">Relationships</emphasis>
:—
<taxonomicName id="62DB15FEE753F661FE04772C1390FDE5" authorityName="Koldaeva" authorityYear="2021" box="[387,640,554,580]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="fallax">
<emphasis id="97AFB26FE753F661FE04772C1390FDE5" box="[387,640,554,580]" italics="true" pageId="8" pageNumber="43">Chrysosplenium fallax</emphasis>
</taxonomicName>
belongs to the ser.
<taxonomicName id="62DB15FEE753F661FCD1772C128EFDE5" box="[854,926,554,580]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="8" pageNumber="44" phylum="Tracheophyta" rank="series" series="Pilosa">
<emphasis id="97AFB26FE753F661FCD1772C128EFDE5" box="[854,926,554,580]" italics="true" pageId="8" pageNumber="43">Pilosa</emphasis>
</taxonomicName>
, being closest to
<taxonomicName id="62DB15FEE753F661FBD8772C15C8FDE5" authorityName="Maximowicz" authorityYear="1859" box="[1119,1240,554,580]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="pilosum">
<emphasis id="97AFB26FE753F661FBD8772C15C8FDE5" box="[1119,1240,554,580]" italics="true" pageId="8" pageNumber="43">C. pilosum</emphasis>
</taxonomicName>
and
<taxonomicName id="62DB15FEE753F661FA8A772C149DFDE5" authorityName="Franchet" authorityYear="1890" box="[1293,1421,554,580]" class="Magnoliopsida" family="Saxifragaceae" genus="Chrysosplenium" kingdom="Plantae" order="Saxifragales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="villosum">
<emphasis id="97AFB26FE753F661FA8A772C149DFDE5" box="[1293,1421,554,580]" italics="true" pageId="8" pageNumber="43">C. villosum</emphasis>
</taxonomicName>
. A list of differences from the species for which it is usually mistaken are presented in
<tableCitation id="E8595BC6E753F661FBB777481596FDC9" box="[1072,1158,590,616]" captionStart="TABLE 2" captionStartId="3.[136,224,160,184]" captionText="TABLE 2. Comparison of the main morphological characters of the related Chrysosplenium taxa." pageId="8" pageNumber="43">Table 2</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>