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<mods:title id="3072187AF4DEB6EBC9A658534DA7F088">A survey of Palaearctic Dictynidae (Araneae). 1. Taxonomic notes on Dictynomorpha Spassky, 1939, Brigittea Lehtinen, 1967 and Lathys Simon, 1884</mods:title>
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<treatment id="03F6879CB656FFC0FF01E6CF2F72BAC0" ID-DOI="http://doi.org/10.5281/zenodo.6094416" ID-GBIF-Taxon="119598494" ID-Zenodo-Dep="6094416" LSID="urn:lsid:plazi:treatment:03F6879CB656FFC0FF01E6CF2F72BAC0" httpUri="http://treatment.plazi.org/id/03F6879CB656FFC0FF01E6CF2F72BAC0" lastPageId="7" lastPageNumber="136" pageId="6" pageNumber="135">
<subSubSection id="C3456501B656FFC1FF01E6CF2E4ABA3C" box="[151,589,1279,1305]" pageId="6" pageNumber="135" type="nomenclature">
<paragraph id="8BE0368AB656FFC1FF01E6CF2E4ABA3C" blockId="6.[151,589,1279,1305]" box="[151,589,1279,1305]" pageId="6" pageNumber="135">
<heading id="D0A881E6B656FFC1FF01E6CF2E4ABA3C" bold="true" box="[151,589,1279,1305]" fontSize="11" level="1" pageId="6" pageNumber="135" reason="1">
<emphasis id="B92BEA98B656FFC1FF01E6CF2E4ABA3C" bold="true" box="[151,589,1279,1305]" pageId="6" pageNumber="135">
<taxonomicName id="4C5F4D09B656FFC1FF01E6CF2DC4BA3C" ID-CoL="3D7P" ID-ENA="1872029" authority="Lehtinen, 1967" authorityName="Lehtinen" authorityYear="1967" box="[151,451,1279,1305]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B656FFC1FF01E6CF2D06BA3C" bold="true" box="[151,257,1279,1305]" italics="true" pageId="6" pageNumber="135">Brigittea</emphasis>
<bibRefCitation id="EFCE4B7BB656FFC1FE9EE6CF2DC4BA3C" author="Lehtinen" box="[264,451,1279,1305]" pageId="6" pageNumber="135" refString="Lehtinen, P. T. (1967) Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici, 4, 199 - 468." type="journal article" year="1967">Lehtinen, 1967</bibRefCitation>
</taxonomicName>
, reinstated
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3456501B656FFC1FF01E7722DD9BAB1" pageId="6" pageNumber="135" type="reference_group">
<paragraph id="8BE0368AB656FFC1FF01E7722DD9BAB1" blockId="6.[151,510,1345,1428]" pageId="6" pageNumber="135">
<treatmentCitationGroup id="AB4F11A4B656FFC1FF01E7722DD9BAB1" pageId="6" pageNumber="135">
<taxonomicName id="4C5F4D09B656FFC1FF01E7722DC7BA72" ID-CoL="3D7P" authority="Lehtinen 1967: 219" authorityName="Lehtinen" authorityPageNumber="219" authorityYear="1967" box="[151,448,1345,1367]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B656FFC1FF01E7722CF6BA72" box="[151,241,1346,1367]" italics="true" pageId="6" pageNumber="135">Brigittea</emphasis>
<treatmentCitation id="0AFE109BB656FFC1FF61E7712DC7BA72" author="Lehtinen" box="[247,448,1345,1367]" page="219" pageId="6" pageNumber="135" year="1967">
<bibRefCitation id="EFCE4B7BB656FFC1FF61E7712D8ABA72" author="Lehtinen" box="[247,397,1345,1367]" pageId="6" pageNumber="135" refString="Lehtinen, P. T. (1967) Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici, 4, 199 - 468." type="journal article" year="1967">Lehtinen 1967</bibRefCitation>
: 219
</treatmentCitation>
</taxonomicName>
, 360.
<taxonomicName id="4C5F4D09B656FFC1FF01E7502DFDBA50" ID-CoL="3D7P" authority="Miller &amp; Svaton 1978: 9" authorityName="Miller &amp; Svaton" authorityPageNumber="9" authorityYear="1978" box="[151,506,1375,1397]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B656FFC1FF01E7502CF6BA50" box="[151,241,1376,1397]" italics="true" pageId="6" pageNumber="135">Brigittea</emphasis>
:
<treatmentCitation id="0AFE109BB656FFC1FF68E76F2DFDBA50" author="Miller" box="[254,506,1375,1397]" page="9" pageId="6" pageNumber="135" year="1978">
<bibRefCitation id="EFCE4B7BB656FFC1FF68E76F2DE6BA50" author="Miller" box="[254,481,1375,1397]" pageId="6" pageNumber="135" refString="Miller, F. &amp; Svaton, J. (1978) Einige seltene und bisher unbekannte Spinnenarten aus der Slowakei. Annotationes Zoologicae et Botanicae, 126, 1 - 19." type="journal article" year="1978">Miller &amp; Svatoň 1978</bibRefCitation>
: 9
</treatmentCitation>
</taxonomicName>
.
<taxonomicName id="4C5F4D09B656FFC1FF01E74F2DDDBAB1" ID-CoL="44XN" authority="Wunderlich 1987: 224" authorityName="Wunderlich" authorityPageNumber="224" authorityYear="1987" box="[151,474,1406,1428]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B656FFC1FF01E74F2CE0BAB1" box="[151,231,1407,1428]" italics="true" pageId="6" pageNumber="135">Dictyna</emphasis>
:
<treatmentCitation id="0AFE109BB656FFC1FF62E74E2DDDBAB1" author="Wunderlich" box="[244,474,1406,1428]" page="224" pageId="6" pageNumber="135" year="1987">
<bibRefCitation id="EFCE4B7BB656FFC1FF62E74E2DAFBAB1" author="Wunderlich" box="[244,424,1406,1428]" pageId="6" pageNumber="135" refString="Wunderlich, J. (1987) Die Spinnen der Kanarischen Inseln und Madeiras: Adaptive Radiation, Biogeographie, Revisionen und Neubeschreibungen. Taxonomy &amp; Ecology, 1, 1 - 435. [Verlag J. Wunderlich, Straubenhardt]" type="journal article" year="1987">Wunderlich 1987</bibRefCitation>
: 224
</treatmentCitation>
</taxonomicName>
.
