treatments-xml/data/03/F6/87/03F687EEFFC9FF97FF3CF49FAB8ECA8D.xml

<document ID-DOI="10.1093/zoolinnean/zlaa091" ID-GBIF-Dataset="f7e2e892-6f4e-45e2-b85d-174e93c49766" ID-ISSN="0024-4082" ID-Zenodo-Dep="5724151" ID-ZooBank="B7203900-1F11-40CD-B4D5-115BAD1B96FE" checkinTime="1619628871472" checkinUser="carolina" docAuthor="Royo-Torres, Rafael, Cobos, Alberto, Mocho, Pedro &amp; Alcalá, Luis" docDate="2021" docId="03F687EEFFC9FF97FF3CF49FAB8ECA8D" docLanguage="en" docName="ZoolJLinnSoc.191.201-227.pdf.imf" docOrigin="Zoological Journal of the Linnean Society 191" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.6:ZoolJLinnSoc.2017-.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-.journal_article" docStyleVersion="6" docTitle="Losillasaurus CASANOVAS ET AL. 2001" docType="treatment" docVersion="7" lastPageNumber="220" masterDocId="FFCFFF96FFDBFF84FFE5F109A843CC31" masterDocTitle="Origin and evolution of turiasaur dinosaurs set by means of a new ‘ rosetta’ specimen from Spain" masterLastPageNumber="227" masterPageNumber="201" pageNumber="219" updateTime="1638551938812" updateUser="ExternalLinkService">
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<mods:title>Origin and evolution of turiasaur dinosaurs set by means of a new ‘ rosetta’ specimen from Spain</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Royo-Torres, Rafael</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cobos, Alberto</mods:namePart>
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<mods:namePart>Mocho, Pedro</mods:namePart>
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<mods:namePart>Alcalá, Luis</mods:namePart>
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<mods:title>Zoological Journal of the Linnean Society</mods:title>
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<mods:date>2021</mods:date>
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<mods:number>2020-09-03</mods:number>
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<mods:number>191</mods:number>
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<mods:identifier type="DOI">10.1093/zoolinnean/zlaa091</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5724198" ID-GBIF-Taxon="190872627" ID-Zenodo-Dep="5724198" LSID="urn:lsid:plazi:treatment:03F687EEFFC9FF97FF3CF49FAB8ECA8D" httpUri="http://treatment.plazi.org/id/03F687EEFFC9FF97FF3CF49FAB8ECA8D" lastPageId="19" lastPageNumber="220" pageId="18" pageNumber="219">
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<paragraph blockId="18.[217,688,1430,1455]" box="[217,688,1430,1455]" pageId="18" pageNumber="219">
<heading box="[217,688,1430,1455]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="219" reason="2">
<taxonomicName authority="CASANOVAS ET AL., 2001" authorityName="CASANOVAS ET AL." authorityYear="2001" box="[217,688,1430,1455]" class="Reptilia" family="Eusauropoda" genus="Losillasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="219" phylum="Chordata" rank="genus">
<emphasis box="[217,401,1430,1453]" italics="true" pageId="18" pageNumber="219">LOSILLASAURUS</emphasis>
CASANOVAS 
<emphasis box="[548,612,1433,1453]" italics="true" pageId="18" pageNumber="219">ET AL</emphasis>
., 2001
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="18.[145,762,1502,1555]" pageId="18" pageNumber="219">
<emphasis box="[145,237,1503,1524]" italics="true" pageId="18" pageNumber="219">Figures:</emphasis>
See 
<bibRefCitation author="Casanovas ML &amp; Santafe JV &amp; Sanz JL" box="[294,540,1502,1524]" pageId="18" pageNumber="219" pagination="99 - 122" refId="ref14361" refString="Casanovas ML, Santafe JV, Sanz JL. 2001. Losillasaurus giganteus, un nuevo sauropodo del transito Jurasico - Cretacico de la cuenca de &quot; Los Serranos &quot; (Valencia, Espana). Paleontologia i Evolucio 32 - 33: 99 - 122." type="journal article" year="2001">
Casanovas 
