213 lines
32 KiB
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213 lines
32 KiB
XML
<document id="F8F2C9CF0959753083D0161A021683AD" ID-DOI="10.5281/zenodo.5170203" ID-ISSN="1942-1354" ID-Zenodo-Dep="5170203" ID-ZooBank="AB1A57F0-7CB4-4830-920B-DF219740A596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628281492031" checkinUser="felipe" docAuthor="Blahnik, Roger J. & Holzenthal, Ralph W." docDate="2017" docId="03F687A7FFA7F867FF01BC46412BF8CF" docLanguage="en" docName="InsectaMundi.0602.1-251.pdf" docOrigin="Insecta Mundi 2017 (602)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Mortoniella (Nanotrichia) Blahnik & Holzenthal 2017, new subgenus" docType="treatment" docVersion="4" lastPageNumber="95" masterDocId="FFCFFFDFFFF8F807FFC1B865400BFFB4" masterDocTitle="Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *" masterLastPageNumber="251" masterPageNumber="1" pageNumber="94" updateTime="1699039969677" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:titleInfo id="DC8D49C41BF2041978D049DC1E1E758B">
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<mods:title id="244A7596B9DB4415F6F46CF393D66EC9">Revision of the northern South American species of Mortoniella Ulmer 1906 (Trichoptera: Glossosomatidae: Protoptilinae) *</mods:title>
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<mods:name id="32146DAA6977A8A0C5385390F0BF6B50" type="personal">
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<mods:namePart id="B1C5F41F80C1902FEF8B23CB0DF1449E">Blahnik, Roger J.</mods:namePart>
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<mods:name id="6859F6CABE8585E5C22B546822124AAD" type="personal">
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<mods:namePart id="B5D7830B6A2FA3750D9CBF195F849D64">Holzenthal, Ralph W.</mods:namePart>
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<mods:typeOfResource id="7868BFEFA5D17AEB4CDCDEA24F4A456C">text</mods:typeOfResource>
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<mods:title id="7CE7C8FE17D0086F0869E2357E19DC47">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="B7D0670CE88BF2CB71F9B2A48365967B">
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<mods:date id="EF11032829BFFF367C735FAC1568C0CE">2017</mods:date>
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<mods:detail id="337629921413D6371A6297753882BAD4" type="pubDate">
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<mods:number id="24041954CFB5EF32B5E8337976C254F5">2017-12-29</mods:number>
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<mods:detail id="F650D5C4E71AE717049459D7A223A482" type="volume">
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<mods:number id="C061CA635B0EBB1D7112AA93743F959A">2017</mods:number>
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<mods:detail id="75A854E94DBBD623DB0806C180B74093" type="issue">
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<mods:number id="EFEF7F6A93775F78886E99AA9CD14CD5">602</mods:number>
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<mods:classification id="F12792D8CC4FDBAB2953D2529D977D33">journal article</mods:classification>
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<mods:identifier id="955BBD298494529A110C5B6337164E51" type="DOI">10.5281/zenodo.5170203</mods:identifier>
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<mods:identifier id="4869562E5F2055FC58CC51EFF3951C9F" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="9C7E45576D3E879D422FBBF38E386613" type="Zenodo-Dep">5170203</mods:identifier>
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<treatment id="03F687A7FFA7F867FF01BC46412BF8CF" ID-DOI="http://doi.org/10.5281/zenodo.5186335" ID-Zenodo-Dep="5186335" LSID="urn:lsid:plazi:treatment:03F687A7FFA7F867FF01BC46412BF8CF" httpUri="http://treatment.plazi.org/id/03F687A7FFA7F867FF01BC46412BF8CF" lastPageId="96" lastPageNumber="95" pageId="95" pageNumber="94">
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<subSubSection id="C345653AFFA7F858FF01BC46431EFB8F" box="[192,789,1059,1083]" pageId="95" pageNumber="94" type="nomenclature">
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<paragraph id="8BE036B1FFA7F858FF01BC46431EFB8F" blockId="95.[192,794,1059,1148]" box="[192,789,1059,1083]" pageId="95" pageNumber="94">
