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<document id="A018899B673A3C92045E52C878FBDD80" ID-CLB-Dataset="5777" ID-DOI="10.1206/669.1" ID-GBIF-Dataset="724cd8a5-13f2-4484-b409-9c77061ce444" ID-ISSN="0003-0082" ID-Zenodo-Dep="4713076" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1619136733944" checkinUser="felipe" docAuthor="Dimitrov, Dimitar, Álvarez-Padilla, Fernando &amp; Hormiga, Gustavo" docDate="2010" docId="03F687C8593B227874A8A9E6FCA2FB57" docLanguage="en" docName="AmMusNovit.2010.3683.1-19.pdf" docOrigin="American Museum Novitates 3683" docSource="http://www.bioone.org/doi/abs/10.1206/669.1" docStyle="DocumentStyle:70DDF1BDF6846024993F05262BAE3001.3:AmMusNovit.2000-2010.journal_article.type1" docStyleId="70DDF1BDF6846024993F05262BAE3001" docStyleName="AmMusNovit.2000-2010.journal_article.type1" docStyleVersion="3" docTitle="Dolichognatha quinquemucronata Dimitrov &amp; Álvarez-Padilla &amp; Hormiga 2010, new combination" docType="treatment" docVersion="5" lastPageNumber="15" masterDocId="FFCFFFB0593122767432AC24FFCFFFA9" masterDocTitle="On the Phylogenetic Placement of the Spider Genus Atimiosa Simon, 1895, and the Circumscription of Dolichognatha O. P. - Cambridge, 1869 (Tetragnathidae, Araneae)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="11" updateTime="1698937526220" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="CC2F40220B8DAB4B5CB9F06474B199DB">On the Phylogenetic Placement of the Spider Genus Atimiosa Simon, 1895, and the Circumscription of Dolichognatha O. P. - Cambridge, 1869 (Tetragnathidae, Araneae)</mods:title>
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<paragraph id="8BE036DE593B227C74A8A9E6FDC4FA5E" blockId="10.[154,626,1474,1527]" pageId="10" pageNumber="11">
<taxonomicName id="4C5F4D5D593B227C74A8A9E6FE8CFA5E" ID-CoL="3757Q" authority="(Simon, 1895)" authorityName="Dimitrov &amp; Álvarez-Padilla &amp; Hormiga" authorityYear="2010" baseAuthorityName="Simon" baseAuthorityYear="1895" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="quinquemucronata" status="comb. nov.">
<emphasis id="B92BEACC593B227C74A8A9E6FDDFFA73" box="[154,528,1474,1498]" italics="true" pageId="10" pageNumber="11">Dolichognatha quinquemucronata</emphasis>
(Simon, 1895)
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,
<emphasis id="B92BEACC593B227C7562A9FBFDC4FA5E" bold="true" box="[336,523,1503,1527]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="A21857B7593B227C7562A9FBFDC4FA5E" box="[336,523,1503,1527]" pageId="10" pageNumber="11" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</paragraph>
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<paragraph id="8BE036DE593B227C7505AA34FE1AF981" blockId="10.[311,469,1552,1576]" box="[311,469,1552,1576]" pageId="10" pageNumber="11">
<heading id="D0A881B2593B227C7505AA34FE1AF981" box="[311,469,1552,1576]" centered="true" fontSize="36" level="3" pageId="10" pageNumber="11" reason="7">
<figureCitation id="13642A5B593B227C7505AA34FE1AF981" box="[311,469,1552,1576]" captionStart="Fig" captionStartId="2.[151,189,966,988]" captionTargetBox="[131,1224,210,932]" captionTargetId="figure-197@2.[131,1224,210,933]" captionTargetPageId="2" captionText="Fig. 1. Dolichognatha quinquemucronata (Simon, 1895), new combination. Subadult female holotype from Sri Lanka (MNHM 16140): ventral (A), dorsal (B). Cephalothorax: dorsal (C), frontal (D). Abdomen and genital area, ventral (E)." figureDoi="http://doi.org/10.5281/zenodo.4713080" httpUri="https://zenodo.org/record/4713080/files/figure.png" pageId="10" pageNumber="11">Figures 1AE</figureCitation>
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<subSubSection id="C3456555593B227C74B2AA6BFDE0F93A" pageId="10" pageNumber="11" type="reference_group">