<emphasis box="[421,475,1502,1524]" italics="true" pageId="18" pageNumber="219">et al.</emphasis>
, 2001
</bibRefCitation>
: figs 1–7; 
<figureCitation box="[647,755,1502,1524]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-10="Figure 11" captionStart-11="Figure 12" captionStart-12="Figure 13" captionStart-13="Figure 14" captionStart-14="Figure 15" captionStart-15="Figure 16" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStart-4="Figure 5" captionStart-5="Figure 6" captionStart-6="Figure 7" captionStart-7="Figure 8" captionStart-8="Figure 9" captionStart-9="Figure 10" captionStartId-0="1.[163,241,1693,1715]" captionStartId-1="2.[145,224,1652,1674]" captionStartId-10="13.[163,243,885,907]" captionStartId-11="14.[145,223,1646,1668]" captionStartId-12="15.[163,242,1011,1033]" captionStartId-13="16.[146,224,1002,1024]" captionStartId-14="17.[161,239,1001,1023]" captionStartId-15="18.[146,224,946,968]" captionStartId-2="5.[164,243,1514,1536]" captionStartId-3="6.[145,225,1139,1161]" captionStartId-4="7.[160,239,847,869]" captionStartId-5="8.[145,224,983,1005]" captionStartId-6="9.[162,242,1112,1134]" captionStartId-7="10.[145,225,931,953]" captionStartId-8="11.[163,244,1118,1140]" captionStartId-9="12.[145,223,1280,1302]" captionTargetBox-0="[262,1342,196,1651]" captionTargetBox-1="[148,1428,198,1608]" captionTargetBox-10="[166,1412,202,842]" captionTargetBox-11="[209,1358,210,1592]" captionTargetBox-12="[260,1345,202,970]" captionTargetBox-13="[151,1422,197,960]" captionTargetBox-14="[260,1345,200,946]" captionTargetBox-15="[154,1425,202,902]" captionTargetBox-2="[170,1443,199,1470]" captionTargetBox-3="[154,1428,198,1096]" captionTargetBox-4="[163,1446,195,808]" captionTargetBox-5="[148,1428,198,929]" captionTargetBox-6="[165,1443,197,1065]" captionTargetBox-7="[149,1429,199,888]" captionTargetBox-8="[165,1443,198,1076]" captionTargetBox-9="[151,1424,201,1235]" captionTargetId-0="figure-26@1.[257,1348,195,1653]" captionTargetId-1="figure-24@2.[145,1431,195,1612]" captionTargetId-10="figure-569@13.[163,1450,195,845]" captionTargetId-11="figure-113@14.[207,1362,195,1607]" captionTargetId-12="figure-512@15.[257,1348,195,972]" captionTargetId-13="figure-475@16.[145,1428,195,962]" captionTargetId-14="figure-443@17.[257,1348,195,961]" captionTargetId-15="figure-430@18.[145,1431,195,907]" captionTargetId-2="figure-136@5.[163,1450,195,1474]" captionTargetId-3="figure-323@6.[145,1431,195,1099]" captionTargetId-4="figure-543@7.[163,1446,195,808]" captionTargetId-5="figure-378@8.[145,1431,195,943]" captionTargetId-6="figure-359@9.[163,1446,195,1073]" captionTargetId-7="figure-424@10.[145,1432,195,891]" captionTargetId-8="figure-365@11.[163,1446,195,1079]" captionTargetId-9="figure-325@12.[145,1431,195,1241]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-10="13" captionTargetPageId-11="14" captionTargetPageId-12="15" captionTargetPageId-13="16" captionTargetPageId-14="17" captionTargetPageId-15="18" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionTargetPageId-9="12" captionText-0="Figure 1. Reconstructions of the skull of Turiasaurus (A) and Losillasaurus (B) in the lateral view. Left premaxilla, maxilla and teeth (MAP-6005) of Losillasaurus giganteus (San Lorenzo specimen) in labial or lateral (C) and lingual or medial (D) views. Right premaxillary tooth (MAP-6013) (E); and right maxillary teeth (MAP-6014) (F), (MAP-6015) (G), (MAP-6016) (H), (MAP-6017) (I), (MAP-6018) (J), (MAP-6019) (K) in lingual views." captionText-1="Figure 2. Left dentary and teeth (MAP-6008) of Losillasaurus giganteus (San Lorenzo specimen) in lingual view (A); left dentary teeth (MAP-6045) (d1), (MAP-6031) (d2), (MAP-6035) (d3), (MAP-6037) (d4), (MAP-6036) (d7), (MAP-6038) (d8), (MAP-6039) (d9), (MAP-6040) (d10), (MAP-6020) (d13), (MAP-6021) (d14) in lingual views. Right dentary and teeth (MAP- 6009) of Losillasaurus giganteus (San Lorenzo specimen) in lingual view (B). Right dentary teeth (MAP-6032) (d2) and (MAP-6033) (d3) of Losillasaurus giganteus (San Lorenzo specimen) in lingual view." captionText-10="Figure 11. Chevrons of Losillasaurus giganteus (San Lorenzo specimen) in posterior view: from anterior to posterior, chevron 1 to chevron 9 (MAP-6085 to MAP-66093), chevron 12 (MAP-6096), chevron 14 to 19 (MAP 6098 to MAP-6103), chevron 21 (MAP 6015) and chevron 22 (MAP-6016)." captionText-11="Figure 12. Left ulna (MAP-6111) of Losillasaurus giganteus (San Lorenzo specimen) in anterior (A), lateral (B), posterior (C), anteromedial (D), medial (E), anterolateral (F) and dorsal (G, H) views." captionText-12="Figure 13. Right femur (MAP-6113) of Losillasaurus giganteus (San Lorenzo specimen) in dorsal (A), ventral (B), medial (C), posterior (D), lateral (E) and anterior (F) views." captionText-13="Figure 14. Right tibia (MAP-6115) of Losillasaurus giganteus (San Lorenzo specimen) in dorsal (A), ventral (B), medial (C), posterior (D), anterior (E) and lateral (F) views." captionText-14="Figure 15. Right fibula (MAP-6116) of Losillasaurus giganteus (San Lorenzo specimen) in medial (A), dorsal (B), anterior (C), lateral (D) and posterior (E) views." captionText-15="Figure 16. Comparison between astragali. Left astragalus (CPT-1244) in dorsal (A) and posterior (D) views of Turiasaurus riodevensis (paratype) and right (B, E) (MAP-6119) and left (C, F) astragalus (MAP-6117) of Losillasaurus giganteus (San Lorenzo specimen) in dorsal (B, C) and posterior (E, F) views." captionText-2="Figure 3. Right premaxillary and maxillary teeth of Losillasaurus giganteus (San Lorenzo specimen): premaxillary teeth (MAP-6013) in lingual (A), labial (B), mesial (C) and apical (D) views; maxillary teeth (MAP-6014) in lingual (E), labial (F) and apical (G) views; maxillary teeth (MAP-6015) in lingual (H), labial (I), mesial (J) and apical (K) views; maxillary teeth (MAP-6016) in lingual (L), labial (M) and apical (N) views; maxillary teeth (MAP-6017) in lingual (O), labial (P), mesial (Q) and apical (R) views; maxillary teeth (MAP-6018) in labial (S), lingual (T), mesial (U) and apical (V) views; and maxillary teeth (MAP-6019) in lingual (W), labial (X), mesial (Y) and apical (Z) views." captionText-3="Figure 4. Dentary teeth of Losillasaurus giganteus (San Lorenzo specimen): right dentary teeth (MAP-6032) in lingual (A), labial (B), mesial (C), distal (D) and apical (E) views; left dentary teeth (MAP-6035) in lingual (F), labial (G), distal (H), mesial (I) and apical (E) views; left dentary teeth (MAP-6036) in lingual (K), labial (L), distal (M), mesial (N) and apical (O) views; left dentary teeth (MAP-6038) in lingual (P), labial (Q), distal (R), mesial (S) and apical (T) views; left dentary teeth (MAP-6039) in lingual (U), labial (V), distal (W) and apical (X) views; right dentary teeth (MAP-6034) in lingual (Y), labial (Z), distal (A’) and apical (B’) views; and left dentary teeth (MAP-6021) in labial (C’), lingual (D’), distal (E’) and apical (F’) views." captionText-4="Figure 5. Longitudinal grooves in the right premaxillary teeth (MAP-6013) in the labial view (A) and left dentary teeth (MAP-6035) in labial (B) and lingual (C) views of Losillasaurus giganteus (San Lorenzo specimen)." captionText-5="Figure 6. Comparison between Turiasaurus and Losillasaurus teeth: right premaxillary teeth (MAP-6013) in labial (A) and apical (B) views of Losillasaurus giganteus (San Lorenzo specimen); right maxillary teeth (MAP-6019) in labial (C) and apical (D) views of Losillasaurus giganteus (San Lorenzo specimen); left maxillary teeth (CPT-1262) in labial (E) and apical (F) views of Turiasaurus riodevensis; left maxillary teeth (CPT-3941) in labial (G) and apical (H) views of Turiasaurus riodevensis." captionText-6="Figure 7. Dorsal spines (MAP-6047) of Losillasaurus giganteus (San Lorenzo specimen) in anterior (A), dorsal (B) and posterior views. Dorsal spines (MCNV Lo-11) of Losillasaurus giganteus (paratype) in posterior view (D). Dorsal spines (MCNV Lo-17b) of Losillasaurus giganteus (paratype) in posterior view (E). Dorsal spine (CPT-2688) of Turiasaurus riodevensis (paratype) in posterior (F) and anterior (G) views. Dorsal spine (CPT-1633) of Turiasaurus riodevensis (referred to material) in posterior (H) and anterior (I) views." captionText-7="Figure 8. Caudal vertebrae in posterior views of Losillasaurus giganteus (San Lorenzo specimen): 4th (MAP-6050), 6th (MAP-6052), 7th (MAP-1846b), 8th (MAP-6053), 9th (MAP-6054), 10th (MAP-6055), 11th (MAP-6056), 12th (MAP-6057), 13th (MAP-6058), 14th (MAP-6059), 15th (MAP-6062), 16th (MAP-6061), 17th (MAP-6060), 18th (MAP-6063), 19th (MAP-6064), 20th (MAP-6066), 21st (MAP-6065), 23rd (MAP-6068), 24th (MAP-6069), 25th (MAP-6070), 26th (MAP-6071), 27th (MAP-6072), 28th (MAP-6073), 29th (MAP-6074), 30th (MAP-6075), 31st (MAP-6076), 33rd (MAP-6078), 34th (MAP-6079), 35th (MAP-6080) and 40th (MAP-6083)." captionText-8="Figure 9. Caudal vertebrae in lateral views of Losillasaurus giganteus (San Lorenzo specimen): 5th (MAP-1846a), 6th (MAP-6052), 7th (MAP-1846b), 8th (MAP-6053), 9th (MAP-6054), 10th (MAP-6055), 11th (MAP-6056), 12th (MAP-6057), 13th (MAP-6058), 14th (MAP-6059), 15th (MAP-6062), 16th (MAP-6061), 17th (MAP-6060), 18th (MAP-6063), 19th (MAP-6064), 20th (MAP-6066), 21st (MAP-6065), 23rd (MAP-6068), 24th (MAP-6069), 25th (MAP-6070), 26th (MAP-6071), 27th (MAP-6072), 28th (MAP-6073), 29th (MAP-6074), 30th (MAP-6075), 31st (MAP-6076), 33rd (MAP-6078), 34th (MAP-6079), 35th (MAP-6080) and 40th (MAP-6083)." captionText-9="Figure 10. Comparison of the caudal spines of Losillasaurus giganteus (CPT-1846, San Lorenzo specimen) in posterior (A), right lateral (B), left lateral (C), anterior (D) and dorsal (E) views and Turiasaurus riodevensis (CPT-1611, Puntal de Santa Cruz) in posterior (F), right lateral (G), anterior (H) and dorsal (I) views." figureDoi-0="http://doi.org/10.5281/zenodo.5724153" figureDoi-1="http://doi.org/10.5281/zenodo.5724155" figureDoi-10="http://doi.org/10.5281/zenodo.5724173" figureDoi-11="http://doi.org/10.5281/zenodo.5724175" figureDoi-12="http://doi.org/10.5281/zenodo.5724177" figureDoi-13="http://doi.org/10.5281/zenodo.5724179" figureDoi-14="http://doi.org/10.5281/zenodo.5724181" figureDoi-15="http://doi.org/10.5281/zenodo.5724183" figureDoi-2="http://doi.org/10.5281/zenodo.5724157" figureDoi-3="http://doi.org/10.5281/zenodo.5724159" figureDoi-4="http://doi.org/10.5281/zenodo.5724161" figureDoi-5="http://doi.org/10.5281/zenodo.5724163" figureDoi-6="http://doi.org/10.5281/zenodo.5724165" figureDoi-7="http://doi.org/10.5281/zenodo.5724167" figureDoi-8="http://doi.org/10.5281/zenodo.5724169" figureDoi-9="http://doi.org/10.5281/zenodo.5724171" httpUri-0="https://zenodo.org/record/5724153/files/figure.png" httpUri-1="https://zenodo.org/record/5724155/files/figure.png" httpUri-10="https://zenodo.org/record/5724173/files/figure.png" httpUri-11="https://zenodo.org/record/5724175/files/figure.png" httpUri-12="https://zenodo.org/record/5724177/files/figure.png" httpUri-13="https://zenodo.org/record/5724179/files/figure.png" httpUri-14="https://zenodo.org/record/5724181/files/figure.png" httpUri-15="https://zenodo.org/record/5724183/files/figure.png" httpUri-2="https://zenodo.org/record/5724157/files/figure.png" httpUri-3="https://zenodo.org/record/5724159/files/figure.png" httpUri-4="https://zenodo.org/record/5724161/files/figure.png" httpUri-5="https://zenodo.org/record/5724163/files/figure.png" httpUri-6="https://zenodo.org/record/5724165/files/figure.png" httpUri-7="https://zenodo.org/record/5724167/files/figure.png" httpUri-8="https://zenodo.org/record/5724169/files/figure.png" httpUri-9="https://zenodo.org/record/5724171/files/figure.png" pageId="18" pageNumber="219">Figs 1–16</figureCitation>
; Supporting Information, File S3.
</paragraph>
</subSubSection>
<subSubSection lastPageId="19" lastPageNumber="220" pageId="18" pageNumber="219" type="materials_examined">
<paragraph blockId="18.[145,760,1587,1640]" pageId="18" pageNumber="219">
<emphasis box="[145,302,1588,1609]" italics="true" pageId="18" pageNumber="219">Type species:</emphasis>
<taxonomicName authority="Casanovas et al. (2001)" authorityName="Casanovas" authorityYear="2001" class="Reptilia" genus="Losillasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="219" phylum="Chordata" rank="species" species="Giganteus">
<emphasis box="[325,620,1587,1609]" italics="true" pageId="18" pageNumber="219">Losillasaurus giganteus</emphasis>
<bibRefCitation author="Casanovas ML &amp; Santafe JV &amp; Sanz JL" pageId="18" pageNumber="219" pagination="99 - 122" refId="ref14361" refString="Casanovas ML, Santafe JV, Sanz JL. 2001. Losillasaurus giganteus, un nuevo sauropodo del transito Jurasico - Cretacico de la cuenca de &quot; Los Serranos &quot; (Valencia, Espana). Paleontologia i Evolucio 32 - 33: 99 - 122." type="journal article" year="2001">
Casanovas 
<emphasis box="[145,201,1618,1640]" italics="true" pageId="18" pageNumber="219">et al.</emphasis>
(2001)
</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph blockId="18.[145,760,1673,1726]" pageId="18" pageNumber="219">
<materialsCitation ID-GBIF-Occurrence="3412228302" collectionCode="MCNV" location="Museo de Ciencias Naturales" municipality="Anterior" pageId="18" pageNumber="219" specimenCount="1" stateProvince="Comunitat Valenciana" typeStatus="holotype">
<emphasis box="[145,259,1673,1694]" italics="true" pageId="18" pageNumber="219">
<typeStatus box="[145,253,1673,1694]" pageId="18" pageNumber="219">Holotype</typeStatus>
:
</emphasis>
<collectingMunicipality box="[272,376,1673,1695]" pageId="18" pageNumber="219">Anterior</collectingMunicipality>
caudal vertebra housed in the 
<location LSID="urn:lsid:plazi:treatment:03F687EEFFC9FF97FF3CF49FAB8ECA8D:8E806023FFC9FF96FF74F7A1A99DCA8C" box="[145,478,1703,1725]" municipality="Anterior" name="Museo de Ciencias Naturales" pageId="18" pageNumber="219" stateProvince="Comunitat Valenciana">Museo de Ciencias Naturales</location>
de 
<collectingRegion box="[517,614,1703,1725]" country="Spain" name="Comunitat Valenciana" pageId="18" pageNumber="219">Valencia</collectingRegion>
(
<collectionCode box="[628,712,1704,1726]" country="Italy" httpUri="http://grbio.org/cool/yhsn-iyjf" name="Museo Civico di Storia Naturale, Venice" pageId="18" pageNumber="219">MCNV</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="18.[145,762,1758,1811]" pageId="18" pageNumber="219">
<materialsCitation ID-GBIF-Occurrence="3412228301" collectionCode="MCNV" location="Museo de Ciencias Naturales" municipality="Two" pageId="18" pageNumber="219" specimenCount="1" stateProvince="Comunitat Valenciana" typeStatus="paratype">
<emphasis box="[145,256,1759,1780]" italics="true" pageId="18" pageNumber="219">
<typeStatus box="[145,250,1759,1780]" pageId="18" pageNumber="219">Paratype</typeStatus>
:
</emphasis>
<collectingMunicipality box="[276,325,1759,1780]" pageId="18" pageNumber="219">Two</collectingMunicipality>
anterior caudal vertebrae housed in the 
<location LSID="urn:lsid:plazi:treatment:03F687EEFFC9FF97FF3CF49FAB8ECA8D:8E806023FFC9FF96FF59F7F4AA44CB22" box="[188,519,1789,1811]" municipality="Two" name="Museo de Ciencias Naturales" pageId="18" pageNumber="219" stateProvince="Comunitat Valenciana">Museo de Ciencias Naturales</location>
de 
<collectingRegion box="[556,653,1789,1811]" country="Spain" name="Comunitat Valenciana" pageId="18" pageNumber="219">Valencia</collectingRegion>
(
<collectionCode box="[667,751,1789,1811]" country="Italy" httpUri="http://grbio.org/cool/yhsn-iyjf" name="Museo Civico di Storia Naturale, Venice" pageId="18" pageNumber="219">MCNV</collectionCode>
)
</materialsCitation>
.
</paragraph>
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<emphasis box="[145,443,1841,1862]" italics="true" pageId="18" pageNumber="219">Type locality and horizon:</emphasis>
La 
<collectingCountry box="[492,583,1841,1863]" name="Canada" pageId="18" pageNumber="219">Cañada</collectingCountry>
site (Valencia), V i l l a r d e l A r z o b i s p o Fo r m a t i o n (C a s a n o v a s 
<emphasis box="[809,874,1071,1093]" italics="true" pageId="18" pageNumber="219">et al.</emphasis>
, 2001; 
<bibRefCitation author="Royo-Torres R &amp; Cobos A &amp; Alcala L" box="[973,1282,1071,1093]" pageId="18" pageNumber="219" pagination="1925 - 1927" refId="ref16034" refString="Royo-Torres R, Cobos A, Alcala L. 2006. A giant European dinosaur and a new sauropod clade. Science 314: 1925 - 1927." type="journal article" year="2006">
Royo-Torres 
<emphasis box="[1137,1203,1071,1093]" italics="true" pageId="18" pageNumber="219">et al.</emphasis>
, 2006
</bibRefCitation>
). Dated as Kimmeridgian according to the data of 
<bibRefCitation author="Campos-Soto S &amp; Benito MI &amp; Cobos A &amp; Caus E &amp; Quijada IE &amp; Suarez-Gonzalez P &amp; Mas R &amp; Royo-Torres R &amp; Alcala L" pageId="18" pageNumber="219" pagination="471 - 510" refId="ref14227" refString="Campos-Soto S, Benito MI, Cobos A, Caus E, Quijada IE, Suarez-Gonzalez P, Mas R, Royo-Torres R, Alcala L. 2019. Revisiting the age and palaeoenvironments of the Upper Jurassic - Lower Cretaceous? Dinosaur-bearing sedimentary record of eastern Spain: implications for Iberian palaeogeography. Journal of Iberian Geology 45: 471 - 510." type="journal article" year="2019">
Campos-Soto 
<emphasis box="[809,867,1133,1154]" italics="true" pageId="18" pageNumber="219">et al.</emphasis>
(2019)
</bibRefCitation>
.
</paragraph>
<paragraph blockId="18.[809,1426,1195,1677]" pageId="18" pageNumber="219">
<emphasis box="[809,1063,1195,1217]" italics="true" pageId="18" pageNumber="219">Referred material:</emphasis>
The material referred to 
<taxonomicName box="[809,974,1226,1247]" class="Reptilia" genus="Losillasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="219" phylum="Chordata" rank="genus">
<emphasis box="[809,974,1226,1247]" italics="true" pageId="18" pageNumber="219">Losillasaurus</emphasis>
</taxonomicName>
comes from the same specimen from where the 
<typeStatus box="[940,1043,1257,1279]" pageId="18" pageNumber="219">holotype</typeStatus>
and 
<typeStatus box="[1107,1215,1257,1278]" pageId="18" pageNumber="219">paratype</typeStatus>
were selected. It consists of a skull fragment and partial postcranial skeleton (see Supporting Information, File S1). A second specimen, described in this paper, is from the San Lorenzo site (Riodeva, Teruel, 
<collectingCountry box="[1159,1226,1379,1401]" name="Spain" pageId="18" pageNumber="219">Spain</collectingCountry>
) in the Villar del Arzobispo Formation. It consists of a partial skull with teeth and partial postcranial skeleton (see the previous description section and Supporting Information, File S1). A third specimen referred to as 
<taxonomicName box="[1237,1397,1502,1523]" class="Reptilia" genus="Losillasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="219" phylum="Chordata" rank="genus">
<emphasis box="[1237,1397,1502,1523]" italics="true" pageId="18" pageNumber="219">Losillasaurus</emphasis>
</taxonomicName>
is a complete anterior caudal vertebra (
<collectionCode box="[1238,1295,1533,1554]" pageId="18" pageNumber="219">SHN</collectionCode>
180) found in Baleal (Peniche municipality, 
<collectingCountry box="[1176,1275,1563,1585]" name="Portugal" pageId="18" pageNumber="219">Portugal</collectingCountry>
) in the Praia de Amoreira-Porto Novo Formation, dated as Upper Kimmeridgian–Lower Tithonian (
<bibRefCitation author="Manuppella G &amp; Antunes MT &amp; Pais J &amp; Ramalho MM &amp; Rey J" pageId="18" pageNumber="219" refId="ref15397" refString="Manuppella G, Antunes MT, Pais J, Ramalho MM, Rey J. 1999. Noticia explicativa da Folha 30 A (Lourinha). Lisboa: Departamento de Geologia do Instituto Geologico e Mineiro, 83." type="book" year="1999">
Manuppella 
<emphasis box="[1359,1418,1625,1646]" italics="true" pageId="18" pageNumber="219">et al.</emphasis>
, 1999
</bibRefCitation>
; 
<bibRefCitation author="Mocho P &amp; Royo-Torres R &amp; Malafaia E &amp; Escaso F &amp; Ortega F" box="[876,1094,1655,1677]" pageId="18" pageNumber="219" pagination="23 - 36" refId="ref15689" refString="Mocho P, Royo-Torres R, Malafaia E, Escaso F, Ortega F. 2017 b. First occurrences of non-neosauropod eusauropod procoelous caudal vertebrae in the Portuguese Upper Jurassic record. Geobios 50: 23 - 36." type="journal article" year="2017">
Mocho 
<emphasis box="[957,1013,1655,1677]" italics="true" pageId="18" pageNumber="219">et al.</emphasis>
, 2017b
</bibRefCitation>
).
</paragraph>
<paragraph blockId="18.[809,1425,1718,1893]" lastBlockId="19.[827,1442,1394,1447]" lastPageId="19" lastPageNumber="220" pageId="18" pageNumber="219">
<emphasis box="[809,1222,1718,1739]" italics="true" pageId="18" pageNumber="219">
Revised diagnosis: 
<taxonomicName box="[1053,1222,1718,1739]" class="Reptilia" genus="Losillasaurus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="18" pageNumber="219" phylum="Chordata" rank="genus">Losillasaurus</taxonomicName>
</emphasis>
is diagnosed by eight autapomorphies: (1) (new) lateral surface of the premaxilla and maxilla with dorsoventrally elongated grooves convergent with diplodocids and 
<taxonomicName authority="(Mannion et al., 2019)" baseAuthorityName="Mannion" baseAuthorityYear="2019" box="[809,1278,1841,1863]" class="Reptilia" family="Diplodocidae" genus="Nemegtosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Saurischia" pageId="18" pageNumber="219" phylum="Chordata" rank="genus">
<emphasis box="[809,995,1841,1862]" italics="true" pageId="18" pageNumber="219">Nemegtosaurus</emphasis>
(
<bibRefCitation author="Mannion PD &amp; Upchurch P &amp; Schhwarz D &amp; Wings O" box="[1014,1267,1841,1863]" pageId="18" pageNumber="219" pagination="784 - 909" refId="ref15347" refString="Mannion PD, Upchurch P, Schhwarz D, Wings O. 2019. Taxonomic affinities of the putative titanosaurs from the Late Jurassic Tendaguru Formation of Tanzania: phylogenetic and biogeographic implications for eusauropod dinosaur evolution. Zoological Journal of the Linnean Society 185: 784 - 909." type="journal article" year="2019">
Mannion 
<emphasis box="[1131,1193,1841,1862]" italics="true" pageId="18" pageNumber="219">et al.</emphasis>
, 2019
</bibRefCitation>
)
</taxonomicName>
; (2) (new) a maxillary tooth with a secondary apex on the distal edge; (3) markedly curved neural spines of the proximal caudal vertebrae that in the first and second caudal vertebrae produce a pronounced cutlass-like shape in the lateral view (
<bibRefCitation author="Casanovas ML &amp; Santafe JV &amp; Sanz JL" box="[393,652,1486,1508]" pageId="19" pageNumber="220" pagination="99 - 122" refId="ref14361" refString="Casanovas ML, Santafe JV, Sanz JL. 2001. Losillasaurus giganteus, un nuevo sauropodo del transito Jurasico - Cretacico de la cuenca de &quot; Los Serranos &quot; (Valencia, Espana). Paleontologia i Evolucio 32 - 33: 99 - 122." type="journal article" year="2001">
Casanovas 
<emphasis box="[525,582,1486,1508]" italics="true" pageId="19" pageNumber="220">et al.</emphasis>
, 2001
</bibRefCitation>
; 
<bibRefCitation author="Upchurch P &amp; Barrett PM &amp; Dodson P" pageId="19" pageNumber="220" pagination="259 - 324" refId="ref16615" refString="Upchurch P, Barrett PM, Dodson P. 2004 a. Sauropoda. In: Weishampel DB, Dodson P, Osmolska H, eds. The Dinosauria, 2 nd edn. Berkeley: University of California Press, 259 - 324." type="book chapter" year="2004">
Upchurch 
<emphasis box="[163,224,1517,1538]" italics="true" pageId="19" pageNumber="220">et al.</emphasis>
, 2004a
</bibRefCitation>
); (4) (new) presence of a dorsoventral ridge in the anterolateral surface of the spine at least between the fourth and tenthcaudal vertebrae (it is not clearly present in the first three caudal vertebrae and disappears in the 11 
<superScript attach="left" box="[562,577,1638,1650]" fontSize="5" pageId="19" pageNumber="220">th</superScript>
caudal vertebra); (5) (new) caudal neural spines with a shallow dorsal groove with directed anteroposteriorly, bigger in the anteriormost caudal vertebrae, especially in the first and second bones and shallow from the third to the 30 
<superScript attach="left" box="[191,206,1792,1804]" fontSize="5" pageId="19" pageNumber="220">th</superScript>
caudal vertebrae; (6) (new) the long-axis of the obturator foramen is perpendicular to the long-axis of the pubis; (7) (new) the lateral trochanter of the fibula is concave and rugose; and (8) (new) the tibial condyle of the femur is twice as large as the fibula condyle.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5724185" ID-Zenodo-Dep="5724185" httpUri="https://zenodo.org/record/5724185/files/figure.png" pageId="19" pageNumber="220" startId="19.[165,243,1181,1203]" targetBox="[166,1443,202,1139]" targetPageId="19">
<paragraph blockId="19.[163,1443,1181,1350]" pageId="19" pageNumber="220">
<emphasis bold="true" box="[165,280,1181,1203]" pageId="19" pageNumber="220">Figure 17.</emphasis>
<emphasis box="[289,552,1181,1203]" italics="true" pageId="19" pageNumber="220">Narindasaurus thevenini</emphasis>
: right premaxillary-maxillary teeth (MNHN MAJ 423) in mesial (A), distal (B), lingual (C), labial (D) and apical (E) views; right tibia (MNHN MAJ 428) in medial view (F); left pubis (MNHN MAJ 430) in lateral view (G); anterior caudal vertebrae (MNHN MAJ 424) in anterior (H), right lateral (I), ventral (J) and left lateral (K) views; posterior caudal vertebrae (MNHN MAJ 426) in posterior (L), right lateral (M) and dorsal (N) views; distal chevron (MNHN MAJ 429) in right lateral view (O); middle-anterior chevron (MNHN MAJ 425) in dorsal (P), right lateral (Q), posterior (R) and anterior (S) views; and right fibula in lateral (T) and anterior (U) views from the same specimen.
</paragraph>
</caption>
<paragraph blockId="19.[827,1443,1487,1724]" pageId="19" pageNumber="220">
<emphasis box="[827,1079,1487,1509]" italics="true" pageId="19" pageNumber="220">Additional comments:</emphasis>
The character, anteroposterior length at the base of the neural spines of the proximal caudal vertebrae being approximately half the height of the spine (ratio = 0.5), was included in the original diagnosis (
<bibRefCitation author="Casanovas ML &amp; Santafe JV &amp; Sanz JL" box="[959,1232,1610,1632]" pageId="19" pageNumber="220" pagination="99 - 122" refId="ref14361" refString="Casanovas ML, Santafe JV, Sanz JL. 2001. Losillasaurus giganteus, un nuevo sauropodo del transito Jurasico - Cretacico de la cuenca de &quot; Los Serranos &quot; (Valencia, Espana). Paleontologia i Evolucio 32 - 33: 99 - 122." type="journal article" year="2001">
Casanovas 
<emphasis box="[1097,1159,1610,1632]" italics="true" pageId="19" pageNumber="220">et al.</emphasis>
, 2001
</bibRefCitation>
) and accepted in 
<bibRefCitation author="Upchurch P &amp; Barrett PM &amp; Dodson P" box="[827,1087,1641,1663]" pageId="19" pageNumber="220" pagination="259 - 324" refId="ref16615" refString="Upchurch P, Barrett PM, Dodson P. 2004 a. Sauropoda. In: Weishampel DB, Dodson P, Osmolska H, eds. The Dinosauria, 2 nd edn. Berkeley: University of California Press, 259 - 324." type="book chapter" year="2004">
Upchurch 
<emphasis box="[943,997,1641,1662]" italics="true" pageId="19" pageNumber="220">et al.</emphasis>
(2004a)
</bibRefCitation>
. However, this is not considered valid as the spines are compressed by taphonomic deformation.
</paragraph>
</subSubSection>
</treatment>
</document>