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</paragraph>
</subSubSection>
<subSubSection id="C3456501B656FFC1FF01E7F02F22BAFD" box="[151,805,1472,1497]" pageId="6" pageNumber="135" type="materials_examined">
<paragraph id="8BE0368AB656FFC1FF01E7F02F22BAFD" blockId="6.[151,1436,1472,2038]" box="[151,805,1472,1497]" pageId="6" pageNumber="135">
<emphasis id="B92BEA98B656FFC1FF01E7F02D2CBAFC" bold="true" box="[151,299,1472,1497]" pageId="6" pageNumber="135">
<typeStatus id="54E48828B656FFC1FF01E7F02CD5BAFC" box="[151,210,1472,1497]" pageId="6" pageNumber="135">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="4C5F4D09B656FFC1FEACE7F12E8CBAFC" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[314,651,1472,1497]" class="Gastropoda" family="Muricidae" genus="Aranea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="135" phylum="Mollusca" rank="species" species="latens">
<emphasis id="B92BEA98B656FFC1FEACE7F12DD1BAFC" box="[314,470,1472,1497]" italics="true" pageId="6" pageNumber="135">Aranea latens</emphasis>
Fabricius, 1775
</taxonomicName>
[=
<taxonomicName id="4C5F4D09B656FFC1FD24E7F22F13BAFC" box="[690,788,1472,1497]" class="Gastropoda" family="Muricidae" genus="Aranea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="135" phylum="Mollusca" rank="species" species="latens">
<emphasis id="B92BEA98B656FFC1FD24E7F22F13BAFC" box="[690,788,1472,1497]" italics="true" pageId="6" pageNumber="135">B. latens</emphasis>
</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection id="C3456501B656FFC1FF51E7D42D0DB904" pageId="6" pageNumber="135" type="diagnosis">
<paragraph id="8BE0368AB656FFC1FF51E7D42D0DB904" blockId="6.[151,1436,1472,2038]" pageId="6" pageNumber="135">
<emphasis id="B92BEA98B656FFC1FF51E7D42D46BAD8" bold="true" box="[199,321,1508,1533]" pageId="6" pageNumber="135">Diagnosis.</emphasis>
<taxonomicName id="4C5F4D09B656FFC1FEDEE7D62DABBAD8" box="[328,428,1510,1533]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B656FFC1FEDEE7D62DABBAD8" box="[328,428,1510,1533]" italics="true" pageId="6" pageNumber="135">Brigittea</emphasis>
</taxonomicName>
is the only genus among Dictyninae with ctenidia on the male palpal tibia and a bipartite cribellum.
</paragraph>
</subSubSection>
<subSubSection id="C3456501B656FFC0FF51E41C2F72BAC0" lastPageId="7" lastPageNumber="136" pageId="6" pageNumber="135" type="discussion">
<paragraph id="8BE0368AB656FFC1FF51E41C2D8AB83B" blockId="6.[151,1436,1472,2038]" pageId="6" pageNumber="135">
<emphasis id="B92BEA98B656FFC1FF51E41C2D4FB960" bold="true" box="[199,328,1580,1605]" pageId="6" pageNumber="135">Comments</emphasis>
. The genus was created for eight species
<taxonomicName id="4C5F4D09B656FFC1FCB9E41E2867B960" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[815,1120,1580,1605]" class="Gastropoda" family="Muricidae" genus="Aranea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="135" phylum="Mollusca" rank="species" species="latens">
<emphasis id="B92BEA98B656FFC1FCB9E41E2F94B960" box="[815,915,1580,1605]" italics="true" pageId="6" pageNumber="135">B. latens</emphasis>
(Fabricius, 1775)
</taxonomicName>
(♂♀, West Palaearctic),
<taxonomicName id="4C5F4D09B656FFC1FA12E41E2D82B94C" authority="Lucas, 1850" authorityName="Lucas" authorityYear="1850" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="civica">
<emphasis id="B92BEA98B656FFC1FA12E41E2CDDB94C" italics="true" pageId="6" pageNumber="135">B. civica</emphasis>
(Lucas, 1850)
</taxonomicName>
(♂♀, West Palaearctic),
<taxonomicName id="4C5F4D09B656FFC1FD39E4622FC9B94C" authority="Lehtinen, 1967" authorityName="Lehtinen" authorityYear="1967" box="[687,974,1616,1641]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="denisi">
<emphasis id="B92BEA98B656FFC1FD39E4622F12B94C" box="[687,789,1616,1641]" italics="true" pageId="6" pageNumber="135">B. denisi</emphasis>
<bibRefCitation id="EFCE4B7BB656FFC1FCB7E4602FC9B94C" author="Lehtinen" box="[801,974,1616,1641]" pageId="6" pageNumber="135" refString="Lehtinen, P. T. (1967) Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici, 4, 199 - 468." type="journal article" year="1967">Lehtinen, 1967</bibRefCitation>
</taxonomicName>
(♂♀,
<collectingCountry id="F348761AB656FFC1FB89E4602864B94C" box="[1055,1123,1616,1641]" name="Niger" pageId="6" pageNumber="135">Niger</collectingCountry>
),
<taxonomicName id="4C5F4D09B656FFC1FBEDE4622905B94C" box="[1147,1282,1618,1641]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="B92BEA98B656FFC1FBEDE4622905B94C" box="[1147,1282,1618,1641]" italics="true" pageId="6" pageNumber="135">B. innocens</emphasis>
</taxonomicName>
(O. Pickard- Cambridge, 1872) (♂♀, West Palaearctic),
<taxonomicName id="4C5F4D09B656FFC1FD37E4462FD4B9AB" authority="Simon, 1905" authorityName="Simon" authorityYear="1905" box="[673,979,1653,1678]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="turbida">
<emphasis id="B92BEA98B656FFC1FD37E4462F1CB9AB" box="[673,795,1653,1678]" italics="true" pageId="6" pageNumber="135">B. turbida</emphasis>
(
<bibRefCitation id="EFCE4B7BB656FFC1FCA7E4452FCDB9AB" author="Simon" box="[817,970,1653,1678]" pageId="6" pageNumber="135" refString="Simon, E. (1905) Voyage de M. Maurice Maindron dans l'Inde meridionale. Arachnides. Annales de la Societe Entomologique de France, 74, 160 - 180." type="journal article" year="1905">Simon, 1905</bibRefCitation>
)
</taxonomicName>
(♀,
<collectingCountry id="F348761AB656FFC1FB83E4452848B9AB" box="[1045,1103,1653,1678]" name="India" pageId="6" pageNumber="135">India</collectingCountry>
&amp;
<collectingCountry id="F348761AB656FFC1FB14E44528FCB9AB" box="[1154,1275,1653,1678]" name="Sri Lanka" pageId="6" pageNumber="135">Sri Lanka</collectingCountry>
),
<taxonomicName id="4C5F4D09B656FFC1FA80E4462D68B994" authority="Caporiacco, 1949" authorityName="Caporiacco" authorityYear="1949" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="tullgreni">
<emphasis id="B92BEA98B656FFC1FA80E446299CB9AB" box="[1302,1435,1653,1678]" italics="true" pageId="6" pageNumber="135">B. tullgreni</emphasis>
(Caporiacco, 1949)
</taxonomicName>
(♂,
<collectingCountry id="F348761AB656FFC1FE32E4A92DF6B994" box="[420,497,1689,1713]" name="Kenya" pageId="6" pageNumber="135">Kenya</collectingCountry>
),
<taxonomicName id="4C5F4D09B656FFC1FD92E4AA2F15B994" authority="Simon, 1873" authorityName="Simon" authorityYear="1873" box="[516,786,1688,1713]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B92BEA98B656FFC1FD92E4AA2E6EB994" box="[516,617,1690,1713]" italics="true" pageId="6" pageNumber="135">B. vicina</emphasis>
(Simon, 1873)
</taxonomicName>
(♂♀,West Palaearctic) and
<taxonomicName id="4C5F4D09B656FFC1FBDBE4AA295DB994" authority="Tikader, 1966" authorityName="Tikader" authorityYear="1966" box="[1101,1370,1688,1713]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="umai">
<emphasis id="B92BEA98B656FFC1FBDBE4AA28A3B994" box="[1101,1188,1690,1713]" italics="true" pageId="6" pageNumber="135">B. umai</emphasis>
(Tikader, 1966)
</taxonomicName>
(♂♀,
<collectingCountry id="F348761AB656FFC1FF01E48D2CD2B9F3" box="[151,213,1725,1750]" name="India" pageId="6" pageNumber="135">India</collectingCountry>
).
<bibRefCitation id="EFCE4B7BB656FFC1FF7EE48D2D9AB9F3" author="Lehtinen" box="[232,413,1725,1750]" pageId="6" pageNumber="135" refString="Lehtinen, P. T. (1967) Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici, 4, 199 - 468." type="journal article" year="1967">Lehtinen (1967)</bibRefCitation>
created a table with characters for 23
<taxonomicName id="4C5F4D09B656FFC1FCDFE48D2FC3B9F3" box="[841,964,1725,1750]" class="Arachnida" family="Dictynidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="family">Dictynidae</taxonomicName>
genera. According to this table,
<emphasis id="B92BEA98B656FFC1FAA7E48E299CB9F0" box="[1329,1435,1726,1749]" italics="true" pageId="6" pageNumber="135">
<taxonomicName id="4C5F4D09B656FFC1FAA7E48E2990B9F0" box="[1329,1431,1726,1749]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">Brigittea</taxonomicName>
,
</emphasis>
unlike
<emphasis id="B92BEA98B656FFC1FF71E4D22D40B9DC" box="[231,327,1762,1785]" italics="true" pageId="6" pageNumber="135">
<taxonomicName id="4C5F4D09B656FFC1FF71E4D22D45B9DC" box="[231,322,1762,1785]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">Dictyna</taxonomicName>
,
</emphasis>
has a bipartite cribellum, ventral boss and anterior teeth on the male chelicerae, and a spiraled structure of the vulva.
</paragraph>
<paragraph id="8BE0368AB656FFC1FF51E5182DC4B8AC" blockId="6.[151,1436,1472,2038]" pageId="6" pageNumber="135">
<bibRefCitation id="EFCE4B7BB656FFC1FF51E5182DDEB864" author="Miller" box="[199,473,1832,1857]" pageId="6" pageNumber="135" refString="Miller, F. &amp; Svaton, J. (1978) Einige seltene und bisher unbekannte Spinnenarten aus der Slowakei. Annotationes Zoologicae et Botanicae, 126, 1 - 19." type="journal article" year="1978">Miller &amp; Svatoň (1978)</bibRefCitation>
revised European
<taxonomicName id="4C5F4D09B656FFC1FD25E51A2F10B864" box="[691,791,1834,1857]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B656FFC1FD25E51A2F10B864" box="[691,791,1834,1857]" italics="true" pageId="6" pageNumber="135">Brigittea</emphasis>
</taxonomicName>
and provided an additional character that separates this genus from
<taxonomicName id="4C5F4D09B656FFC1FEB5E57E2D7BB840" box="[291,380,1870,1893]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B656FFC1FEB5E57E2D7BB840" box="[291,380,1870,1893]" italics="true" pageId="6" pageNumber="135">Dictyna</emphasis>
</taxonomicName>
widely separated receptacles [contiguous in the
<typeStatus id="54E48828B656FFC1FC5DE57E2FFCB843" box="[971,1019,1870,1894]" pageId="6" pageNumber="135">type</typeStatus>
species,
<taxonomicName id="4C5F4D09B656FFC1FBF1E57E2CDFB8AC" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="species" species="arundinacea">
<emphasis id="B92BEA98B656FFC1FBF1E57E291DB843" box="[1127,1306,1869,1894]" italics="true" pageId="6" pageNumber="135">D. arundinacea</emphasis>
(Linnaeus, 1758)
</taxonomicName>
and related species].
</paragraph>
<paragraph id="8BE0368AB656FFC0FF51E5A62EE2BE39" blockId="6.[151,1436,1472,2038]" lastBlockId="7.[151,1437,151,1365]" lastPageId="7" lastPageNumber="136" pageId="6" pageNumber="135">
<bibRefCitation id="EFCE4B7BB656FFC1FF51E5A62DB7B88B" box="[199,432,1941,1966]" pageId="6" pageNumber="135" refString="Merrett, P., Locket, G. H. &amp; Millidge, A. F. (1985) A check list of British spiders. Bulletin of the British arachnological Society, 6, 381 - 403." type="journal article">
Merrett
<emphasis id="B92BEA98B656FFC1FEB5E5A62D52B88B" box="[291,341,1941,1966]" italics="true" pageId="6" pageNumber="135">et al</emphasis>
. (1985)
</bibRefCitation>
, in a faunistic paper, doubted the validity of
<taxonomicName id="4C5F4D09B656FFC1FC38E5A62815B888" box="[942,1042,1942,1965]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B656FFC1FC38E5A62815B888" box="[942,1042,1942,1965]" italics="true" pageId="6" pageNumber="135">Brigittea</emphasis>
</taxonomicName>
stating that “
<emphasis id="B92BEA98B656FFC1FB3DE5A628C1B888" box="[1195,1222,1942,1965]" italics="true" pageId="6" pageNumber="135">D.</emphasis>
[
<emphasis id="B92BEA98B656FFC1FB46E5A62913B888" box="[1232,1300,1942,1965]" italics="true" pageId="6" pageNumber="135">ictyna</emphasis>
]
<taxonomicName id="4C5F4D09B656FFC1FAB3E5A5296FB88B" box="[1317,1384,1941,1966]" class="Gastropoda" family="Muricidae" genus="Aranea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="6" pageNumber="135" phylum="Mollusca" rank="species" species="latens">
<emphasis id="B92BEA98B656FFC1FAB3E5A5296FB88B" box="[1317,1384,1941,1966]" italics="true" pageId="6" pageNumber="135">latens</emphasis>
</taxonomicName>
was included in the new genus
<taxonomicName id="4C5F4D09B656FFC1FE52E58A2E2FB8F4" box="[452,552,1978,2001]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="135" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B656FFC1FE52E58A2E2FB8F4" box="[452,552,1978,2001]" italics="true" pageId="6" pageNumber="135">Brigittea</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFCE4B7BB656FFC1FDC7E5882F00B8F4" author="Lehtinen" box="[593,775,1976,2001]" pageId="6" pageNumber="135" refString="Lehtinen, P. T. (1967) Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici, 4, 199 - 468." type="journal article" year="1967">Lehtinen (1967)</bibRefCitation>
on account of its bipartite cribellum and palpal characters; this was followed by
<bibRefCitation id="EFCE4B7BB656FFC1FE1AE5ED2E39B8D0" author="Brignoli" box="[396,574,2013,2038]" pageId="6" pageNumber="135" refString="Brignoli, P. M. (1983) A catalogue of the Araneae described between 1940 and 1981. Manchester University Press, Manchester, 755 pp." type="book" year="1983">Brignoli (1983)</bibRefCitation>
, but since there seems to be some doubt about the validity of this genus we have decided to leave
<taxonomicName id="4C5F4D09B657FFC0FE01E2A72DDDBF95" box="[407,474,151,176]" class="Gastropoda" family="Muricidae" genus="Aranea" higherTaxonomySource="CoL" kingdom="Animalia" order="Neogastropoda" pageId="7" pageNumber="136" phylum="Mollusca" rank="species" species="latens">
<emphasis id="B92BEA98B657FFC0FE01E2A72DDDBF95" box="[407,474,151,176]" italics="true" pageId="7" pageNumber="136">latens</emphasis>
</taxonomicName>
in
<taxonomicName id="4C5F4D09B657FFC0FD97E2A92E5DBF95" box="[513,602,153,176]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FD97E2A92E5DBF95" box="[513,602,153,176]" italics="true" pageId="7" pageNumber="136">Dictyna</emphasis>
</taxonomicName>
for the present”.
<bibRefCitation id="EFCE4B7BB657FFC0FCB2E2A72FFDBF95" author="Wunderlich" box="[804,1018,151,176]" pageId="7" pageNumber="136" refString="Wunderlich, J. (1987) Die Spinnen der Kanarischen Inseln und Madeiras: Adaptive Radiation, Biogeographie, Revisionen und Neubeschreibungen. Taxonomy &amp; Ecology, 1, 1 - 435. [Verlag J. Wunderlich, Straubenhardt]" type="journal article" year="1987">Wunderlich (1987)</bibRefCitation>
used these arguments in favour of a formal synonymy of the two genera and suggested that this taxon be regarded as the “
<taxonomicName id="4C5F4D09B657FFC0FBF6E28D2904BFF0" box="[1120,1283,188,213]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="latens">
<emphasis id="B92BEA98B657FFC0FBF6E28D2904BFF0" box="[1120,1283,188,213]" italics="true" pageId="7" pageNumber="136">Dictyna latens</emphasis>
</taxonomicName>
-Gruppe”. He extended this group to eight species by adding
<taxonomicName id="4C5F4D09B657FFC0FD22E2D12FF4BFDD" authority="Schmidt, 1968" authorityName="Schmidt" authorityYear="1968" box="[692,1011,223,248]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="guanchae">
<emphasis id="B92BEA98B657FFC0FD22E2D12F42BFDD" box="[692,837,223,248]" italics="true" pageId="7" pageNumber="136">D. guanchae</emphasis>
Schmidt, 1968
</taxonomicName>
and
<taxonomicName id="4C5F4D09B657FFC0FBBBE2D1299BBFDD" authority="Wunderlich, 1987" authorityName="Wunderlich" authorityYear="1987" box="[1069,1436,223,248]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="agaetensis">
<emphasis id="B92BEA98B657FFC0FBBBE2D128CFBFDD" box="[1069,1224,225,248]" italics="true" pageId="7" pageNumber="136">D. agaetensis</emphasis>
<bibRefCitation id="EFCE4B7BB657FFC0FB47E2EF299BBFDD" author="Wunderlich" box="[1233,1436,223,248]" pageId="7" pageNumber="136" refString="Wunderlich, J. (1987) Die Spinnen der Kanarischen Inseln und Madeiras: Adaptive Radiation, Biogeographie, Revisionen und Neubeschreibungen. Taxonomy &amp; Ecology, 1, 1 - 435. [Verlag J. Wunderlich, Straubenhardt]" type="journal article" year="1987">Wunderlich, 1987</bibRefCitation>
</taxonomicName>
(currently these names are considered as synonyms).
</paragraph>
<paragraph id="8BE0368AB657FFC0FF51E31828FBBEAD" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="136">
However, while studying
<taxonomicName id="4C5F4D09B657FFC0FE60E3172E76BE65" box="[502,625,295,320]" class="Arachnida" family="Dictynidae" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="family">Dictynidae</taxonomicName>
from
<collectingCountry id="F348761AB657FFC0FD56E3172F41BE65" box="[704,838,295,320]" name="Kazakhstan" pageId="7" pageNumber="136">Kazakhstan</collectingCountry>
, we found additional characters which support the validity of
<taxonomicName id="4C5F4D09B657FFC0FEB6E37D2D83BE41" box="[288,388,333,356]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FEB6E37D2D83BE41" box="[288,388,333,356]" italics="true" pageId="7" pageNumber="136">Brigittea</emphasis>
</taxonomicName>
. The male palp in all Dictyninae and partially in Tricholathysinae has a very similar conformation; therefore somatic characters play an important role in the recognition of these genera.
</paragraph>
<paragraph id="8BE0368AB657FFC0FF51E3A42ECCBD18" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="136">
The bipartite cribellum is known in only five genera of Dictyninae and was not reported in Tricholathysinae.
<taxonomicName id="4C5F4D09B657FFC0FF01E3892CFCBEF5" box="[151,251,441,464]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FF01E3892CFCBEF5" box="[151,251,441,464]" italics="true" pageId="7" pageNumber="136">Brigittea</emphasis>
</taxonomicName>
is the only genus among Dictyninae with ctenidia on the male palpal tibia and a bipartite cribellum. The cribellum of
<taxonomicName id="4C5F4D09B657FFC0FEB0E3ED2D8DBED1" box="[294,394,477,500]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FEB0E3ED2D8DBED1" box="[294,394,477,500]" italics="true" pageId="7" pageNumber="136">Brigittea</emphasis>
</taxonomicName>
has only one anterior row of setae (
<emphasis id="B92BEA98B657FFC0FC8EE3EC2F33BED6" box="[792,820,476,499]" italics="true" pageId="7" pageNumber="136">As</emphasis>
), whereas the remaining cribellate area is covered only with fine spigots (
<emphasis id="B92BEA98B657FFC0FEFBE0312D8EBD3D" box="[365,393,513,536]" italics="true" pageId="7" pageNumber="136">Fs</emphasis>
). This contrasts with that in
<taxonomicName id="4C5F4D09B657FFC0FD75E0312F3BBD3D" box="[739,828,513,536]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FD75E0312F3BBD3D" box="[739,828,513,536]" italics="true" pageId="7" pageNumber="136">Dictyna</emphasis>
</taxonomicName>
in which the anterior two-thirds of the cribellum is covered with setae and one third with fine spigots.
</paragraph>
<paragraph id="8BE0368AB657FFC0FF51E07728D3BC31" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="136">
The male chelicerae each have a distinct baso-lateral tooth (
<emphasis id="B92BEA98B657FFC0FC35E0792FBBBD45" box="[931,956,585,608]" italics="true" pageId="7" pageNumber="136">Bt</emphasis>
) in
<taxonomicName id="4C5F4D09B657FFC0FC61E079285CBD45" box="[1015,1115,585,608]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FC61E079285CBD45" box="[1015,1115,585,608]" italics="true" pageId="7" pageNumber="136">Brigittea</emphasis>
</taxonomicName>
(
<figureCitation id="13642A0FB657FFC0FBE6E07728C5BD45" box="[1136,1218,583,608]" captionStart="FIGURES 26 36" captionStartId="8.[151,264,1925,1948]" captionTargetBox="[259,1327,193,1888]" captionTargetId="figure@8.[248,1339,193,1893]" captionTargetPageId="8" captionText="FIGURES 26 36. Dictyna arundinacea (26 30), Brigittea latens (31 36). 26, 31 Male palp, ventral. 27, 32 Ctenidia, dorsoretrolateral. 28, 33 Chelicera, mesal. 29, 34 Tip of conductor, retrolateral. 30, 35 Same, ventral. 36 Tip of embolus. Abbreviations: Bt—retrobasal tooth of chelicera; Cr—cheliceral ridge; Ct—cheliceral teeth." httpUri="https://zenodo.org/record/243582/files/figure.png" pageId="7" pageNumber="136">Fig 33</figureCitation>
) which is weakly developed in
<taxonomicName id="4C5F4D09B657FFC0FEA2E05D2D8BBDA1" box="[308,396,621,644]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FEA2E05D2D8BBDA1" box="[308,396,621,644]" italics="true" pageId="7" pageNumber="136">Dictyna</emphasis>
</taxonomicName>
. Such a tooth is found not only in the
<typeStatus id="54E48828B657FFC0FCC3E05D2F82BDA0" box="[853,901,621,645]" pageId="7" pageNumber="136">type</typeStatus>
species, but well documented for
<taxonomicName id="4C5F4D09B657FFC0FA80E05D299BBDA1" box="[1302,1436,621,644]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="B92BEA98B657FFC0FA80E05D299BBDA1" box="[1302,1436,621,644]" italics="true" pageId="7" pageNumber="136">B. innocens</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCE4B7BB657FFC0FF08E0BF2D60BD8D" author="IJland" box="[158,359,655,680]" pageId="7" pageNumber="136" refString="IJland, S., van Helsdingen, P. J. &amp; Miller, J. (2012) On some spiders from Gargano, Apulia, Italy. Nieuwsbrief Spinnenwerkgroep Nederland, 32, 2 - 20." type="journal article" year="2012">
IJland
<emphasis id="B92BEA98B657FFC0FF7DE0A12D1CBD8D" box="[235,283,655,680]" italics="true" pageId="7" pageNumber="136">et al</emphasis>
. 2012
</bibRefCitation>
: fig. 4) and
<taxonomicName id="4C5F4D09B657FFC0FE7DE0A12E70BD8D" box="[491,631,655,680]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="guanchae">
<emphasis id="B92BEA98B657FFC0FE7DE0A12E70BD8D" box="[491,631,655,680]" italics="true" pageId="7" pageNumber="136">B. guanchae</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFCE4B7BB657FFC0FD13E0BF2FB7BD8D" author="Wunderlich" box="[645,944,655,680]" pageId="7" pageNumber="136" refString="Wunderlich, J. (1987) Die Spinnen der Kanarischen Inseln und Madeiras: Adaptive Radiation, Biogeographie, Revisionen und Neubeschreibungen. Taxonomy &amp; Ecology, 1, 1 - 435. [Verlag J. Wunderlich, Straubenhardt]" type="journal article" year="1987">Wunderlich 1987: fig. 590</bibRefCitation>
) as well.
<taxonomicName id="4C5F4D09B657FFC0FB8DE0A128CFBD8D" box="[1051,1224,655,680]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="latens">
<emphasis id="B92BEA98B657FFC0FB8DE0A128CFBD8D" box="[1051,1224,655,680]" italics="true" pageId="7" pageNumber="136">Brigittea latens</emphasis>
</taxonomicName>
has also cheliceral teeth (
<emphasis id="B92BEA98B657FFC0FF4BE0852CF0BDE9" box="[221,247,693,716]" italics="true" pageId="7" pageNumber="136">Ct</emphasis>
) lacking in
<taxonomicName id="4C5F4D09B657FFC0FEE8E0852E6DBDE8" box="[382,618,692,717]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="arundinacea">
<emphasis id="B92BEA98B657FFC0FEE8E0852E6DBDE8" box="[382,618,692,717]" italics="true" pageId="7" pageNumber="136">Dictyna arundinacea</emphasis>
</taxonomicName>
(cf.
<figureCitation id="13642A0FB657FFC0FD08E0842F19BDE9" box="[670,798,692,717]" captionStart="FIGURES 26 36" captionStartId="8.[151,264,1925,1948]" captionTargetBox="[259,1327,193,1888]" captionTargetId="figure@8.[248,1339,193,1893]" captionTargetPageId="8" captionText="FIGURES 26 36. Dictyna arundinacea (26 30), Brigittea latens (31 36). 26, 31 Male palp, ventral. 27, 32 Ctenidia, dorsoretrolateral. 28, 33 Chelicera, mesal. 29, 34 Tip of conductor, retrolateral. 30, 35 Same, ventral. 36 Tip of embolus. Abbreviations: Bt—retrobasal tooth of chelicera; Cr—cheliceral ridge; Ct—cheliceral teeth." httpUri="https://zenodo.org/record/243582/files/figure.png" pageId="7" pageNumber="136">Figs 28, 33</figureCitation>
). The cheliceral ridge (
<emphasis id="B92BEA98B657FFC0FBB4E0852847BDE9" box="[1058,1088,693,716]" italics="true" pageId="7" pageNumber="136">Cr</emphasis>
) of the
<typeStatus id="54E48828B657FFC0FB00E08528C1BDE8" box="[1174,1222,693,717]" pageId="7" pageNumber="136">type</typeStatus>
species of
<taxonomicName id="4C5F4D09B657FFC0FAD5E085299BBDE9" box="[1347,1436,693,716]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FAD5E085299BBDE9" box="[1347,1436,693,716]" italics="true" pageId="7" pageNumber="136">Dictyna</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D09B657FFC0FF5EE0E92D2BBDD5" box="[200,300,729,752]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FF5EE0E92D2BBDD5" box="[200,300,729,752]" italics="true" pageId="7" pageNumber="136">Brigittea</emphasis>
</taxonomicName>
also differs in its length. Also, the profile of the chelicera in
<taxonomicName id="4C5F4D09B657FFC0FC7AE0E92842BDD5" box="[1004,1093,729,752]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FC7AE0E92842BDD5" box="[1004,1093,729,752]" italics="true" pageId="7" pageNumber="136">Dictyna</emphasis>
</taxonomicName>
is strongly concave (
<figureCitation id="13642A0FB657FFC0FAACE0E72988BDD5" box="[1338,1423,727,752]" captionStart="FIGURES 26 36" captionStartId="8.[151,264,1925,1948]" captionTargetBox="[259,1327,193,1888]" captionTargetId="figure@8.[248,1339,193,1893]" captionTargetPageId="8" captionText="FIGURES 26 36. Dictyna arundinacea (26 30), Brigittea latens (31 36). 26, 31 Male palp, ventral. 27, 32 Ctenidia, dorsoretrolateral. 28, 33 Chelicera, mesal. 29, 34 Tip of conductor, retrolateral. 30, 35 Same, ventral. 36 Tip of embolus. Abbreviations: Bt—retrobasal tooth of chelicera; Cr—cheliceral ridge; Ct—cheliceral teeth." httpUri="https://zenodo.org/record/243582/files/figure.png" pageId="7" pageNumber="136">Fig. 28</figureCitation>
), while in
<taxonomicName id="4C5F4D09B657FFC0FF6FE0CD2D5ABC31" box="[249,349,765,788]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FF6FE0CD2D5ABC31" box="[249,349,765,788]" italics="true" pageId="7" pageNumber="136">Brigittea</emphasis>
</taxonomicName>
it is almost straight (
<figureCitation id="13642A0FB657FFC0FDDEE0CC2E9DBC31" box="[584,666,764,789]" captionStart="FIGURES 26 36" captionStartId="8.[151,264,1925,1948]" captionTargetBox="[259,1327,193,1888]" captionTargetId="figure@8.[248,1339,193,1893]" captionTargetPageId="8" captionText="FIGURES 26 36. Dictyna arundinacea (26 30), Brigittea latens (31 36). 26, 31 Male palp, ventral. 27, 32 Ctenidia, dorsoretrolateral. 28, 33 Chelicera, mesal. 29, 34 Tip of conductor, retrolateral. 30, 35 Same, ventral. 36 Tip of embolus. Abbreviations: Bt—retrobasal tooth of chelicera; Cr—cheliceral ridge; Ct—cheliceral teeth." httpUri="https://zenodo.org/record/243582/files/figure.png" pageId="7" pageNumber="136">Fig. 33</figureCitation>
) or convex (pl. IV, fig.
<quantity id="4CA79B6FB657FFC0FC09E0CC2FC2BC30" box="[927,965,764,789]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="136" unit="in" value="1.0">1 in</quantity>
<bibRefCitation id="EFCE4B7BB657FFC0FC5AE0CC28C0BC31" author="Miller" box="[972,1223,764,789]" pageId="7" pageNumber="136" refString="Miller, F. &amp; Svaton, J. (1978) Einige seltene und bisher unbekannte Spinnenarten aus der Slowakei. Annotationes Zoologicae et Botanicae, 126, 1 - 19." type="journal article" year="1978">Miller &amp; Svatoň 1978</bibRefCitation>
).
</paragraph>
<paragraph id="8BE0368AB657FFC0FF51E12F2E99BB11" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="136">
The
<typeStatus id="54E48828B657FFC0FF6CE1102D2DBC1D" box="[250,298,800,824]" pageId="7" pageNumber="136">type</typeStatus>
species of both genera differ essentially in their structure of the ctenidia. In
<emphasis id="B92BEA98B657FFC0FB1DE11128EDBC1D" box="[1163,1258,801,824]" italics="true" pageId="7" pageNumber="136">
<taxonomicName id="4C5F4D09B657FFC0FB1DE11128E0BC1D" box="[1163,1255,801,824]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">Dictyna</taxonomicName>
,
</emphasis>
ctenidia are set in separate alveoli, while in
<taxonomicName id="4C5F4D09B657FFC0FE4EE1752E3BBC79" box="[472,572,837,860]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FE4EE1752E3BBC79" box="[472,572,837,860]" italics="true" pageId="7" pageNumber="136">Brigittea</emphasis>
</taxonomicName>
, they are located in the same alveolus. The ctenidia in
<taxonomicName id="4C5F4D09B657FFC0FB38E1752958BC78" box="[1198,1375,836,861]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="arundinacea">
<emphasis id="B92BEA98B657FFC0FB38E1752958BC78" box="[1198,1375,836,861]" italics="true" pageId="7" pageNumber="136">D. arundinacea</emphasis>
</taxonomicName>
have distinct micro-ridges which are lacking in
<taxonomicName id="4C5F4D09B657FFC0FDF9E1592ED7BCA5" box="[623,720,871,896]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="latens">
<emphasis id="B92BEA98B657FFC0FDF9E1592ED7BCA5" box="[623,720,871,896]" italics="true" pageId="7" pageNumber="136">B. latens</emphasis>
</taxonomicName>
. The tip of the lower arm of the conductor is also different in
<taxonomicName id="4C5F4D09B657FFC0FA17E1592D22BC80" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="arundinacea">
<emphasis id="B92BEA98B657FFC0FA17E1592D22BC80" italics="true" pageId="7" pageNumber="136">D. arundinacea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D09B657FFC0FECBE1BD2DC7BC80" box="[349,448,908,933]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="latens">
<emphasis id="B92BEA98B657FFC0FECBE1BD2DC7BC80" box="[349,448,908,933]" italics="true" pageId="7" pageNumber="136">B. latens</emphasis>
</taxonomicName>
, but this difference is not important as the shape of this tip in
<taxonomicName id="4C5F4D09B657FFC0FB13E1BD28D9BC81" box="[1157,1246,909,932]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FB13E1BD28D9BC81" box="[1157,1246,909,932]" italics="true" pageId="7" pageNumber="136">Dictyna</emphasis>
</taxonomicName>
is very variable. More important differences are found in the shape of embolus. In
<emphasis id="B92BEA98B657FFC0FCEFE1812FDFBCED" box="[889,984,945,968]" italics="true" pageId="7" pageNumber="136">
<taxonomicName id="4C5F4D09B657FFC0FCEFE1812FD2BCED" box="[889,981,945,968]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">Dictyna</taxonomicName>
,
</emphasis>
it is filamentous and the tip is gradually tapering, while in
<taxonomicName id="4C5F4D09B657FFC0FEF7E1E52DC5BCC8" box="[353,450,980,1005]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="latens">
<emphasis id="B92BEA98B657FFC0FEF7E1E52DC5BCC8" box="[353,450,980,1005]" italics="true" pageId="7" pageNumber="136">B. latens</emphasis>
</taxonomicName>
it is modified (
<figureCitation id="13642A0FB657FFC0FDFAE1E42EB8BCC8" box="[620,703,980,1005]" captionStart="FIGURES 26 36" captionStartId="8.[151,264,1925,1948]" captionTargetBox="[259,1327,193,1888]" captionTargetId="figure@8.[248,1339,193,1893]" captionTargetPageId="8" captionText="FIGURES 26 36. Dictyna arundinacea (26 30), Brigittea latens (31 36). 26, 31 Male palp, ventral. 27, 32 Ctenidia, dorsoretrolateral. 28, 33 Chelicera, mesal. 29, 34 Tip of conductor, retrolateral. 30, 35 Same, ventral. 36 Tip of embolus. Abbreviations: Bt—retrobasal tooth of chelicera; Cr—cheliceral ridge; Ct—cheliceral teeth." httpUri="https://zenodo.org/record/243582/files/figure.png" pageId="7" pageNumber="136">Fig. 36</figureCitation>
). This character was overlooked by
<bibRefCitation id="EFCE4B7BB657FFC0FBC6E1E42903BCC8" author="Lehtinen" box="[1104,1284,980,1005]" pageId="7" pageNumber="136" refString="Lehtinen, P. T. (1967) Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici, 4, 199 - 468." type="journal article" year="1967">Lehtinen (1967)</bibRefCitation>
and
<bibRefCitation id="EFCE4B7BB657FFC0FAACE1E42D3ABB35" author="Miller" pageId="7" pageNumber="136" refString="Miller, F. &amp; Svaton, J. (1978) Einige seltene und bisher unbekannte Spinnenarten aus der Slowakei. Annotationes Zoologicae et Botanicae, 126, 1 - 19." type="journal article" year="1978">Miller &amp; Svatoň (1978)</bibRefCitation>
. A modified embolus might prove to be the “best” character that separates the two very similar genera
<taxonomicName id="4C5F4D09B657FFC0FF7EE62D2D46BB11" box="[232,321,1053,1076]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FF7EE62D2D46BB11" box="[232,321,1053,1076]" italics="true" pageId="7" pageNumber="136">Dictyna</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D09B657FFC0FEE1E62C2E90BB10" authority="Lehtinen 1967" authorityName="Lehtinen" authorityYear="1967" box="[375,663,1052,1077]" class="Arachnida" family="Dictynidae" genus="Emblyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FEE1E62C2DDCBB10" box="[375,475,1052,1077]" italics="true" pageId="7" pageNumber="136">Emblyna</emphasis>
(
<bibRefCitation id="EFCE4B7BB657FFC0FE7FE62C2E88BB10" author="Lehtinen" box="[489,655,1052,1077]" pageId="7" pageNumber="136" refString="Lehtinen, P. T. (1967) Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici, 4, 199 - 468." type="journal article" year="1967">Lehtinen 1967</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BE0368AB657FFC0FF51E60F28AABBC2" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="136">
Equally, the epigynes of
<taxonomicName id="4C5F4D09B657FFC0FE4AE6712E8BBB7D" box="[476,652,1087,1112]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="arundinacea">
<emphasis id="B92BEA98B657FFC0FE4AE6712E8BBB7D" box="[476,652,1087,1112]" italics="true" pageId="7" pageNumber="136">D. arundinacea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D09B657FFC0FD55E6712F21BB7D" box="[707,806,1087,1112]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="latens">
<emphasis id="B92BEA98B657FFC0FD55E6712F21BB7D" box="[707,806,1087,1112]" italics="true" pageId="7" pageNumber="136">B. latens</emphasis>
</taxonomicName>
are entirely different in all respects.
<taxonomicName id="4C5F4D09B657FFC0FB54E6712968BB7D" box="[1218,1391,1087,1112]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="latens">
<emphasis id="B92BEA98B657FFC0FB54E6712968BB7D" box="[1218,1391,1087,1112]" italics="true" pageId="7" pageNumber="136">Brigittea latens</emphasis>
</taxonomicName>
has no epigynal sulci, which are present in all
<emphasis id="B92BEA98B657FFC0FDE7E6552F5ABB59" box="[625,861,1125,1148]" italics="true" pageId="7" pageNumber="136">
<taxonomicName id="4C5F4D09B657FFC0FDE7E6552F08BB59" box="[625,783,1125,1148]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="sensu">
Dictyna
<taxonomicNameLabel id="A21857E3B657FFC0FD47E6552F08BB59" box="[721,783,1125,1148]" pageId="7" pageNumber="136" sensu="stricto">sensu</taxonomicNameLabel>
</taxonomicName>
stricto
</emphasis>
.
<taxonomicName id="4C5F4D09B657FFC0FCFAE655281DBB59" box="[876,1050,1125,1148]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="civica">
<emphasis id="B92BEA98B657FFC0FCFAE655281DBB59" box="[876,1050,1125,1148]" italics="true" pageId="7" pageNumber="136">Brigittea civica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D09B657FFC0FBC6E65528B3BB59" box="[1104,1204,1125,1148]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B92BEA98B657FFC0FBC6E65528B3BB59" box="[1104,1204,1125,1148]" italics="true" pageId="7" pageNumber="136">B. vicina</emphasis>
</taxonomicName>
have sulci, although they are much shorter than in
<taxonomicName id="4C5F4D09B657FFC0FE73E6B92E39BB85" box="[485,574,1161,1184]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FE73E6B92E39BB85" box="[485,574,1161,1184]" italics="true" pageId="7" pageNumber="136">Dictyna</emphasis>
</taxonomicName>
. The most significant differences lie in the shape of “receptacles”, which are contiguous in
<taxonomicName id="4C5F4D09B657FFC0FEA8E69D2D90BBE1" box="[318,407,1197,1220]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FEA8E69D2D90BBE1" box="[318,407,1197,1220]" italics="true" pageId="7" pageNumber="136">Dictyna</emphasis>
</taxonomicName>
and forming a V-shaped structure together with the ducts; in contrast, in
<emphasis id="B92BEA98B657FFC0FA92E69D2969BBE1" box="[1284,1390,1197,1220]" italics="true" pageId="7" pageNumber="136">
<taxonomicName id="4C5F4D09B657FFC0FA92E69D296DBBE1" box="[1284,1386,1197,1220]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">Brigittea</taxonomicName>
,
</emphasis>
the receptacles do not touch each other and together with ducts form a spiraled (coiled) structure.
</paragraph>
<paragraph id="8BE0368AB657FFC0FF51E6C4291BBA71" blockId="7.[151,1437,151,1365]" pageId="7" pageNumber="136">
Based on the reasons mentioned above, we decided to resurrect
<taxonomicName id="4C5F4D09B657FFC0FC2DE6C52818BA29" box="[955,1055,1269,1292]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FC2DE6C52818BA29" box="[955,1055,1269,1292]" italics="true" pageId="7" pageNumber="136">Brigittea</emphasis>
</taxonomicName>
as a valid genus with following species:
<taxonomicName id="4C5F4D09B657FFC0FF61E7292D5DBA15" box="[247,346,1303,1328]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="latens">
<emphasis id="B92BEA98B657FFC0FF61E7292D5DBA15" box="[247,346,1303,1328]" italics="true" pageId="7" pageNumber="136">B. latens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5F4D09B657FFC0FEFFE7292DCCBA15" box="[361,459,1305,1328]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="civica">
<emphasis id="B92BEA98B657FFC0FEFFE7292DCCBA15" box="[361,459,1305,1328]" italics="true" pageId="7" pageNumber="136">B. civica</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5F4D09B657FFC0FE4CE7292E5ABA15" box="[474,605,1305,1328]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="B92BEA98B657FFC0FE4CE7292E5ABA15" box="[474,605,1305,1328]" italics="true" pageId="7" pageNumber="136">B. innocens</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5F4D09B657FFC0FDFAE7292ED6BA15" box="[620,721,1305,1328]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="vicina">
<emphasis id="B92BEA98B657FFC0FDFAE7292ED6BA15" box="[620,721,1305,1328]" italics="true" pageId="7" pageNumber="136">B. vicina</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5F4D09B657FFC0FC9EE7292852BA15" authority="Schmidt, 1968" authorityName="Schmidt" authorityYear="1968" box="[776,1109,1303,1328]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="guanchae" status="comb. nov.">
<emphasis id="B92BEA98B657FFC0FC9EE7292F92BA15" box="[776,917,1303,1328]" italics="true" pageId="7" pageNumber="136">B. guanchae</emphasis>
(Schmidt, 1968)
</taxonomicName>
,
<taxonomicNameLabel id="A21857E3B657FFC0FBF4E72728CEBA15" box="[1122,1225,1303,1328]" pageId="7" pageNumber="136">
<emphasis id="B92BEA98B657FFC0FBF4E72728CEBA15" bold="true" box="[1122,1225,1303,1328]" pageId="7" pageNumber="136">comb. n.</emphasis>
</taxonomicNameLabel>
(ex
<taxonomicName id="4C5F4D09B657FFC0FB6AE7292952BA15" box="[1276,1365,1305,1328]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FB6AE7292952BA15" box="[1276,1365,1305,1328]" italics="true" pageId="7" pageNumber="136">Dictyna</emphasis>
</taxonomicName>
) from the Canary Islands. The latter species was attributed to the
<taxonomicName id="4C5F4D09B657FFC0FCBDE70D2FC9BA70" box="[811,974,1340,1365]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="latens">
<emphasis id="B92BEA98B657FFC0FCBDE70D2FC9BA70" box="[811,974,1340,1365]" italics="true" pageId="7" pageNumber="136">Dictyna latens</emphasis>
</taxonomicName>
group by
<bibRefCitation id="EFCE4B7BB657FFC0FBD6E70C2910BA71" author="Wunderlich" box="[1088,1303,1340,1365]" pageId="7" pageNumber="136" refString="Wunderlich, J. (1987) Die Spinnen der Kanarischen Inseln und Madeiras: Adaptive Radiation, Biogeographie, Revisionen und Neubeschreibungen. Taxonomy &amp; Ecology, 1, 1 - 435. [Verlag J. Wunderlich, Straubenhardt]" type="journal article" year="1987">Wunderlich (1987)</bibRefCitation>
.
</paragraph>
<paragraph id="8BE0368AB657FFC0FF01E7B52F72BAC0" blockId="7.[151,1436,1412,1509]" pageId="7" pageNumber="136">
Four species from Afrotropical and South Asian regions considered in
<taxonomicName id="4C5F4D09B657FFC0FC20E7B5281DBAB9" box="[950,1050,1413,1436]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FC20E7B5281DBAB9" box="[950,1050,1413,1436]" italics="true" pageId="7" pageNumber="136">Brigittea</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFCE4B7BB657FFC0FBD3E7B428FCBAB8" author="Lehtinen" box="[1093,1275,1412,1437]" pageId="7" pageNumber="136" refString="Lehtinen, P. T. (1967) Classification of the cribellate spiders and some allied families, with notes on the evolution of the suborder Araneomorpha. Annales Zoologici Fennici, 4, 199 - 468." type="journal article" year="1967">Lehtinen (1967)</bibRefCitation>
(
<emphasis id="B92BEA98B657FFC0FA9AE7B52D7CBAE5" italics="true" pageId="7" pageNumber="136">
<taxonomicName id="4C5F4D09B657FFC0FA9AE7B52972BAB8" box="[1292,1397,1412,1437]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="denisi">D. denisi</taxonomicName>
,
<taxonomicName id="4C5F4D09B657FFC0FA17E7B52CEBBAE5" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="turbida">D. turbida</taxonomicName>
,
<taxonomicName id="4C5F4D09B657FFC0FF6EE7992D7CBAE5" box="[248,379,1447,1472]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="tullgreni">D. tullgreni</taxonomicName>
</emphasis>
and
<taxonomicName id="4C5F4D09B657FFC0FE24E7992E0BBAE5" box="[434,524,1449,1472]" class="Arachnida" family="Dictynidae" genus="Brigittea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="species" species="umai">
<emphasis id="B92BEA98B657FFC0FE24E7992E0BBAE5" box="[434,524,1449,1472]" italics="true" pageId="7" pageNumber="136">D. umai</emphasis>
</taxonomicName>
) remain in
<taxonomicName id="4C5F4D09B657FFC0FD06E7992EEEBAE5" box="[656,745,1449,1472]" class="Arachnida" family="Dictynidae" genus="Dictyna" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="136" phylum="Arthropoda" rank="genus">
<emphasis id="B92BEA98B657FFC0FD06E7992EEEBAE5" box="[656,745,1449,1472]" italics="true" pageId="7" pageNumber="136">Dictyna</emphasis>
</taxonomicName>
. These species are not properly known and their descriptions lack information about a divided cribellum or modified chelicera.
</paragraph>
</subSubSection>
</treatment>
</document>