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<heading id="D0A881DDFFA7F858FF01BC46431EFB8F" bold="true" box="[192,789,1059,1083]" fontSize="10" level="2" pageId="95" pageNumber="94" reason="3">
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<emphasis id="B92BEAA3FFA7F858FF01BC46431EFB8F" bold="true" box="[192,789,1059,1083]" pageId="95" pageNumber="94">
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<taxonomicName id="4C5F4D32FFA7F858FF01BC464233FB8F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[192,568,1059,1083]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="95" pageNumber="94" phylum="Tracheophyta" rank="subGenus" status="subgen. nov." subGenus="Nanotrichia">
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<emphasis id="B92BEAA3FFA7F858FF01BC464163FB8F" bold="true" box="[192,360,1059,1083]" italics="true" pageId="95" pageNumber="94">Mortoniella</emphasis>
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(
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<emphasis id="B92BEAA3FFA7F858FEBCBC464226FB8F" bold="true" box="[381,557,1059,1083]" italics="true" pageId="95" pageNumber="94">Nanotrichia</emphasis>
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)
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFA7F858FD8BBC46431EFB8F" box="[586,789,1059,1083]" pageId="95" pageNumber="94" rank="subGenus">new subgenus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C345653AFFA7F867FF01BC01412BF8CF" lastPageId="96" lastPageNumber="95" pageId="95" pageNumber="94" type="materials_examined">
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<paragraph id="8BE036B1FFA7F858FF01BC014311FBCF" blockId="95.[192,794,1059,1148]" box="[192,794,1123,1148]" pageId="95" pageNumber="94">
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<typeStatus id="54E48813FFA7F858FF01BC0140F4FBC8" box="[192,255,1124,1148]" pageId="95" pageNumber="94">Type</typeStatus>
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species:
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<taxonomicName id="4C5F4D32FFA7F858FEB5BC06431DFBCF" authority="Flint, 1974" authorityName="Flint" authorityYear="1974" box="[372,790,1123,1147]" class="Insecta" family="Glossosomatidae" genus="Mexitrichia" kingdom="Animalia" order="Trichoptera" pageId="95" pageNumber="94" phylum="Arthropoda" rank="species" species="pacuara">
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<emphasis id="B92BEAA3FFA7F858FEB5BC064271FBCF" box="[372,634,1123,1147]" italics="true" pageId="95" pageNumber="94">Mexitrichia pacuara</emphasis>
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<bibRefCitation id="EFCE4B40FFA7F858FD42BC06431DFBCF" author="Flint, O. S., Jr." box="[643,790,1123,1147]" pageId="95" pageNumber="94" pagination="1 - 30" refId="ref106579" refString="Flint, O. S., Jr. 1974. Studies of Neotropical caddisflies, XVIII: New species of Rhyacophilidae and Glossosomatidae (Trichoptera). Smithsonian Contributions to Zoology 169: 1 - 30." type="journal article" year="1974">Flint, 1974</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BE036B1FFA7F867FF31BCC643A9FE0F" blockId="95.[192,1441,1187,1916]" lastBlockId="96.[192,1440,195,444]" lastPageId="96" lastPageNumber="95" pageId="95" pageNumber="94">
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This new subgenus can be distinguished from the subgenus
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<taxonomicName id="4C5F4D32FFA7F858FC30BCC6448CFB0F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[1009,1159,1187,1211]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="95" pageNumber="94" phylum="Tracheophyta" rank="genus">
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<emphasis id="B92BEAA3FFA7F858FC30BCC6448CFB0F" box="[1009,1159,1187,1211]" italics="true" pageId="95" pageNumber="94">Mortoniella</emphasis>
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</taxonomicName>
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by several characters considered collectively. One of these, the small size of the species, is suggested by the name assigned to the subgenus. The most distinctive character defining the subgenus is the reduction of the hind wing venation to include only fork
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<collectionCode id="ED4EAE74FFA7F858FD59BD6142B9FAAF" box="[664,690,1284,1307]" pageId="95" pageNumber="94">II</collectionCode>
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.
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<collectionCode id="ED4EAE74FFA7F858FD02BD6642DDFAAF" box="[707,726,1283,1307]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="95" pageNumber="94" type="Herbarium">A</collectionCode>
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similar reduction occurs in some species and species subgroups of the leroda group of the subgenus
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<taxonomicName id="4C5F4D32FFA7F858FCF2BD4643C2FA8F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[819,969,1315,1339]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="95" pageNumber="94" phylum="Tracheophyta" rank="genus">
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<emphasis id="B92BEAA3FFA7F858FCF2BD4643C2FA8F" box="[819,969,1315,1339]" italics="true" pageId="95" pageNumber="94">Mortoniella</emphasis>
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</taxonomicName>
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, but generally the costal margin of these species has a distinct bend or inflection (
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<figureCitation id="13642A34FFA7F858FCDDBD264396FAEF" box="[796,925,1347,1371]" captionStart="Figures 100-101" captionStartId="238.[192,291,1836,1858]" captionText="Figures 100-101. Mortoniella (M.), leroda group species, wings. 100—Mortoniella limona Flint [limona subgroup]. A—forewing; B—hind wing. 101—Mortoniella applanata, n. sp. [unplaced to subgroup]. A— forewing; B—hind wing." pageId="95" pageNumber="94">Fig. 101B</figureCitation>
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). This is much less distinct in species of
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<taxonomicName id="4C5F4D32FFA7F858FF01BD064157FACF" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[192,348,1379,1403]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="95" pageNumber="130" phylum="Tracheophyta" rank="subGenus" subGenus="Nanotrichia">
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<emphasis id="B92BEAA3FFA7F858FF01BD064157FACF" box="[192,348,1379,1403]" italics="true" pageId="95" pageNumber="94">Nanotrichia</emphasis>
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</taxonomicName>
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, such that the overall wing shape is more spear-like (
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<figureCitation id="13642A34FFA7F858FBC2BD064528FACF" box="[1027,1315,1379,1403]" captionStart="Figures 102-104" captionStartId="239.[192,287,1807,1829]" captionTargetBox="[337,1299,350,1638]" captionTargetId="figure-70@239.[333,1300,350,1638]" captionTargetPageId="239" captionText="Figures 102-104. Mortoniella (Nanotrichia) species, wings. 102—Mortoniella eduardoi (Rueda Martín and Gibon) [velasquezi group]. A—forewing; B—hind wing. 103—Mortoniella collegarum (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing. 104—Mortoniella usseglioi (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing." figureDoi="http://doi.org/10.5281/zenodo.5170409" httpUri="https://zenodo.org/record/5170409/files/figure.png" pageId="95" pageNumber="94">Fig. 102B, 103B, 104B</figureCitation>
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). Another character synapomorphy that seems to be consistent for the included species is that the middle legs have only 3 spurs (rarely 2).
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<collectionCode id="ED4EAE74FFA7F858FDB2BDC6428DFA0F" box="[627,646,1443,1467]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="95" pageNumber="94" type="Herbarium">A</collectionCode>
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similar reduction occurs in various species in the leroda and bilineata groups of the subgenus
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<taxonomicName id="4C5F4D32FFA7F858FDAFBDA6430FFA6F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[622,772,1475,1499]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="95" pageNumber="94" phylum="Tracheophyta" rank="genus">
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<emphasis id="B92BEAA3FFA7F858FDAFBDA6430FFA6F" box="[622,772,1475,1499]" italics="true" pageId="95" pageNumber="94">Mortoniella</emphasis>
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</taxonomicName>
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, but the usual spur formula in both of these groups is 0:4:4. Additionally, the anterior margin of segment IX of males in the subgenus
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<taxonomicName id="4C5F4D32FFA7F858FB27BD864589FA4F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[1254,1410,1507,1531]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="95" pageNumber="94" phylum="Tracheophyta" rank="subGenus" subGenus="Nanotrichia">
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<emphasis id="B92BEAA3FFA7F858FB27BD864589FA4F" box="[1254,1410,1507,1531]" italics="true" pageId="95" pageNumber="94">Nanotrichia</emphasis>
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</taxonomicName>
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is distinctly produced in its ventral ½ and the posterior margin is nearly linear. The lobes of the segment are separated dorsally by ½ or more of the width of the segment (with several exceptions). Species in the bilineata group of the subgenus
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<taxonomicName id="4C5F4D32FFA7F858FCC5BE264393F9EF" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[772,920,1603,1627]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="95" pageNumber="94" phylum="Tracheophyta" rank="genus">
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<emphasis id="B92BEAA3FFA7F858FCC5BE264393F9EF" box="[772,920,1603,1627]" italics="true" pageId="95" pageNumber="94">Mortoniella</emphasis>
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</taxonomicName>
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also have the anterior margin of segment IX produced in the ventral ½ and the lobes widely separated dorsally, but these species are larger and have retained fork
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<collectionCode id="ED4EAE74FFA7F858FE29BEE141F0F92F" box="[488,507,1668,1691]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="95" pageNumber="94" type="Museum">V</collectionCode>
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in the hind wing, and usually also fork
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<collectionCode id="ED4EAE74FFA7F858FC31BEE1441DF92F" box="[1008,1046,1668,1691]" pageId="95" pageNumber="94">III</collectionCode>
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. In general, the very shortened phallobase and relatively simple structure of tergum X of species in
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<taxonomicName id="4C5F4D32FFA7F858FB96BEC644F8F90F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[1111,1267,1699,1723]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="95" pageNumber="130" phylum="Tracheophyta" rank="subGenus" subGenus="Nanotrichia">
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<emphasis id="B92BEAA3FFA7F858FB96BEC644F8F90F" box="[1111,1267,1699,1723]" italics="true" pageId="95" pageNumber="94">Nanotrichia</emphasis>
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</taxonomicName>
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also support their separation from the subgenus
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<taxonomicName id="4C5F4D32FFA7F858FD4DBEA64329F96F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[652,802,1731,1755]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="95" pageNumber="94" phylum="Tracheophyta" rank="genus">
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<emphasis id="B92BEAA3FFA7F858FD4DBEA64329F96F" box="[652,802,1731,1755]" italics="true" pageId="95" pageNumber="94">Mortoniella</emphasis>
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</taxonomicName>
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. Two very distinctive species groups are assigned to
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<taxonomicName id="4C5F4D32FFA7F858FF27BE86418DF94F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[230,390,1763,1787]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="95" pageNumber="130" phylum="Tracheophyta" rank="subGenus" subGenus="Nanotrichia">
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<emphasis id="B92BEAA3FFA7F858FF27BE86418DF94F" box="[230,390,1763,1787]" italics="true" pageId="95" pageNumber="94">Nanotrichia</emphasis>
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</taxonomicName>
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, the ormina and velasquezi groups. These are most convincingly defined by differences in the female genitalia. Of these, the ormina group is the more variable and the placement of
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<taxonomicName id="4C5F4D32FFA7F858FEB0BF41435BF88F" authority="Blahnik and Holzenthal" authorityName="Blahnik and Holzenthal. The" authorityYear="2008" box="[369,848,1827,1851]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="95" pageNumber="94" phylum="Tracheophyta" rank="species" species="rodmani">
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<emphasis id="B92BEAA3FFA7F858FEB0BF414200F88F" box="[369,523,1827,1851]" italics="true" pageId="95" pageNumber="94">M. rodmani</emphasis>
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Blahnik and Holzenthal
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</taxonomicName>
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and
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<taxonomicName id="4C5F4D32FFA7F858FC57BF41443DF88F" authority="Blahnik & Holzenthal, 2017" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[918,1078,1827,1851]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="95" pageNumber="94" phylum="Tracheophyta" rank="species" species="simplicis" status="sp. nov.">
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<emphasis id="B92BEAA3FFA7F858FC57BF41443DF88F" box="[918,1078,1827,1851]" italics="true" pageId="95" pageNumber="94">M. simplicis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21857D8FFA7F858FB88BF41449BF88F" box="[1097,1168,1828,1851]" pageId="95" pageNumber="94" rank="species">n. sp.</taxonomicNameLabel>
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within this group is conjectural, since females are not known for these species. Some species in both the ormina and velasquezi groups have modified scale-like setae paralleling the major veins of the fore- and/or hind wings.
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<collectionCode id="ED4EAE74FF98F867FEA2B8A6417DFF6F" box="[355,374,195,219]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="96" pageNumber="95" type="Herbarium">A</collectionCode>
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similar development occurs in a few species of the leroda group of the subgenus
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<taxonomicName id="4C5F4D32FF98F867FF01B886415DFF4F" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[192,342,227,251]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="96" pageNumber="95" phylum="Tracheophyta" rank="genus">
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<emphasis id="B92BEAA3FF98F867FF01B886415DFF4F" box="[192,342,227,251]" italics="true" pageId="96" pageNumber="95">Mortoniella</emphasis>
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</taxonomicName>
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. The character is mostly hidden by the normal setae present and may not be readily evident. The scales seem to be readily lost in alcohol preserved specimens. In some species of the velasquezi group, there is a more general and readily apparent field of scale-like setae on the hind wing. These are of somewhat different morphology than the scale-like setae paralleling the veins (stalked basally and upright, rather than leaf-like and flattened). Whether the presence of scale-like setae represents an independent character development in the two groups, or an apomorphy for the group, lost in some taxa, is uncertain.
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</paragraph>
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<paragraph id="8BE036B1FF98F867FF31B9864125FC2F" blockId="96.[192,1441,483,1915]" pageId="96" pageNumber="95">
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<emphasis id="B92BEAA3FF98F867FF31B9864133FE4F" box="[240,312,483,507]" italics="true" pageId="96" pageNumber="95">Adult</emphasis>
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—Length of forewing: 1.7-4.0 mm; females slightly larger than males. Forewing with forks
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<collectionCode id="ED4EAE74FF98F867FEC8BA61411DFDAF" box="[265,278,516,539]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&quot;Alexandru Ioan Cuza&quot; University" pageId="96" pageNumber="95" type="Herbarium">I</collectionCode>
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,
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<collectionCode id="ED4EAE74FF98F867FEE5BA614136FDAF" box="[292,317,516,539]" pageId="96" pageNumber="95">II</collectionCode>
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, and
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<collectionCode id="ED4EAE74FF98F867FE42BA6141A3FDAF" box="[387,424,516,539]" pageId="96" pageNumber="95">III</collectionCode>
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, forks
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<collectionCode id="ED4EAE74FF98F867FE3FBA614202FDAF" box="[510,521,516,539]" country="Romania" lsid="urn:lsid:biocol.org:col:14415" name="&quot;Alexandru Ioan Cuza&quot; University" pageId="96" pageNumber="95" type="Herbarium">I</collectionCode>
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and
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<collectionCode id="ED4EAE74FF98F867FD88BA614254FDAF" box="[585,607,516,539]" pageId="96" pageNumber="95">II</collectionCode>
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usually sessile, sometimes with fork
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<collectionCode id="ED4EAE74FF98F867FB83BA614453FDAF" box="[1090,1112,516,539]" pageId="96" pageNumber="95">II</collectionCode>
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stalked (
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<figureCitation id="13642A34FF98F867FB11BA66455AFDAF" box="[1232,1361,515,539]" captionStart="Figures 102-104" captionStartId="239.[192,287,1807,1829]" captionTargetBox="[337,1299,350,1638]" captionTargetId="figure-70@239.[333,1300,350,1638]" captionTargetPageId="239" captionText="Figures 102-104. Mortoniella (Nanotrichia) species, wings. 102—Mortoniella eduardoi (Rueda Martín and Gibon) [velasquezi group]. A—forewing; B—hind wing. 103—Mortoniella collegarum (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing. 104—Mortoniella usseglioi (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing." figureDoi="http://doi.org/10.5281/zenodo.5170409" httpUri="https://zenodo.org/record/5170409/files/figure.png" pageId="96" pageNumber="95">Fig. 104A</figureCitation>
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); hind wing with fork
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<collectionCode id="ED4EAE74FF98F867FE44BA414190FD8F" box="[389,411,548,571]" pageId="96" pageNumber="95">II</collectionCode>
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only (
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<figureCitation id="13642A34FF98F867FE2ABA46431BFD8F" box="[491,784,547,571]" captionStart="Figures 102-104" captionStartId="239.[192,287,1807,1829]" captionTargetBox="[337,1299,350,1638]" captionTargetId="figure-70@239.[333,1300,350,1638]" captionTargetPageId="239" captionText="Figures 102-104. Mortoniella (Nanotrichia) species, wings. 102—Mortoniella eduardoi (Rueda Martín and Gibon) [velasquezi group]. A—forewing; B—hind wing. 103—Mortoniella collegarum (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing. 104—Mortoniella usseglioi (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing." figureDoi="http://doi.org/10.5281/zenodo.5170409" httpUri="https://zenodo.org/record/5170409/files/figure.png" pageId="96" pageNumber="95">Fig. 102B, 103B, 104B</figureCitation>
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), basal forks of Rs and
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<collectionCode id="ED4EAE74FF98F867FBFDBA41445EFD8F" box="[1084,1109,548,571]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="96" pageNumber="95" type="Herbarium">M</collectionCode>
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veins at about midlength and narrowly separated (ormina group), or with basal fork of Rs more proximal (velasquezi group). Crossveins of forewing (
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<emphasis id="B92BEAA3FF98F867FE36BA01430EFDCF" box="[503,773,612,635]" italics="true" pageId="96" pageNumber="95">r, s, r-m, m, m-cu, cu</emphasis>
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) linear or nearly so and usually hyaline; hind wing with
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<emphasis id="B92BEAA3FF98F867FEC2BAE1413AFD2F" box="[259,305,644,667]" italics="true" pageId="96" pageNumber="95">r-m</emphasis>
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only (ormina group,
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<figureCitation id="13642A34FF98F867FD80BAE6431FFD2F" box="[577,788,643,667]" captionStart="Figures 102-104" captionStartId="239.[192,287,1807,1829]" captionTargetBox="[337,1299,350,1638]" captionTargetId="figure-70@239.[333,1300,350,1638]" captionTargetPageId="239" captionText="Figures 102-104. Mortoniella (Nanotrichia) species, wings. 102—Mortoniella eduardoi (Rueda Martín and Gibon) [velasquezi group]. A—forewing; B—hind wing. 103—Mortoniella collegarum (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing. 104—Mortoniella usseglioi (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing." figureDoi="http://doi.org/10.5281/zenodo.5170409" httpUri="https://zenodo.org/record/5170409/files/figure.png" pageId="96" pageNumber="95">Fig. 103B, 104B</figureCitation>
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), or with crossveins absent (velasquezi group,
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<figureCitation id="13642A34FF98F867FAAFBAE6410DFD0F" captionStart="Figures 102-104" captionStartId="239.[192,287,1807,1829]" captionTargetBox="[337,1299,350,1638]" captionTargetId="figure-70@239.[333,1300,350,1638]" captionTargetPageId="239" captionText="Figures 102-104. Mortoniella (Nanotrichia) species, wings. 102—Mortoniella eduardoi (Rueda Martín and Gibon) [velasquezi group]. A—forewing; B—hind wing. 103—Mortoniella collegarum (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing. 104—Mortoniella usseglioi (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing." figureDoi="http://doi.org/10.5281/zenodo.5170409" httpUri="https://zenodo.org/record/5170409/files/figure.png" pageId="96" pageNumber="95">Fig. 102B</figureCitation>
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). Costal margin of hind wing without distinct inflection, wings more or less spear-shaped. Forewing rather broadly rounded apically in velasquezi group (
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<figureCitation id="13642A34FF98F867FC39BAA64471FD6F" box="[1016,1146,707,731]" captionStart="Figures 102-104" captionStartId="239.[192,287,1807,1829]" captionTargetBox="[337,1299,350,1638]" captionTargetId="figure-70@239.[333,1300,350,1638]" captionTargetPageId="239" captionText="Figures 102-104. Mortoniella (Nanotrichia) species, wings. 102—Mortoniella eduardoi (Rueda Martín and Gibon) [velasquezi group]. A—forewing; B—hind wing. 103—Mortoniella collegarum (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing. 104—Mortoniella usseglioi (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing." figureDoi="http://doi.org/10.5281/zenodo.5170409" httpUri="https://zenodo.org/record/5170409/files/figure.png" pageId="96" pageNumber="95">Fig. 102A</figureCitation>
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), narrow and angulate in ormina group (
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<figureCitation id="13642A34FF98F867FE5FBA86427BFD4F" box="[414,624,739,763]" captionStart="Figures 102-104" captionStartId="239.[192,287,1807,1829]" captionTargetBox="[337,1299,350,1638]" captionTargetId="figure-70@239.[333,1300,350,1638]" captionTargetPageId="239" captionText="Figures 102-104. Mortoniella (Nanotrichia) species, wings. 102—Mortoniella eduardoi (Rueda Martín and Gibon) [velasquezi group]. A—forewing; B—hind wing. 103—Mortoniella collegarum (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing. 104—Mortoniella usseglioi (Rueda Martín and Gibon) [ormina group]. A—forewing; B—hind wing." figureDoi="http://doi.org/10.5281/zenodo.5170409" httpUri="https://zenodo.org/record/5170409/files/figure.png" pageId="96" pageNumber="95">Fig. 103A, 104A</figureCitation>
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), apex of hind wing angulate. Fore- and/or hind wing frequently with adpressed scale-like setae paralleling veins; some species of velasquezi group also with patch of upright scale-like or hair-like setae on hind wing. Spur formula 0:3:4 (0:2:4, as variant in
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<taxonomicName id="4C5F4D32FF98F867FABEBB41413BFCEF" authorityName="Blahnik & Holzenthal" authorityYear="2017" baseAuthorityName="Rueda Martin and Gibon" baseAuthorityYear="2008" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="96" pageNumber="95" phylum="Tracheophyta" rank="species" species="usseglioi">
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<emphasis id="B92BEAA3FF98F867FABEBB41413BFCEF" italics="true" pageId="96" pageNumber="95">M. usseglioi</emphasis>
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</taxonomicName>
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). Overall color varying from light brown to dark brown; forewing usually marked with white or whitish setae at anastamosis, in some species of ormina group with small white spots apically.
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</paragraph>
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<paragraph id="8BE036B1FF98F867FF31BBC64381F94F" blockId="96.[192,1441,483,1915]" pageId="96" pageNumber="95">
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<emphasis id="B92BEAA3FF98F867FF31BBC641A6FC0F" box="[240,429,931,955]" italics="true" pageId="96" pageNumber="95">Male genitalia</emphasis>
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—Ventral process of segment
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<collectionCode id="ED4EAE74FF98F867FCF8BBC14353FC0F" box="[825,856,932,955]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="96" pageNumber="95">VI</collectionCode>
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relatively elongate, narrow, and posteriorly projecting (length usually 2 times width or more). Segment IX with anterolateral margin rounded and produced in ventral ½, posterolateral margin nearly straight; segment deeply mesally excised dorsally and ventrally, forming lateral lobes, lobes usually separated dorsally by ½ or more of width of segment. Tergum X well developed, setose, usually with deep V-shaped or U-shaped apicomesal invagination (
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<taxonomicName id="4C5F4D32FF98F867FDC8BC214295FBEF" baseAuthorityName="Flint" baseAuthorityYear="1963" box="[521,670,1091,1115]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="96" pageNumber="95" phylum="Tracheophyta" rank="species" species="aequalis">
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<emphasis id="B92BEAA3FF98F867FDC8BC214295FBEF" box="[521,670,1091,1115]" italics="true" pageId="96" pageNumber="95">M. aequalis</emphasis>
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</taxonomicName>
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an exception); apicolateral lobes usually simple in structure and not strongly sclerotized. Inferior appendages directly fused to one another and to ventral margin of phallic ensemble, generally short and reduced in ormina group, or in velasquezi group with more distinct dorsolateral process and apex sometimes elongate and fused to mesal spinelike process; appendages always with paired mesal pockets accommodating rod-like projections of phallobase, these very enlarged in velasquezi group, apices of pockets with relatively short to very elongate spine-like projections, typically narrow in ormina group, widened in velasquezi group. Basal segments of parameres variable, usually relatively simple in structure in velasquezi group (short, rounded and fused to one another), often more modified in ormina group, sometimes with elongate rod-like projections; apical segments (paramere appendages) usually present, short to elongate, typically more or less rod-like, often strongly curved. Phallobase short or very short, always with, paired rod-like projections from the apicoventral margin, these short in ormina group (sometimes with apex flared), long and with apices flared in velasquezi group; dorsal margin with moderately elongate mesal spine, relatively simple in structure in some, more often with lateral margins projecting; dorsal phallic spine without basal articulation with phallicata, apex strongly reflexed in most. Phallicata usually short and only indistinctly sclerotized, with distinctly sclerotized, rounded, lateral projection in velasquezi group. Endophallic membrane very variable, usually relatively short and simple in structure, sometimes with dorsal or lateral spines or spine-like structures or with membranous lateral lobes; in velasquezi group with pair of ventral sclerites or spine-like projections and dorsomesal projection, typically with small spines or short spine-like projection; phallotremal spines not usually evident (
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<taxonomicName id="4C5F4D32FF98F867FE64BE81426EF94F" baseAuthorityName="Rueda Martin and Gibon" baseAuthorityYear="2008" box="[421,613,1763,1787]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="96" pageNumber="95" phylum="Tracheophyta" rank="species" species="collegarum">
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<emphasis id="B92BEAA3FF98F867FE64BE81426EF94F" box="[421,613,1763,1787]" italics="true" pageId="96" pageNumber="95">M. collegarum</emphasis>
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</taxonomicName>
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a notable exception).
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</paragraph>
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<paragraph id="8BE036B1FF98F867FF31BF66412BF8CF" blockId="96.[192,1441,483,1915]" pageId="96" pageNumber="95">
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<emphasis id="B92BEAA3FF98F867FF31BF66417EF8AF" box="[240,373,1795,1819]" italics="true" pageId="96" pageNumber="95">Etymology</emphasis>
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—This subgenus is named
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<taxonomicName id="4C5F4D32FF98F867FD09BF664368F8AF" authorityName="Blahnik & Holzenthal" authorityYear="2017" box="[712,867,1795,1819]" class="Magnoliopsida" family="Apocynaceae" genus="Mortoniella" kingdom="Plantae" order="Gentianales" pageId="96" pageNumber="130" phylum="Tracheophyta" rank="subGenus" subGenus="Nanotrichia">
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<emphasis id="B92BEAA3FF98F867FD09BF664368F8AF" box="[712,867,1795,1819]" italics="true" pageId="96" pageNumber="95">Nanotrichia</emphasis>
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</taxonomicName>
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from the Greek words
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<emphasis id="B92BEAA3FF98F867FB42BF6144C5F8AF" box="[1155,1230,1796,1819]" italics="true" pageId="96" pageNumber="95">nanos</emphasis>
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, meaning small, and referring to the small size of the species in the subgenus, and
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<emphasis id="B92BEAA3FF98F867FBCFBF464440F88F" box="[1038,1099,1827,1851]" italics="true" pageId="96" pageNumber="95">trich</emphasis>
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|
- for hair, a term generally applied to species in the order
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<taxonomicName id="4C5F4D32FF98F867FD8ABF2642E8F8EF" baseAuthorityName="Holzenthal and Calor" baseAuthorityYear="2017" box="[587,739,1859,1883]" class="Insecta" kingdom="Animalia" order="Trichoptera" pageId="96" pageNumber="95" phylum="Arthropoda" rank="order">Trichoptera</taxonomicName>
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because of their hairy wings. The name is feminine in gender.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |