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<document ID-CLB-Dataset="6047" ID-DOI="http://dx.doi.org/10.3897/zookeys.1044.58844" ID-GBIF-Dataset="7bae8d8a-c95f-44bc-84b7-a8de8a0f1ae9" ID-PMC="PMC8222277" ID-Pensoft-Pub="1313-2970-1044-269" ID-Pensoft-UUID="8EC99E5110F45866A56F56BA7EA3D3AB" ID-PubMed="34183880" ID-ZooBank="DE81899437314028BBE9C53C4CE220AC" ModsDocID="1313-2970-1044-269" checkinTime="1623875423107" checkinUser="pensoft" docAuthor="Giachino, Pier Mauro, Eberhard, Stefan &amp; Perina, Giulia" docDate="2021" docId="09074B2F2DE85F688FC4283571657045" docLanguage="en" docName="ZooKeys 1044: 269-337" docOrigin="ZooKeys 1044" docPubDate="2021-06-16" docSource="http://dx.doi.org/10.3897/zookeys.1044.58844" docTitle="Pilbaraphanus Giachino &amp; Eberhard &amp; Perina 2021, gen. nov." docType="treatment" docUuid="F300FB18-16FE-468B-B9BA-CC5C0926CFDF" docUuidSource="ZooBank" docVersion="6" id="8EC99E5110F45866A56F56BA7EA3D3AB" lastPageNumber="269" masterDocId="8EC99E5110F45866A56F56BA7EA3D3AB" masterDocTitle="A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia" masterLastPageNumber="337" masterPageNumber="269" pageNumber="269" updateTime="1732669819433" updateUser="ExternalLinkService">
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<mods:title id="766C5934A5629E214FCF51CBBD40F4A0">A rich fauna of subterranean short-range endemic Anillini (Coleoptera, Carabidae, Trechinae) from semi-arid regions of Western Australia</mods:title>
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<mods:namePart id="3AB3BBEB93889304A4D09BB4C680405C">Giachino, Pier Mauro</mods:namePart>
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<mods:affiliation id="EE146699B8ED3E203025DFC6FD08165F">World Biodiversity Association onlus. Private: via della Trinita 13, I- 10010 San Martino Canavese (TO), Italy</mods:affiliation>
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<mods:namePart id="FDF3FE36FB91AFB362D869001A141B4D">Eberhard, Stefan</mods:namePart>
<mods:affiliation id="C457F7CD119DAD4200C64541C98F4D23">Subterranean Ecology Pty Ltd, 227 Coningham Road, Coningham, TAS 7054, Australia</mods:affiliation>
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<mods:namePart id="508D7A6F7446B15837D4E7ED27BBF1CB">Perina, Giulia</mods:namePart>
<mods:nameIdentifier id="1BBE7624DC25CF78450EB133EFF906D5" type="ORCID">https://orcid.org/0000-0002-0349-3803</mods:nameIdentifier>
<mods:affiliation id="3208723BFF25E1D1D829ABAA3CF74949">Collections and Research, Western Australian Museum, 49 Kew Street, Welshpool, WA 6106, Australia</mods:affiliation>
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<mods:date id="1475090FA32FCC37D97D0268B0886B16">2021</mods:date>
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<mods:identifier id="F0E08FCFF757E54E7F12A27F3C63A736" type="DOI">http://dx.doi.org/10.3897/zookeys.1044.58844</mods:identifier>
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<treatment id="09074B2F2DE85F688FC4283571657045" ID-GBIF-Taxon="182754089" LSID="urn:lsid:zoobank.org:act:F300FB18-16FE-468B-B9BA-CC5C0926CFDF" httpUri="http://treatment.plazi.org/id/09074B2F2DE85F688FC4283571657045" lastPageNumber="269" pageId="0" pageNumber="269" scope_family="Carabidae" scope_order="Coleoptera" scope_subFamily="Trechinae">
<subSubSection id="7E1B4195A901B7E30E4FD8D406617E04" pageId="0" pageNumber="269" type="nomenclature">
<paragraph id="B705D35C93C687DA50E2D0A3D25C5C4C" pageId="0" pageNumber="269">
<taxonomicName id="D8C15C2136DB09B5670D01FF04A369A3" ID-CoL="8MJDR" LSID="http://zoobank.org/F300FB18-16FE-468B-B9BA-CC5C0926CFDF" authority="Giachino &amp; Eberhard &amp; Perina, 2021" authorityName="Giachino &amp; Eberhard &amp; Perina" authorityYear="2021" class="Insecta" family="Carabidae" genus="Pilbaraphanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilbaraphanus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="genus" status="gen. nov.">Pilbaraphanus</taxonomicName>
<taxonomicNameLabel id="A6D417B951AF33AD0F3129AD40FE569A" pageId="0" pageNumber="269">gen. nov.</taxonomicNameLabel>
<figureCitation id="2C60D45A03D18887E72E3C5BCDE73EE6" captionStart="Figures 1014" captionStartId="F4" captionText="Figures 10 - 14. Pilbaraphanus gen. et spp. nov., habitus (10, 13) right metafemur and metatrochanter in ventral view (11, 14) aedeagus in lateral view (12) 10 - 12 P. chichesterianus sp. nov., HT ♂ 13, 14 P. bilybarianus sp. nov. HT ♀. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1044.58844.figures10-14" httpUri="https://binary.pensoft.net/fig/553763" pageId="0" pageNumber="269">Figs 10-14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C54B87AFBDCE9D6CC6389A6E1F96B76A" pageId="0" pageNumber="269" type="type species">
<paragraph id="9AB0DD86670F4841231CD26F4968B143" pageId="0" pageNumber="269">Type species.</paragraph>
<paragraph id="B93C8CBCB67958587E1440DE25D9F9E2" pageId="0" pageNumber="269">
<taxonomicName id="A8100987C3DA18AEE8936C0FEB907FD2" authorityName="Giachino &amp; Eberhard &amp; Perina" authorityYear="2021" class="Insecta" family="Carabidae" genus="Pilbaraphanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilbaraphanus chichesterianus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="species" species="chichesterianus">
<emphasis id="E5701F893407D582D627B3DEA42E773A" italics="true" pageId="0" pageNumber="269">Pilbaraphanus chichesterianus</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection id="60AEF433B9A4CC344CC458225C7706FB" pageId="0" pageNumber="269" type="diagnosis">
<paragraph id="83CF41342531754C0510ABC6D3E02292" pageId="0" pageNumber="269">Diagnosis.</paragraph>
<paragraph id="340033F8A56F213CCEFBFCD67977092A" pageId="0" pageNumber="269">
Genus characterised by very long and sharp metatrochanters; dentate metafemora; elytra not reduced at tip; elytral disc bearing three setae and longitudinal grooves absent; 9th pore of the umbilicate series in the normal position (placed after the 8th); aedeagus with median lobe not very curved, size of basal bulb normal and parameres bearing two apical setae. Labial tooth present and smooth. Differs from
<taxonomicName id="02EC3F1E980C3B11C2F7BC2B373AD6A0" authorityName="Baehr" authorityYear="2017" class="Insecta" family="Carabidae" genus="Magnanillus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Magnanillus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="24F0E68E7A8FAA2047F2E27BD9C3733F" italics="true" pageId="0" pageNumber="269">Magnanillus</emphasis>
</taxonomicName>
Baehr, 2017 by the absence of longitudinal elytral grooves.
</paragraph>
</subSubSection>
<subSubSection id="8D3AC966F9395AA7FC80FBD8678B27C7" pageId="0" pageNumber="269" type="description">
<paragraph id="F24B6B6DA30AC767113DED8B00B5F838" pageId="0" pageNumber="269">Description.</paragraph>
<paragraph id="BF57EF663D9EB307DFCD6D8A66187370" pageId="0" pageNumber="269">Species of medium size (TL mm 1.37-2.18) and anophthalmous. Integument depigmented but well sclerified and covered with sparse pubescence.</paragraph>
<paragraph id="50F713F2B369A9F0B966864E187DF997" pageId="0" pageNumber="269">
<emphasis id="F7D1CF9E5422AFAF86DAB9EE7C7B77F2" bold="true" italics="true" pageId="0" pageNumber="269">Head</emphasis>
from normal to large size, slightly narrower than pronotum; mandibles short and simple, without hyperplasias. Maxillary palpi ovoidal and swollen. Labium transverse and articulated; mentum not fused with submentum. Labial tooth present and smooth. Antennae moniliform, without particular features.
</paragraph>
<paragraph id="E879C0535186D5E7189441D0A4CC9A8C" pageId="0" pageNumber="269">
<emphasis id="6369491FE1CC00B2A280465ECAB5FDC6" bold="true" italics="true" pageId="0" pageNumber="269">Pronotum</emphasis>
subquadrate, with sides not or only slightly sinuate at the basal third, smooth or denticulated before basal angles. Basal angles right, sharp, not rounded; basal border as wide as, or only slightly narrower, than the anterior border; presence of two marginal setae, the posterior one placed just before basal angles.
</paragraph>
<paragraph id="94FC7D77BD1E62A083DDAC3F10825ED2" pageId="0" pageNumber="269">
<emphasis id="AA37F836D28E21D088F6E90F331C10FA" bold="true" italics="true" pageId="0" pageNumber="269">Elytra</emphasis>
subrectangular and elongated, separately rounded and depressed, not truncate and not emarginated apically, with longitudinal groove absent. Elytral striae absent (except sutural stria). Lateral margin starting from humeral area, distinctly crenulate up to at least half-length. Scutellar pore present, large and umbilicate; umbilicate series of type B (sensu
<bibRefCitation id="80707F4F6100DC5F9583A5BCF61B66C3" author="Jeannel, R" journalOrPublisher="Memoires du Museum national d'Histoire naturelle, Paris (A)" pageId="0" pageNumber="269" pagination="33 - 204" refId="B33" refString="Jeannel, R, 1963. Monographie des &quot; Anillini &quot;, Bembidiides endoges (Coleoptera Trechidae). Memoires du Museum national d'Histoire naturelle, Paris (A) 28: 33 - 204" title="Monographie des &quot; Anillini &quot;, Bembidiides endoges (Coleoptera Trechidae)." volume="28" year="1963">Jeannel 1963</bibRefCitation>
;
<bibRefCitation id="4D3EDC00961E1A9B3BDCE544A2F04CF3" author="Giachino, PM" journalOrPublisher="Biodiversity Journal, monograph" pageId="0" pageNumber="269" pagination="1 - 112" refId="B16" refString="Giachino, PM, Vailati, D, 2011. Review of the Anillina of Greece (Coleoptera, Carabidae, Bembidiini). Biodiversity Journal, monograph 1: 1 - 112" title="Review of the Anillina of Greece (Coleoptera, Carabidae, Bembidiini)." volume="1" year="2011">Giachino and Vailati 2011</bibRefCitation>
); disc bearing three discal setae.
</paragraph>
<paragraph id="ACDB9C4406F4B9948426454FAFC948AE" pageId="0" pageNumber="269">
<emphasis id="44CB2F6DADAED18C0B7D5F50B69E0E56" bold="true" italics="true" pageId="0" pageNumber="269">Legs</emphasis>
relatively short and stumpy. Unarmed profemora; posterior edge of metafemora dentate, metatrochanters very long and sharp, two dilated protarsomeres in the male.
</paragraph>
<paragraph id="1B1DD55000C2822BC170C29ABC146BD4" pageId="0" pageNumber="269">
<emphasis id="610E996EBD37AA295EF6070B54875291" bold="true" italics="true" pageId="0" pageNumber="269">Aedeagus</emphasis>
relatively large, median lobe long and not very curved, with basal bulb of normal size. Parameres long and bearing two apical setae. Endophallus with poorly sclerified phanerae.
</paragraph>
</subSubSection>
<subSubSection id="8C229F38185E062B0ADADDF368433356" pageId="0" pageNumber="269" type="etymology">
<paragraph id="F048BFC709AED58E83CBCA301B32A4B3" pageId="0" pageNumber="269">Etymology.</paragraph>
<paragraph id="74E0623A5DE92D0EF3030EA812E89D93" pageId="0" pageNumber="269">
The genus name combines the Pilbara name with the suffix -
<emphasis id="A24EFB69DC073DF4A4B5999468909831" italics="true" pageId="0" pageNumber="269">phanus</emphasis>
to recognise the genus
<taxonomicName id="E37D6387FB220E62BFA585499DCFD5A0" class="Insecta" family="Carabidae" genus="Illaphanus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Illaphanus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="5D2D0EDC99D61280DDDF232368AD61EF" italics="true" pageId="0" pageNumber="269">Illaphanus</emphasis>
</taxonomicName>
. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="CB18E570CFBB58F31C98FE10FC19DDFE" pageId="0" pageNumber="269" type="species included">
<paragraph id="7D6234233F55AF4F001D9131B1657131" pageId="0" pageNumber="269">Species included.</paragraph>
<paragraph id="E148890B708A64D43A1F1EAEBA62FB45" pageId="0" pageNumber="269">Only two species currently belong to this genus:</paragraph>
<paragraph id="8398A2E5254727FB66E80C970EF59DE7" pageId="0" pageNumber="269">
<taxonomicName id="F44308F6672B335125E50E97BDE5CF29" authorityName="Giachino &amp; Eberhard &amp; Perina" authorityYear="2021" class="Insecta" family="Carabidae" genus="Pilbaraphanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilbaraphanus chichesterianus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="species" species="chichesterianus">
<emphasis id="DC2051ED518D0E649C9151E214CA0130" italics="true" pageId="0" pageNumber="269">Pilbaraphanus chichesterianus</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<paragraph id="FCC67636A18F08E41E7A5ABE2DC1F425" pageId="0" pageNumber="269">
<taxonomicName id="3F70A7C3373515F639DD7794BF21E626" authorityName="Giachino &amp; Eberhard &amp; Perina" authorityYear="2021" class="Insecta" family="Carabidae" genus="Pilbaraphanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilbaraphanus bilybarianus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="species" species="bilybarianus">
<emphasis id="1B1F46B16DFC6AA01C24CE45CAB56AEA" italics="true" pageId="0" pageNumber="269">Pilbaraphanus bilybarianus</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<caption id="5AC9DD27B350BA34E36C4A626F2E2BDC" doi="10.3897/zookeys.1044.58844.figures10-14" httpUri="https://binary.pensoft.net/fig/553763" pageId="0" pageNumber="269" start="Figures 1014" startId="F4">
<paragraph id="FA9002791D113B993C09179B96E27E17" pageId="0" pageNumber="269">
<emphasis id="40A317B8E35E92CC0D01A9736B3CF532" bold="true" pageId="0" pageNumber="269">Figures 10-14.</emphasis>
<taxonomicName id="91017E9DA9AC08CF57A29367C66CE368" authorityName="Giachino &amp; Eberhard &amp; Perina" authorityYear="2021" class="Insecta" family="Carabidae" genus="Pilbaraphanus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pilbaraphanus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="genus">
<emphasis id="A619175CADD78FD796DF650AFB678DE3" italics="true" pageId="0" pageNumber="269">Pilbaraphanus</emphasis>
</taxonomicName>
gen. et spp. nov., habitus (
<emphasis id="3A4C21AACCD1B58F845C1E5E88EA43B1" bold="true" pageId="0" pageNumber="269">10, 13</emphasis>
) right metafemur and metatrochanter in ventral view (
<emphasis id="22F86802A9E60AD5289188AF005D6758" bold="true" pageId="0" pageNumber="269">11, 14</emphasis>
) aedeagus in lateral view (
<emphasis id="822EB0E6C5584FEF9F829DD965B97A3D" bold="true" pageId="0" pageNumber="269">12</emphasis>
)
<emphasis id="A4AB4B51BCBE763407E363F008367388" bold="true" pageId="0" pageNumber="269">10-12</emphasis>
<taxonomicName id="735D273D24DBEC4C8265F44045892B7B" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. chichesterianus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="species" species="chichesterianus">
<emphasis id="139B7AF2A1F63114F711CFCA894FFD48" italics="true" pageId="0" pageNumber="269">P. chichesterianus</emphasis>
</taxonomicName>
sp. nov., HT ♂
<emphasis id="120ACD75159292178DBA2AA3A710765C" bold="true" pageId="0" pageNumber="269">13, 14</emphasis>
<taxonomicName id="3A4DA9B4B66F2EA47FC35F452AE18151" class="Insecta" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="P. bilybarianus" order="Coleoptera" pageId="0" pageNumber="269" phylum="Arthropoda" rank="species" species="bilybarianus">
<emphasis id="F6A5211BB1DB4526F502EF987E413B56" italics="true" pageId="0" pageNumber="269">P. bilybarianus</emphasis>
</taxonomicName>
sp. nov. HT ♀. Scale bars: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>