<paragraph id="8BE036DE593B227C74B2AA6BFF27F9D3" blockId="10.[128,654,1615,1683]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB4F11F0593B227C74B2AA6BFF27F9D3" pageId="10" pageNumber="11">
<treatmentCitation id="0AFE10CF593B227C74B2AA6BFDF5F9CB" author="Simon, E." box="[128,570,1615,1634]" page="742" pageId="10" pageNumber="11" year="1894">
<taxonomicName id="4C5F4D5D593B227C74B2AA6BFDF5F9CB" authority="Simon, 1894: 742" authorityName="Simon" authorityPageNumber="742" authorityYear="1894" box="[128,570,1615,1634]" class="Arachnida" family="Tetragnathidae" genus="Atimiosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="quinquemucronata">
<emphasis id="B92BEACC593B227C74B2AA6BFEB0F9CB" box="[128,383,1615,1634]" italics="true" pageId="10" pageNumber="11">Atimiosa quinquemucronata</emphasis>
<bibRefCitation id="EFCE4B2F593B227C75BCAA6BFDF5F9CB" author="Simon, E." box="[398,570,1615,1634]" pageId="10" pageNumber="11" pagination="489 - 760" refId="ref6217" refString="Simon, E. 1894. Histoire naturelle des araignees 1: 489 - 760. Paris: Roret." type="journal article" year="1894">Simon, 1894: 742</bibRefCitation>
</taxonomicName>
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(nomen nudum).
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</paragraph>
<paragraph id="8BE036DE593B227C74B2AAA4FDE0F93A" blockId="10.[128,654,1615,1683]" box="[128,559,1664,1683]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB4F11F0593B227C74B2AAA4FDE0F93A" box="[128,559,1664,1683]" pageId="10" pageNumber="11">
<treatmentCitation id="0AFE10CF593B227C74B2AAA4FDE4F93A" author="Simon, E." box="[128,555,1664,1683]" page="154" pageId="10" pageNumber="11" year="1895">
<taxonomicName id="4C5F4D5D593B227C74B2AAA4FDE4F93A" authority="Simon, 1895 a: 154" authorityName="Simon" authorityPageNumber="154" authorityYear="1895" box="[128,555,1664,1683]" class="Arachnida" family="Tetragnathidae" genus="Atimiosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="quinquemucronata">
<emphasis id="B92BEACC593B227C74B2AAA4FEB7F93A" box="[128,376,1664,1683]" italics="true" pageId="10" pageNumber="11">Atimiosa quinquemucronata</emphasis>
<bibRefCitation id="EFCE4B2F593B227C754DAAA4FDE4F93A" author="Simon, E." box="[383,555,1664,1683]" pageId="10" pageNumber="11" pagination="131 - 160" refId="ref6237" refString="Simon, E. 1895 a. Etudes arachnologiques. 26 e. XLI. Descriptions d'especes et de genres nouveaux de l'ordre des Araneae. Annales de la Societe Entomologique de France 64: 131 - 160." type="journal article" year="1895">Simon, 1895a: 154</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
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<subSubSection id="C3456555593B227C74A8AA81FCF5FB4C" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BE036DE593B227C74A8AA81FD42F914" blockId="10.[128,653,1701,1754]" box="[154,653,1701,1725]" pageId="10" pageNumber="11">
TYPE: Subadult female
<typeStatus id="54E4887C593B227C758EAA81FDEFF914" box="[444,544,1701,1725]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
from
<collectingCountry id="F348764E593B227C765FAA81FD42F914" box="[621,653,1701,1725]" name="Sri Lanka" pageId="10" pageNumber="11">Sri</collectingCountry>
</paragraph>
<paragraph id="8BE036DE593B227C74B2AAE6FCF5FB4C" blockId="10.[128,653,1701,1754]" lastBlockId="10.[701,1228,1170,1754]" pageId="10" pageNumber="11">
Lanka. The only information on the label, other than the collection number and the species name, is Nuw. El. (
<collectionCode id="ED4EAE1B593B227C7028A88BFBB8FB6E" box="[1050,1143,1199,1223]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="10" pageNumber="11">MNHN</collectionCode>
16140, examined).
</paragraph>
</subSubSection>
<subSubSection id="C3456555593B227C76EAA8CFFCA9F973" pageId="10" pageNumber="11" type="discussion">
<paragraph id="8BE036DE593B227C76EAA8CFFCA9F973" blockId="10.[701,1228,1170,1754]" pageId="10" pageNumber="11">
JUSTIFICATION OF THE TRANSFER: The
<typeStatus id="54E4887C593B227C70A9A8CFFB04FAAA" box="[1179,1227,1259,1283]" pageId="10" pageNumber="11">type</typeStatus>
specimen, which is also the only known specimen of this species, is a subadult female (fig. 1A, E), thus it is impossible to study its genital morphology. However, it has all somatic characters diagnostic of
<taxonomicName id="4C5F4D5D593B227C705EA95BFCC7FA1D" authorityName="O.Pickard-Cambridge" authorityYear="1869" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEACC593B227C705EA95BFCC7FA1D" italics="true" pageId="10" pageNumber="11">Dolichognatha</emphasis>
</taxonomicName>
, such as: PME smaller than the PLE, PLE, and ALE separated, chelicerae slender and elongated, and femur IV without trichobothria. Even with 92.5% missing data in the data matrix, the cladistic analyses unambiguously placed this species in a group together with the
<taxonomicName id="4C5F4D5D593B227C77ADAA68FB8EF9CD" authorityName="O.Pickard-Cambridge" authorityYear="1869" box="[927,1089,1612,1636]" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEACC593B227C77ADAA68FB8EF9CD" box="[927,1089,1612,1636]" italics="true" pageId="10" pageNumber="11">Dolichognatha</emphasis>
</taxonomicName>
species included in our analyses and with
<taxonomicName id="4C5F4D5D593B227C701BAA4EFB0BF92B" authorityName="Dimitrov &amp; Álvarez-Padilla &amp; Hormiga" authorityYear="2010" baseAuthorityName="Schmidt and Krause" baseAuthorityYear="1993" box="[1065,1220,1642,1666]" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="comorensis">
<emphasis id="B92BEACC593B227C701BAA4EFB0BF92B" box="[1065,1220,1642,1666]" italics="true" pageId="10" pageNumber="11">A. comorensis</emphasis>
</taxonomicName>
. Based on this cladistic hypothesis (fig. 9), we transfer
<taxonomicName id="4C5F4D5D593B227C7715AA81FBD2F914" authorityName="Simon" authorityYear="1895" box="[807,1053,1701,1725]" class="Arachnida" family="Tetragnathidae" genus="Atimiosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="quinquemucronata">
<emphasis id="B92BEACC593B227C7715AA81FBD2F914" box="[807,1053,1701,1725]" italics="true" pageId="10" pageNumber="11">A. quinquemucronata</emphasis>
</taxonomicName>
to the genus
<taxonomicName id="4C5F4D5D593B227C768FAAE6FC90F973" authorityName="O.Pickard-Cambridge" authorityYear="1869" box="[701,863,1730,1754]" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEACC593B227C768FAAE6FC90F973" box="[701,863,1730,1754]" italics="true" pageId="10" pageNumber="11">Dolichognatha</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF206656593A227D7442AA28FD1BF9D8" ID-DOI="http://doi.org/10.5281/zenodo.4713146" ID-Zenodo-Dep="4713146" httpUri="https://zenodo.org/record/4713146/files/figure.png" pageId="11" pageNumber="12" startId="11.[112,150,1548,1570]" targetBox="[90,1185,210,1513]" targetPageId="11">
<paragraph id="8BE036DE593A227D7442AA28FD1BF9D8" blockId="11.[88,1188,1548,1649]" pageId="11" pageNumber="12">
Fig. 9. Strict consensus of the six MPTs found by TNT under the equal weights parsimony criterion analyzing the complete data set (L
<emphasis id="B92BEACC593A227D75F2AA02FE1BF995" bold="true" box="[448,468,1574,1596]" pageId="11" pageNumber="12">5</emphasis>
1059, CI
<emphasis id="B92BEACC593A227D760BAA02FD82F995" bold="true" box="[569,589,1574,1596]" pageId="11" pageNumber="12">5</emphasis>
0.25, RI
<emphasis id="B92BEACC593A227D769DAA02FD0CF995" bold="true" box="[687,707,1574,1596]" pageId="11" pageNumber="12">5</emphasis>
0.60). Numbers cutting the branches represent Bremer support; over the branches, standard bootstrap/jackknife; below the branches, Poisson bootstrap/ symmetric resampling.
<taxonomicName id="4C5F4D5D593A227D7575AA78FE17F9D8" authorityName="O.Pickard-Cambridge" authorityYear="1869" box="[327,472,1628,1649]" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEACC593A227D7575AA78FE17F9D8" box="[327,472,1628,1649]" italics="true" pageId="11" pageNumber="12">Dolichognatha</emphasis>
</taxonomicName>
is shown in a gray box.
</paragraph>
</caption>
<caption id="DF206656593D227A74A5AA2CFC72F9C4" ID-DOI="http://doi.org/10.5281/zenodo.4713150" ID-Zenodo-Dep="4713150" httpUri="https://zenodo.org/record/4713150/files/figure.png" pageId="12" pageNumber="13" startId="12.[151,189,1544,1566]" targetBox="[145,1211,210,1510]" targetPageId="12">
<paragraph id="8BE036DE593D227A74A5AA2CFC72F9C4" blockId="12.[128,1228,1544,1645]" pageId="12" pageNumber="13">
Fig. 10. Strict consensus of the six MPTs found by TNT under the equal weights parsimony criterion analyzing the matrix without
<taxonomicName id="4C5F4D5D593D227A7582AA06FD4AF99E" authorityName="Simon" authorityYear="1895" box="[432,645,1570,1591]" class="Arachnida" family="Tetragnathidae" genus="Atimiosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="quinquemucronata">
<emphasis id="B92BEACC593D227A7582AA06FD4AF99E" box="[432,645,1570,1591]" italics="true" pageId="12" pageNumber="13">A. quinquemucronata</emphasis>
</taxonomicName>
(L
<emphasis id="B92BEACC593D227A769EAA06FD0FF991" bold="true" box="[684,704,1570,1592]" pageId="12" pageNumber="13">5</emphasis>
1058, CI
<emphasis id="B92BEACC593D227A7714AA06FCF5F991" bold="true" box="[806,826,1570,1592]" pageId="12" pageNumber="13">5</emphasis>
0.25, RI
<emphasis id="B92BEACC593D227A77AEAA06FC7FF991" bold="true" box="[924,944,1570,1592]" pageId="12" pageNumber="13">5</emphasis>
0.60). Numbers cutting the branches represent Bremer support; over the branches, standard bootstrap/jackknife; below the branches, Poisson bootstrap/symmetric resampling.
<taxonomicName id="4C5F4D5D593D227A761DAA7CFD0FF9C4" authorityName="O.Pickard-Cambridge" authorityYear="1869" box="[559,704,1624,1645]" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEACC593D227A761DAA7CFD0FF9C4" box="[559,704,1624,1645]" italics="true" pageId="12" pageNumber="13">Dolichognatha</emphasis>
</taxonomicName>
is shown in a gray box.
</paragraph>
</caption>
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<paragraph id="8BE036DE593C227B7442AA25FE92F9E4" blockId="13.[88,1187,1537,1613]" pageId="13" pageNumber="14">
Fig. 11. Alternative topologies (above and at right, on opposite page) obtained analyzing the data under the implied weighting criterion using different values of the concavity constant
<emphasis id="B92BEACC593C227B77BAAA38FC5AF998" box="[904,917,1564,1585]" italics="true" pageId="13" pageNumber="14">k</emphasis>
(
<taxonomicName id="4C5F4D5D593C227B7796AA38FBB6F998" authorityName="Simon" authorityYear="1895" box="[932,1145,1564,1585]" class="Arachnida" family="Tetragnathidae" genus="Atimiosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="quinquemucronata">
<emphasis id="B92BEACC593C227B7796AA38FBB6F998" box="[932,1145,1564,1585]" italics="true" pageId="13" pageNumber="14">A. quinquemucronata</emphasis>
</taxonomicName>
not included in the analyses).
</paragraph>
</caption>
<subSubSection id="C3456555593F2278760CAF40FD42FCD3" box="[574,653,868,890]" pageId="14" pageNumber="15" type="description">
<paragraph id="8BE036DE593F2278760CAF40FD42FCD3" blockId="14.[574,653,868,890]" box="[574,653,868,890]" pageId="14" pageNumber="15">
<figureCitation id="13642A5B593F2278760CAF40FD42FCD3" box="[574,653,868,890]" captionStart="Fig" captionStartId="13.[112,150,1537,1559]" captionTargetBox="[267,1008,210,1504]" captionTargetId="figure-0@13.[267,1008,210,1504]" captionTargetPageId="13" captionText="Fig. 11. Alternative topologies (above and at right, on opposite page) obtained analyzing the data under the implied weighting criterion using different values of the concavity constant k (A. quinquemucronata not included in the analyses)." figureDoi="http://doi.org/10.5281/zenodo.4713160" httpUri="https://zenodo.org/record/4713160/files/figure.png" pageId="14" pageNumber="15">Fig. 11.</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3456555593F227874A8AF99FF17FAD8" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BE036DE593F227874A8AF99FF17FAD8" blockId="14.[128,654,957,1393]" pageId="14" pageNumber="15">
COMMENTS: There are three other species of
<taxonomicName id="4C5F4D5D593F227874ACAFFEFE8FFC5B" authorityName="O.Pickard-Cambridge" authorityYear="1869" box="[158,320,986,1010]" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEACC593F227874ACAFFEFE8FFC5B" box="[158,320,986,1010]" italics="true" pageId="14" pageNumber="15">Dolichognatha</emphasis>
</taxonomicName>
besides
<taxonomicName id="4C5F4D5D593F227875ADAFFEFD42FC5B" authorityName="Simon" authorityYear="1895" box="[415,653,986,1010]" class="Arachnida" family="Tetragnathidae" genus="Atimiosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="quinquemucronata">
<emphasis id="B92BEACC593F227875ADAFFEFD42FC5B" box="[415,653,986,1010]" italics="true" pageId="14" pageNumber="15">D. quinquemucronata</emphasis>
</taxonomicName>
described from
<collectingCountry id="F348764E593F22787504AFDCFE62FBB9" box="[310,429,1016,1040]" name="Sri Lanka" pageId="14" pageNumber="15">Sri Lanka</collectingCountry>
:
<taxonomicName id="4C5F4D5D593F22787589AFDCFDAFFBB9" baseAuthorityName="Simon" baseAuthorityYear="1895" box="[443,608,1016,1040]" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="incanescens">
<emphasis id="B92BEACC593F22787589AFDCFDAFFBB9" box="[443,608,1016,1040]" italics="true" pageId="14" pageNumber="15">D. incanescens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5F4D5D593F22787640AFDCFF1CFB84" authorityName="O.Pickard-Cambridge" authorityYear="1869" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="nietneri">
<emphasis id="B92BEACC593F22787640AFDCFF1CFB84" italics="true" pageId="14" pageNumber="15">D. nietneri</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C5F4D5D593F22787527A831FEB2FB84" baseAuthorityName="Simon" baseAuthorityYear="1895" box="[277,381,1045,1069]" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="albida">
<emphasis id="B92BEACC593F22787527A831FEB2FB84" box="[277,381,1045,1069]" italics="true" pageId="14" pageNumber="15">D. albida</emphasis>
</taxonomicName>
. This makes a reconciliation of the subadult specimen of
<taxonomicName id="4C5F4D5D593F2278763EA817FF39FBC1" authorityName="Simon" authorityYear="1895" class="Arachnida" family="Tetragnathidae" genus="Atimiosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="quinquemucronata">
<emphasis id="B92BEACC593F2278763EA817FF39FBC1" italics="true" pageId="14" pageNumber="15">D. quinquemucronata</emphasis>
</taxonomicName>
a very difficult task. However, the presence of characteristic abdominal tubercles in
<taxonomicName id="4C5F4D5D593F22787494A8AFFE52FB0A" authorityName="Simon" authorityYear="1895" box="[166,413,1163,1187]" class="Arachnida" family="Tetragnathidae" genus="Atimiosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="quinquemucronata">
<emphasis id="B92BEACC593F22787494A8AFFE52FB0A" box="[166,413,1163,1187]" italics="true" pageId="14" pageNumber="15">D. quinquemucronata</emphasis>
</taxonomicName>
, which differ from those of the other three Sri Lankan species, and its eye pattern, may prove useful if adults with these traits are collected. At present, as the
<typeStatus id="54E4887C593F22787486A924FED7FAB1" box="[180,280,1280,1304]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C5F4D5D593F22787561A925FD84FAB0" authorityName="Simon" authorityYear="1895" box="[339,587,1281,1305]" class="Arachnida" family="Tetragnathidae" genus="Atimiosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="quinquemucronata">
<emphasis id="B92BEACC593F22787561A925FD84FAB0" box="[339,587,1281,1305]" italics="true" pageId="14" pageNumber="15">D. quinquemucronata</emphasis>
</taxonomicName>
is a subadult specimen that cannot be identified, we suggest that it should be treated as nomen nudum.
</paragraph>
</subSubSection>
<subSubSection id="C3456555593F22787491A9BBFE64F9FE" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8BE036DE593F22787491A9BBFDF3FA7D" blockId="14.[163,619,1439,1492]" pageId="14" pageNumber="15">
<taxonomicName id="4C5F4D5D593F22787491A9BBFEBBFA7D" ID-CoL="37572" authority="(Schmidt and Krause, 1993)" authorityName="Dimitrov &amp; Álvarez-Padilla &amp; Hormiga" authorityYear="2010" baseAuthorityName="Schmidt and Krause" baseAuthorityYear="1993" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="comorensis" status="comb. nov.">
<emphasis id="B92BEACC593F22787491A9BBFE08FA1E" box="[163,455,1439,1463]" italics="true" pageId="14" pageNumber="15">Dolichognatha comorensis</emphasis>
(
<bibRefCitation id="EFCE4B2F593F227875EAA9BBFEA3FA7D" author="Schmidt, G. &amp; R. H. Krause" pageId="14" pageNumber="15" pagination="4 - 9" refId="ref6124" refString="Schmidt, G., and R. H. Krause. 1993. Spinnen von den Komoren III: Tetragnathinae und Metinae (Araneida: Araneidae). Teil I. Arachnologisches Magazin 1 (10): 4 - 9." type="journal article" year="1993">Schmidt and Krause, 1993</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B92BEACC593F227875B3A998FDF3FA7D" bold="true" box="[385,572,1468,1492]" pageId="14" pageNumber="15">
<taxonomicNameLabel id="A21857B7593F227875B3A998FDF3FA7D" box="[385,572,1468,1492]" pageId="14" pageNumber="15" rank="species">new combination</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BE036DE593F2278753CA9C9FE30F9AC" blockId="14.[270,511,1517,1541]" box="[270,511,1517,1541]" pageId="14" pageNumber="15">
<heading id="D0A881B2593F2278753CA9C9FE30F9AC" box="[270,511,1517,1541]" centered="true" fontSize="36" level="3" pageId="14" pageNumber="15" reason="7">
<figureCitation id="13642A5B593F2278753CA9C9FE7FF9AC" box="[270,432,1517,1541]" captionStart="Fig" captionStartId="5.[112,150,1569,1591]" captionTargetBox="[100,1174,210,1534]" captionTargetId="figure-1@5.[100,1174,210,1535]" captionTargetPageId="5" captionText="Fig. 2. Dolichognatha comorensis (Schmidt and Krause, 1993), new combination. Male palp: prolateral" figureDoi="http://doi.org/10.5281/zenodo.4713084" httpUri="https://zenodo.org/record/4713084/files/figure.png" pageId="14" pageNumber="15">Figures 2AE</figureCitation>
;
<figureCitation id="13642A5B593F2278758FA9C9FE30F9AC" box="[445,511,1517,1541]" captionStart="Fig" captionStartId="6.[151,189,716,738]" captionTargetBox="[131,1224,210,681]" captionTargetId="figure-427@6.[131,1224,210,682]" captionTargetPageId="6" captionText="Fig. 3. Dolichognatha comorensis (Schmidt and Krause, 1993), new combination holotype from Grande Comoro (RMCA 160.649). Male: ventral (A), lateral (B), frontal (C)." figureDoi="http://doi.org/10.5281/zenodo.4713090" httpUri="https://zenodo.org/record/4713090/files/figure.png" pageId="14" pageNumber="15">3AC</figureCitation>
</heading>
</paragraph>
<paragraph id="8BE036DE593F227874B2AA09FE64F9FE" blockId="14.[128,654,1580,1623]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB4F11F0593F227874B2AA09FE64F9FE" pageId="14" pageNumber="15">
<treatmentCitation id="0AFE10CF593F227874B2AA09FD46F996" author="Schmidt, G. &amp; R. H. Krause" box="[128,649,1580,1600]" page="6" pageId="14" pageNumber="15" year="1993">
<taxonomicName id="4C5F4D5D593F227874B2AA09FD46F996" authority="Schmidt and Krause, 1993: 6" authorityName="Schmidt and Krause" authorityPageNumber="6" authorityYear="1993" box="[128,649,1580,1600]" class="Arachnida" family="Tetragnathidae" genus="Atimiosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="comorensis">
<emphasis id="B92BEACC593F227874B2AA09FE8EF9E9" box="[128,321,1581,1600]" italics="true" pageId="14" pageNumber="15">Atimiosa comorensis</emphasis>
<bibRefCitation id="EFCE4B2F593F22787561AA08FD46F996" author="Schmidt, G. &amp; R. H. Krause" box="[339,649,1580,1599]" pageId="14" pageNumber="15" pagination="4 - 9" refId="ref6124" refString="Schmidt, G., and R. H. Krause. 1993. Spinnen von den Komoren III: Tetragnathinae und Metinae (Araneida: Araneidae). Teil I. Arachnologisches Magazin 1 (10): 4 - 9." type="journal article" year="1993">Schmidt and Krause, 1993: 6</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AFE10CF593F227874A8AA60FE68F9FE" author="Schmidt, G. &amp; R. H. Krause" box="[154,423,1604,1623]" page="5" pageId="14" pageNumber="15" year="1994">
<bibRefCitation id="EFCE4B2F593F227874A8AA60FE68F9FE" author="Schmidt, G. &amp; R. H. Krause" box="[154,423,1604,1623]" pageId="14" pageNumber="15" pagination="3 - 25" refId="ref6167" refString="Schmidt, G., and R. H. Krause. 1994. Spinnen von den Komoren 3: Tetragnathinae und Metinae (Araneida: Araneidae). Arachnologisches Magazin 2 (Sonderausgabe 1): 3 - 25 [reprint of Schmidt and Krause, 1993]." type="journal article" year="1994">Schmidt and Krause, 1994: 5</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3456555593F227874A8AA4EFCDDFCD0" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BE036DE593F227874A8AA4EFEABF973" blockId="14.[128,654,1642,1754]" pageId="14" pageNumber="15">
<materialsCitation id="3B373C83593F227874A8AA4EFEABF973" ID-GBIF-Occurrence="3313391302" collectingDate="1983-11-27" collectionCode="RMCA" collectorName="R. Jocque" country="Comoros" county="Boboni" elevation="600" location="Islands" municipality="Grande Comoro" pageId="14" pageNumber="15" specimenCode="RMCA 160.649" specimenCount="1" typeStatus="holotype">
TYPE:
<typeStatus id="54E4887C593F227874C6AA4EFE90F92B" box="[244,351,1642,1666]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
male from
<collectingCountry id="F348764E593F22787616AA4EFD41F92B" box="[548,654,1642,1666]" name="Comoros" pageId="14" pageNumber="15">Comoros</collectingCountry>
<location id="8E806005593F227874B2AAA3FF1CF936" LSID="urn:lsid:plazi:treatment:03F687C8593B227874A8A9E6FCA2FB57:8E806005593F227874B2AAA3FF1CF936" box="[128,211,1671,1695]" country="Comoros" county="Boboni" municipality="Grande Comoro" name="Islands" pageId="14" pageNumber="15">Islands</location>
,
<collectingMunicipality id="6B84ACA4593F227874D2AAA3FE6DF936" box="[224,418,1671,1695]" pageId="14" pageNumber="15">Grande Comoro</collectingMunicipality>
,
<collectingCounty id="62814E52593F22787582AAA3FDC8F936" box="[432,519,1671,1695]" pageId="14" pageNumber="15">Boboni</collectingCounty>
,
<date id="FFE1101E593F22787626AAA3FD46F936" box="[532,649,1671,1695]" pageId="14" pageNumber="15" value="1983-11-27">
<collectingDate id="EFA5E9F6593F22787626AAA3FD46F936" box="[532,649,1671,1695]" pageId="14" pageNumber="15" value="1983-11-27">27.xi.1983</collectingDate>
</date>
,
<elevation id="0072D1ED593F227874B2AA80FEF2F915" box="[128,317,1700,1724]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="15" unit="m" value="600.0">
elevation
<quantity id="4CA79B3B593F227874CFAA80FEF2F915" box="[253,317,1700,1724]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="15" unit="m" value="600.0">600m</quantity>
</elevation>
,
<collectorName id="26AA5308593F2278756AAA80FE15F915" box="[344,474,1700,1724]" pageId="14" pageNumber="15">R. Jocqué</collectorName>
leg. (
<specimenCode id="DBF99EA5593F22787604AA80FF15F973" collectionCode="RMCA" country="Belgium" httpUri="http://grbio.org/cool/ee3x-fpgk" name="Royal Museum for Central Africa" pageId="14" pageNumber="15" type="Museum">RMCA 160.649</specimenCode>
, examined).
</materialsCitation>
</paragraph>
<paragraph id="8BE036DE593F22787697AF40FCDDFCD0" blockId="14.[677,786,868,889]" box="[677,786,868,889]" pageId="14" pageNumber="15">
<emphasis id="B92BEACC593F22787697AF40FCDDFCD0" box="[677,786,868,889]" italics="true" pageId="14" pageNumber="15">Continued.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C3456555593F227876EAAF99FCA2FB57" pageId="14" pageNumber="15" type="discussion">
<paragraph id="8BE036DE593F227876EAAF99FCA2FB57" blockId="14.[701,1228,957,1278]" pageId="14" pageNumber="15">
JUSTIFICATION FOR THE TRANSFER: Morphology of the male palp of this species (presence of CBEP and metine embolic apophysis; shape of conductor and embolus) as well as eye size and arrangement are consistent with placement in the genus
<taxonomicName id="4C5F4D5D593F2278777CA876FC23FBC3" authorityName="O.Pickard-Cambridge" authorityYear="1869" box="[846,1004,1106,1130]" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEACC593F2278777CA876FC23FBC3" box="[846,1004,1106,1130]" italics="true" pageId="14" pageNumber="15">Dolichognatha</emphasis>
</taxonomicName>
. Transfer is further supported by the results from our cladistic analyses: parsimony analyses under equal and implied weights always, except when
<emphasis id="B92BEACC593F2278704BA88FFB48FB6A" box="[1145,1159,1195,1219]" italics="true" pageId="14" pageNumber="15">k</emphasis>
<emphasis id="B92BEACC593F227870A6A88EFB65FB6B" bold="true" box="[1172,1194,1194,1218]" pageId="14" pageNumber="15">#</emphasis>
3, found
<taxonomicName id="4C5F4D5D593F2278773AA8ECFC6DFB49" authorityName="Schmidt and Krause" authorityYear="1993" box="[776,930,1224,1248]" class="Arachnida" family="Tetragnathidae" genus="Atimiosa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="comorensis">
<emphasis id="B92BEACC593F2278773AA8ECFC6DFB49" box="[776,930,1224,1248]" italics="true" pageId="14" pageNumber="15">A. comorensis</emphasis>
</taxonomicName>
nested within
<taxonomicName id="4C5F4D5D593F22787075A8ECFD2FFB57" authorityName="O.Pickard-Cambridge" authorityYear="1869" class="Arachnida" family="Tetragnathidae" genus="Dolichognatha" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEACC593F22787075A8ECFD2FFB57" italics="true" pageId="14" pageNumber="15">Dolichognatha</emphasis>
</taxonomicName>
(figs. 9, 10).
</paragraph>
</subSubSection>
</treatment>
</document>