179 lines
20 KiB
XML
179 lines
20 KiB
XML
<document id="8E47BE4F858EE3967EBD4E91E7F84C86" ID-DOI="10.5281/zenodo.5170203" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170203" ID-ZooBank="AB1A57F0-7CB4-4830-920B-DF219740A596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628281492031" checkinUser="felipe" docAuthor="Blahnik, Roger J. & Holzenthal, Ralph W." docDate="2017" docId="03F687A7FFB9F845FF01BDA6455DFCEF" docLanguage="en" docName="InsectaMundi.0602.1-251.pdf" docOrigin="Insecta Mundi 2017 (602)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Mortoniella (Mortoniella) akrogeneios Blahnik & Holzenthal 2017, new species" docType="treatment" docVersion="4" lastPageNumber="65" masterDocId="FFCFFFDFFFF8F807FFC1B865400BFFB4" masterDocTitle="Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *" masterLastPageNumber="251" masterPageNumber="1" pageNumber="64" updateTime="1699039969677" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="7FF9AA16270B412FCB6D09DBB46425DE">
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<mods:title id="1499DEAE6F7AC3853AE01ABA1CF16CE9">Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *</mods:title>
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</mods:titleInfo>
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<mods:name id="6F9C94C2F08407CE2DE8177BA7A04ADC" type="personal">
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<mods:namePart id="772553860D3113CDE6EA6F999B77F096">Blahnik, Roger J.</mods:namePart>
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<mods:name id="B52C5F05CD0FA0DBC609CC9F0B12974D" type="personal">
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<mods:roleTerm id="95DFEFFCFD4F45E1E5A3B44351629D40">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="ABD2CE1E9901F490FBBD4704EBD5DB4F">Holzenthal, Ralph W.</mods:namePart>
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<mods:typeOfResource id="599F4CA2821BC7AE5F34FCE5E6AA3A9D">text</mods:typeOfResource>
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<mods:title id="8192EE472E647FF8E84322746AE69296">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="CBDDB1A63E5B310462F8960A9116FB94">
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<mods:date id="C326353AEC9DE84EC4EF6CA321CC1EDD">2017</mods:date>
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<mods:number id="3F1A82A21AEB8324F8D6D79C0B57ADED">2017-12-29</mods:number>
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</mods:detail>
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<mods:detail id="A55B2025A0A9B7B81D40D441774BFC77" type="volume">
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<mods:number id="07D2CBA59A805783F86B7C1B24F3AD94">2017</mods:number>
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<mods:detail id="D26B2BD43134B9554BF16EBEB6519D5B" type="issue">
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<mods:number id="8EEA04DD12570936B9E44EC67164E114">602</mods:number>
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<mods:classification id="385A5EF777AFD763F79F1A38738CE72F">journal article</mods:classification>
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<mods:identifier id="09601005E896FC25616725DFB52DA821" type="DOI">10.5281/zenodo.5170203</mods:identifier>
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<mods:identifier id="2B3386E1A8BE75BE5061534288F99606" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="D233CF836DD751C79C0B8593C4B49842" type="Zenodo-Dep">5170203</mods:identifier>
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<mods:identifier id="259F79823B5F5A0BDAA366958051EF01" type="ZooBank">AB1A57F0-7CB4-4830-920B-DF219740A596</mods:identifier>
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<treatment id="03F687A7FFB9F845FF01BDA6455DFCEF" ID-DOI="http://doi.org/10.5281/zenodo.6487940" ID-Zenodo-Dep="6487940" LSID="urn:lsid:plazi:treatment:03F687A7FFB9F845FF01BDA6455DFCEF" httpUri="http://treatment.plazi.org/id/03F687A7FFB9F845FF01BDA6455DFCEF" lastPageId="66" lastPageNumber="65" pageId="65" pageNumber="64">
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<subSubSection id="C345653AFFB9F846FF01BDA6438CFA6F" box="[192,903,1475,1499]" pageId="65" pageNumber="64" type="nomenclature">
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<paragraph id="8BE036B1FFB9F846FF01BDA6438CFA6F" blockId="65.[192,903,1475,1531]" box="[192,903,1475,1499]" pageId="65" pageNumber="64">
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<heading id="D0A881DDFFB9F846FF01BDA6438CFA6F" bold="true" box="[192,903,1475,1499]" fontSize="10" level="2" pageId="65" pageNumber="64" reason="3">
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<emphasis id="B92BEAA3FFB9F846FF01BDA6438CFA6F" bold="true" box="[192,903,1475,1499]" pageId="65" pageNumber="64">
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<taxonomicName id="4C5F4D32FFB9F846FF01BDA642DAFA6F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[192,721,1475,1499]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="65" pageNumber="64" phylum="Tracheophyta" rank="species" species="akrogeneios" status="sp. nov." subGenus="Mortoniella">
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<emphasis id="B92BEAA3FFB9F846FF01BDA64168FA6F" bold="true" box="[192,355,1475,1499]" italics="true" pageId="65" pageNumber="64">Mortoniella</emphasis>
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(
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<emphasis id="B92BEAA3FFB9F846FEB7BDA64212FA6F" bold="true" box="[374,537,1475,1499]" italics="true" pageId="65" pageNumber="64">Mortoniella</emphasis>
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)
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<emphasis id="B92BEAA3FFB9F846FDEABDA642DAFA6F" bold="true" box="[555,721,1475,1499]" italics="true" pageId="65" pageNumber="64">akrogeneios</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFB9F846FD23BDA6438CFA6F" box="[738,903,1475,1499]" pageId="65" pageNumber="64" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C345653AFFB9F845FF01BD8645ABFD0F" lastPageId="66" lastPageNumber="65" pageId="65" pageNumber="64" type="description">
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<paragraph id="8BE036B1FFB9F846FF01BD86411DFA4F" blockId="65.[192,903,1475,1531]" box="[192,278,1507,1531]" pageId="65" pageNumber="64">
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<figureCitation id="13642A34FFB9F846FF01BD86411DFA4F" box="[192,278,1507,1531]" captionStart="Figure 48" captionStartId="188.[192,276,1836,1858]" captionTargetBox="[343,1303,233,1761]" captionTargetId="figure-57@188.[343,1304,233,1761]" captionTargetPageId="188" captionText="Figure 48. Mortoniella (Mortoniella) akrogeneios, new species, male genitalia. A—lateral; B—segment IX and tergum X, dorsal; C—phallic ensemble, ventral; D—dorsal phallic spine, dorsal; E—ventral process of segment VI, lateral." figureDoi="http://doi.org/10.5281/zenodo.5170303" httpUri="https://zenodo.org/record/5170303/files/figure.png" pageId="65" pageNumber="64">Fig. 48</figureCitation>
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</paragraph>
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<paragraph id="8BE036B1FFB9F846FF31BE4644FAF94F" blockId="65.[192,1440,1571,1787]" pageId="65" pageNumber="64">
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<taxonomicName id="4C5F4D32FFB9F846FF31BE464205F98F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[240,526,1571,1595]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="65" pageNumber="64" phylum="Tracheophyta" rank="species" species="akrogeneios" status="sp. nov.">
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<emphasis id="B92BEAA3FFB9F846FF31BE464205F98F" box="[240,526,1571,1595]" italics="true" pageId="65" pageNumber="64">Mortoniella akrogeneios</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFB9F846FDDCBE414255F98F" box="[541,606,1572,1595]" pageId="65" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
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is superficially most similar to
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<taxonomicName id="4C5F4D32FFB9F846FC1DBE4144A4F98F" authority="(Flint)" baseAuthorityName="Flint" baseAuthorityYear="1981" box="[988,1199,1571,1595]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="65" pageNumber="64" phylum="Tracheophyta" rank="species" species="limona">
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<emphasis id="B92BEAA3FFB9F846FC1DBE414453F98F" box="[988,1112,1571,1595]" italics="true" pageId="65" pageNumber="64">M. limona</emphasis>
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(Flint)
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</taxonomicName>
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, due to the shape of the apex of the dorsal phallic spine, which has a very pronounced rounded ventral projection in both species. However, the ventral projection of
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<taxonomicName id="4C5F4D32FFB9F846FD21BE014391F9CF" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[736,922,1635,1659]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="65" pageNumber="64" phylum="Tracheophyta" rank="species" species="akrogeneios">
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<emphasis id="B92BEAA3FFB9F846FD21BE014391F9CF" box="[736,922,1635,1659]" italics="true" pageId="65" pageNumber="64">M. akrogeneios</emphasis>
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</taxonomicName>
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is narrower (and thus may appear more prominent). Additional diagnostic characters distinguishing it from
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<taxonomicName id="4C5F4D32FFB9F846FC13BEE14442F92F" baseAuthorityName="Flint" baseAuthorityYear="1981" box="[978,1097,1667,1691]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="65" pageNumber="64" phylum="Tracheophyta" rank="species" species="limona">
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<emphasis id="B92BEAA3FFB9F846FC13BEE14442F92F" box="[978,1097,1667,1691]" italics="true" pageId="65" pageNumber="64">M. limona</emphasis>
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</taxonomicName>
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include its apically projecting inferior appendages, short paramere appendages, and shallower emargination of tergum X. Also,
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<taxonomicName id="4C5F4D32FFB9F846FA41BEC14147F96F" authorityName="Blahnik & Holzenthal" authorityYear="2017" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="65" pageNumber="64" phylum="Tracheophyta" rank="species" species="akrogeneios">
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<emphasis id="B92BEAA3FFB9F846FA41BEC14147F96F" italics="true" pageId="65" pageNumber="64">M. akrogeneios</emphasis>
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</taxonomicName>
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has only small rounded, ventral projections on the phallicata, not nearly as projecting as those in
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<taxonomicName id="4C5F4D32FFB9F846FF1EBE814150F94F" baseAuthorityName="Flint" baseAuthorityYear="1981" box="[223,347,1763,1787]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="65" pageNumber="64" phylum="Tracheophyta" rank="species" species="limona">
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<emphasis id="B92BEAA3FFB9F846FF1EBE814150F94F" box="[223,347,1763,1787]" italics="true" pageId="65" pageNumber="64">M. limona</emphasis>
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</taxonomicName>
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and its two closely related species,
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<taxonomicName id="4C5F4D32FFB9F846FCC1BE81438BF94F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[768,896,1763,1787]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="65" pageNumber="64" phylum="Tracheophyta" rank="species" species="gracilis" status="sp. nov.">
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<emphasis id="B92BEAA3FFB9F846FCC1BE81438BF94F" box="[768,896,1763,1787]" italics="true" pageId="65" pageNumber="64">M. gracilis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFB9F846FC4FBE8143C5F94F" box="[910,974,1764,1787]" pageId="65" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
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and
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<taxonomicName id="4C5F4D32FFB9F846FBC8BE8144A8F94F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[1033,1187,1763,1787]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="65" pageNumber="64" phylum="Tracheophyta" rank="species" species="variabilis" status="sp. nov.">
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<emphasis id="B92BEAA3FFB9F846FBC8BE8144A8F94F" box="[1033,1187,1763,1787]" italics="true" pageId="65" pageNumber="64">M. variabilis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFB9F846FB70BE8144FAF94F" box="[1201,1265,1764,1787]" pageId="65" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8BE036B1FFB9F846FF31BF46412BF8CF" blockId="65.[192,1440,1827,1915]" pageId="65" pageNumber="64">
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<emphasis id="B92BEAA3FFB9F846FF31BF464133F88F" box="[240,312,1827,1851]" italics="true" pageId="65" pageNumber="64">Adult</emphasis>
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—Length of forewing: male 3.0-
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<quantity id="4CA79B54FFB9F846FD10BF46433FF88F" box="[721,820,1827,1851]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="65" pageNumber="64" unit="mm" value="3.1">3.1 mm</quantity>
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. Forewing with forks I, II, and III present, hind wing with forks II and III. Spur formula 0:4:4. Overall color (in alcohol) light brown. Wing bar not evident
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</paragraph>
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<paragraph id="8BE036B1FFBAF845FF31B8A645ABFD0F" blockId="66.[192,1441,195,859]" pageId="66" pageNumber="65">
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<emphasis id="B92BEAA3FFBAF845FF31B8A641AFFF6F" box="[240,420,195,219]" italics="true" pageId="66" pageNumber="65">Male genitalia</emphasis>
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—Ventral process of segment VI laterally compressed, ventrally projecting, large, subtriangular, length about equal to width at base, process not retracted anterobasally. Segment IX nearly evenly rounded anterolaterally, length greatest midlaterally, posterolateral margin weakly rounded dorsally, narrowing ventrally; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, separated dorsomesally by much less than ½ width of segment. Tergum X short, lateral margins subparallel, apical margin concave, apicolateral lobes very weakly projecting; apicomesal lobes visible in lateral view, rounded apically; ventrolateral lobes distinct, irregular, projecting. Inferior appendages without dorsolateral projections, apicoventral projections paired, projecting, upturned and acute apically. Mesal pockets of inferior appendage with apical processes short, posterodorsally curved. Paramere appendage short (much shorter than dorsal phallic spine) and relatively thick, apex acute. Dorsal phallic spine, as viewed laterally, with pronounced compressed apical expansion, upturned and acute dorsally, enlarged and rounded ventrally, dorsal and ventral projections connected almost linearly on posterior margin; spine, in dorsal view, very slightly widened in middle, apex acute. Phallicata with dorsal margin somewhat elevated and sclerotized, ventrally with pair of small rounded and sclerotized basal projections. Endophallic membrane short, with 2 pairs of enlarged spines (possibly phallotremal complex), 1 pair extending straight, the other strongly curved; ventromesal spine absent.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C345653AFFBAF845FF31BAA6455DFCEF" pageId="66" pageNumber="65" type="materials_examined">
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||
<paragraph id="8BE036B1FFBAF845FF31BAA642E4FD4F" blockId="66.[192,1441,195,859]" pageId="66" pageNumber="65">
|
||
<materialsCitation id="3B373CECFFBAF845FF31BAA642E4FD4F" collectingDate="1976-02-04" collectionCode="N, NMNH" country="Ecuador" location="Spangler" municipality="Puyo" pageId="66" pageNumber="65" specimenCode="UMSP000097078" specimenCount="1" stateProvince="Pastaza" typeStatus="holotype">
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||
<emphasis id="B92BEAA3FFBAF845FF31BAA641BFFD6F" bold="true" box="[240,436,707,731]" italics="true" pageId="66" pageNumber="65">
|
||
<typeStatus id="54E48813FFBAF845FF31BAA64162FD6F" box="[240,361,707,731]" pageId="66" pageNumber="65" type="holotype">Holotype</typeStatus>
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||
male
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</emphasis>
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(alcohol)—
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<emphasis id="B92BEAA3FFBAF845FDFEBAA64352FD6F" bold="true" box="[575,857,707,731]" pageId="66" pageNumber="65">
|
||
<collectingCountry id="F3487621FFBAF845FDFEBAA642D3FD6F" box="[575,728,707,731]" name="Ecuador" pageId="66" pageNumber="65">ECUADOR</collectingCountry>
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||
:
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||
<collectingRegion id="499BF853FFBAF845FD25BAA64358FD6F" box="[740,851,707,731]" country="Ecuador" name="Pastaza" pageId="66" pageNumber="65">Pastaza</collectingRegion>
|
||
:
|
||
</emphasis>
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||
<collectingMunicipality id="6B84ACCBFFBAF845FCA1BAA14395FD68" box="[864,926,708,732]" pageId="66" pageNumber="65">Puyo</collectingMunicipality>
|
||
(
|
||
<quantity id="4CA79B54FFBAF845FC6CBAA643F1FD6F" box="[941,1018,707,731]" metricMagnitude="4" metricUnit="m" metricValue="2.7" pageId="66" pageNumber="65" unit="km" value="27.0">27 km</quantity>
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||
<collectionCode id="ED4EAE74FFBAF845FBC0BAA64412FD6F" box="[1025,1049,707,731]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="66" pageNumber="65" type="Herbarium">N</collectionCode>
|
||
), Est. Fluvia Metrica,
|
||
<date id="FFE11071FFBAF845FAEFBAA64597FD6F" box="[1326,1436,707,731]" pageId="66" pageNumber="65" value="1976-02-04">
|
||
<collectingDate id="EFA5E999FFBAF845FAEFBAA64597FD6F" box="[1326,1436,707,731]" pageId="66" pageNumber="65" value="1976-02-04">4.ii.1976</collectingDate>
|
||
</date>
|
||
,
|
||
<location id="8E80606AFFBAF845FF01BA864138FD4F" LSID="urn:lsid:plazi:treatment:03F687A7FFB9F845FF01BDA6455DFCEF:8E80606AFFBAF845FF01BA864138FD4F" box="[192,307,739,763]" country="Ecuador" municipality="Puyo" name="Spangler" pageId="66" pageNumber="65" stateProvince="Pastaza">Spangler</location>
|
||
, et al. (
|
||
<specimenCode id="DBF99ECAFFBAF845FE4EBA864261FD4F" box="[399,618,739,763]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="66" pageNumber="65" type="University or college">UMSP000097078</specimenCode>
|
||
) (
|
||
<collectionCode id="ED4EAE74FFBAF845FD43BA8642E8FD4F" box="[642,739,739,763]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="66" pageNumber="65" type="Museum">NMNH</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BE036B1FFBAF845FF31BB664514FCAF" blockId="66.[192,1441,195,859]" box="[240,1311,771,795]" pageId="66" pageNumber="65">
|
||
<emphasis id="B92BEAA3FFBAF845FF31BB6642BAFCAF" bold="true" box="[240,689,771,795]" pageId="66" pageNumber="65">
|
||
<emphasis id="B92BEAA3FFBAF845FF31BB664170FCAE" bold="true" box="[240,379,771,794]" italics="true" pageId="66" pageNumber="65">
|
||
<typeStatus id="54E48813FFBAF845FF31BB664170FCAE" box="[240,379,771,794]" pageId="66" pageNumber="65" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
—
|
||
<collectingCountry id="F3487621FFBAF845FE57BB664224FCAF" box="[406,559,771,795]" name="Ecuador" pageId="66" pageNumber="65">ECUADOR</collectingCountry>
|
||
:
|
||
<collectingRegion id="499BF853FFBAF845FDFDBB6642A0FCAF" box="[572,683,771,795]" country="Ecuador" name="Pastaza" pageId="66" pageNumber="65">Pastaza</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
same data as
|
||
<typeStatus id="54E48813FFBAF845FCA5BB6643D9FCAF" box="[868,978,771,795]" pageId="66" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
–
|
||
<specimenCount id="9D59FD38FFBAF845FC23BB66443AFCAF" box="[994,1073,771,795]" pageId="66" pageNumber="65" type="male">1 male</specimenCount>
|
||
(alcohol) (
|
||
<collectionCode id="ED4EAE74FFBAF845FB73BB664518FCAF" box="[1202,1299,771,795]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="66" pageNumber="65" type="Museum">NMNH</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE036B1FFBAF845FF31BB46455DFCEF" blockId="66.[192,1441,195,859]" pageId="66" pageNumber="65">
|
||
<emphasis id="B92BEAA3FFBAF845FF31BB464179FC8F" box="[240,370,803,827]" italics="true" pageId="66" pageNumber="65">Etymology</emphasis>
|
||
—This species is named
|
||
<taxonomicName id="4C5F4D32FFBAF845FD61BB414353FC8F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[672,856,803,827]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="66" pageNumber="65" phylum="Tracheophyta" rank="species" species="akrogeneios">
|
||
<emphasis id="B92BEAA3FFBAF845FD61BB414353FC8F" box="[672,856,803,827]" italics="true" pageId="66" pageNumber="65">M. akrogeneios</emphasis>
|
||
</taxonomicName>
|
||
from a Greek word meaning “with a prominent chin,” and referring to the projecting ventral margin of the dorsal phallic spine in this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |