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<document id="2849D0242DEFFCC91058280845640338" ID-CLB-Dataset="58541" ID-DOI="10.5852/ejt.2023.879.2155" ID-GBIF-Dataset="67c69252-9e0f-49b8-b7c7-ce62d25e9251" ID-ISSN="2118-9773" ID-Zenodo-Dep="8135424" ID-ZooBank="02FF7797-1F28-4782-B9F6-24CE9A4993D3" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1689079849653" checkinUser="felipe" docAuthor="Bitzilekis, Eleftherios, Vardinoyannis, Katerina &amp; Mylonas, Moisis" docDate="2023" docId="03F68790FFD4FF97FE23FECCFE69FA2C" docLanguage="en" docName="ejt-879_1-23.pdf" docOrigin="European Journal of Taxonomy 879 (1)" docSource="http://dx.doi.org/10.5852/ejt.2023.879.2155" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Metafruticicola coartata subsp. gemina Fuchs &amp; Kaufel 1936" docType="treatment" docVersion="6" lastPageNumber="17" masterDocId="FFCFFFE8FFC4FF87FFF3FFC4FFC5FFB5" masterDocTitle="A new species of Metafruticicola (Gastropoda, Hygromiidae) from the Aegean Archipelago and new anatomical and distributional data on some congeneric species" masterLastPageNumber="23" masterPageNumber="1" pageNumber="17" updateTime="1699536541208" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="7786E2A111A9AD8D1352D5CCD6FF23E4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9FD112341CCD7F1891A7683AD4B13517">
<mods:title id="079CCD805EEF827A7770695D93DA4EE4">A new species of Metafruticicola (Gastropoda, Hygromiidae) from the Aegean Archipelago and new anatomical and distributional data on some congeneric species</mods:title>
</mods:titleInfo>
<mods:name id="EE8C2C89F9625D669D2D52A0904A7C04" type="personal">
<mods:role id="16670959132EABC104A84B006F9BFF0E">
<mods:roleTerm id="76BB310B67EC22C66F8AE5ADA7151510">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="00C8417A9E9390FB08FCB9A98E9ABC6C">Bitzilekis, Eleftherios</mods:namePart>
<mods:nameIdentifier id="2DCFCD9B6169F3AAFEBCB41D18C65940" type="LSID">599CF3C1-6866-472D-AE73-DA281CC99E5D</mods:nameIdentifier>
<mods:affiliation id="5870CBF1804F85BA503DC4647CAA2FE4">Natural History Museum of Crete, School of Sciences and Engineering, University of Crete, 71409 Irakleio, Greece. &amp; Biology Department, School of Sciences and Engineering, University of Crete, Voutes University Campus, 70013 Irakleio, Greece.</mods:affiliation>
<mods:nameIdentifier id="AFB7A39F7F017F228CCEA60C79EF7545" type="email">ebitzi@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="E8EB660AD12C3752B3C31A30843A242A" type="personal">
<mods:role id="B361E40EAE538042EC29D7040F0BDDAA">
<mods:roleTerm id="57CD572DD7E55B16538ED174678FDA5F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DADB4852E4149A73B5C64A19060F5926">Vardinoyannis, Katerina</mods:namePart>
<mods:nameIdentifier id="710C3700A03DA20499D2340704F10764" type="LSID">CFD1E53C-B8A2-44E4-8E61-A0664CDD1067</mods:nameIdentifier>
<mods:affiliation id="101A44FDE5EB35BFB245211A77809631">Natural History Museum of Crete, School of Sciences and Engineering, University of Crete, 71409 Irakleio, Greece.</mods:affiliation>
<mods:nameIdentifier id="7070D3ECCDDF36EC69D6DCCFAEE40EE3" type="email">mollusca@nhmc.uoc.gr</mods:nameIdentifier>
</mods:name>
<mods:name id="DF619350848D400CF6E843D8180D80C4" type="personal">
<mods:role id="D937F6EF24024C061B4AB3EA4167510C">
<mods:roleTerm id="80516E9ADF27F8517772A47EC3ADCD4F">Author</mods:roleTerm>
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<mods:namePart id="ACD001C17ACD7CFA22F296793D2CA005">Mylonas, Moisis</mods:namePart>
<mods:nameIdentifier id="7E9D2266890228B60DEB8E014B3F6DE1" type="LSID">5AF8941E-C2E3-4EA6-8002-9344BE3C6BA9</mods:nameIdentifier>
<mods:affiliation id="11C2A44F1B860A6589BEB6F271E5320E">Natural History Museum of Crete, School of Sciences and Engineering, University of Crete, 71409 Irakleio, Greece. &amp; Biology Department, School of Sciences and Engineering, University of Crete, Voutes University Campus, 70013 Irakleio, Greece.</mods:affiliation>
<mods:nameIdentifier id="335EC684AA7574B43945A9A32DB2E68D" type="email">mylonas@nhmc.uoc.gr</mods:nameIdentifier>
</mods:name>
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<mods:title id="7C1D0D288B10F41C4328BE4CB4B74716">European Journal of Taxonomy</mods:title>
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<mods:part id="EEE3D3154773ABDB3C91B00643532B3B">
<mods:date id="0649FE5A1E222733E08759F214F8213C">2023</mods:date>
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<mods:number id="6A6F861101526D1B9DFFC9843A5E4B38">2023-07-04</mods:number>
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<mods:number id="8D1BEFABE45393A4A60FB60ECC6016BB">879</mods:number>
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<mods:url id="04067F3E1095F8BFDFBE18EDBBD5EB03">http://dx.doi.org/10.5852/ejt.2023.879.2155</mods:url>
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<mods:classification id="DC58A01DF2984242E22148DB96F5F9B6">journal article</mods:classification>
<mods:identifier id="B241F74ABED23F370DA0870115B81DF8" type="CLB-Dataset">58541</mods:identifier>
<mods:identifier id="DC998C7432F28B216B5C23BFDEFDD3E9" type="DOI">10.5852/ejt.2023.879.2155</mods:identifier>
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<treatment id="03F68790FFD4FF97FE23FECCFE69FA2C" ID-DOI="http://doi.org/10.5281/zenodo.8135451" ID-GBIF-Taxon="211051659" ID-Zenodo-Dep="8135451" LSID="urn:lsid:plazi:treatment:03F68790FFD4FF97FE23FECCFE69FA2C" httpUri="http://treatment.plazi.org/id/03F68790FFD4FF97FE23FECCFE69FA2C" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="C345650DFFD4FF97FE23FECCFBA1FE96" box="[464,1124,264,291]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BE03686FFD4FF97FE23FECCFBA1FE96" blockId="16.[464,1124,264,327]" box="[464,1124,264,291]" pageId="16" pageNumber="17">
<heading id="D0A881EAFFD4FF97FE23FECCFBA1FE96" box="[464,1124,264,291]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
<taxonomicName id="4C5F4D05FFD4FF97FE23FECCFBA1FE96" ID-CoL="7HGBS" authority="Fuchs &amp; Kaufel, 1936" authorityName="Fuchs &amp; Kaufel" authorityYear="1936" box="[464,1124,264,291]" class="Gastropoda" family="Hygromiidae" genus="Metafruticicola" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="subSpecies" species="coartata" subSpecies="gemina">
<emphasis id="B92BEA94FFD4FF97FE23FECCFC9DFE96" bold="true" box="[464,856,264,291]" italics="true" pageId="16" pageNumber="17">Metafruticicola coartata gemina</emphasis>
<bibRefCitation id="EFCE4B77FFD4FF97FCACFECDFBA1FE96" author="Fuchs A. &amp; Kaufel F." box="[863,1124,265,291]" pageId="16" pageNumber="17" pagination="541 - 662" refId="ref12029" refString="Fuchs A. &amp; Kaufel F. 1936. Anatomische und systematische Untersuchungen an Land- und Susswasserschnecken aus Griechenland und von den Inseln des Agaischen Meeres. Zeitschrift fur wissenschaftliche Zoologie, Abteilung B. Archiv fur Naturgeschichte (Zeitschrift fur systematische Zoologie) 5 (4): 541 - 662." type="journal article" year="1936">Fuchs &amp; Käufel, 1936</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C345650DFFD4FF97FD22FEE9FCA6FEF2" box="[721,867,301,327]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BE03686FFD4FF97FD22FEE9FCA6FEF2" blockId="16.[464,1124,264,327]" box="[721,867,301,327]" pageId="16" pageNumber="17">
<figureCitation id="13642A03FFD4FF97FD22FEE9FC8DFEF2" box="[721,840,301,327]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[262,1325,265,1852]" captionTargetId="figure-12@17.[262,1325,265,1852]" captionTargetPageId="17" captionText="Fig. 7. Metafruticicola (Westerlundia) coartata. A. M. (Westerlundia) c. coartata Fuchs &amp; Käufel, 1936, Greece, Cyclades, Makra Islet (NHMC 50.4056). Shell. B–D. M. (Westerlundia) c. gemina Fuchs &amp; Käufel, 1936, Greece, Dodecanese, Karavia Nisia, north Islet (NHMC 50.6805). B. Shell. C. General view of genitalia. D. View of penial papilla. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.8135445" httpUri="https://zenodo.org/record/8135445/files/figure.png" pageId="16" pageNumber="17">Fig 7B–D</figureCitation>
,
<figureCitation id="13642A03FFD4FF97FCA7FEE9FCA6FEF2" box="[852,867,301,327]" captionStart="Fig" captionStartId="18.[189,232,1752,1778]" captionTargetBox="[189,1398,644,1708]" captionTargetId="figure-184@18.[189,1398,644,1708]" captionTargetPageId="18" captionText="Fig. 8. Distribution of Metafruticicola (Westerlundia) coartata Fuchs &amp; Käufel, 1936. Known (black) and new (red) distributional records on the presence of the species in various Aegean islands. —Greece, Cyclades: 1. Christiani; 2. Askania; 3. Thirasia; 4. Anafi; 5. Fteno; 6. Pacheia; 7. Makra; 8. Anydros; 9. Mikros Avelas; 10. Machaires; 11. Gramvousa; 12. Donousa; 13. Liadi. — Greece, Dodecanese: 14 Kinaros; 15. Megalo &amp; Mikro Mavro; 16. Levitha; 17. Safonidi; 18. Ofidousa; 19. Pontikousa; 20. Fokionisia; 21. Astypalaia; 22. Chondros; 23. Lianos; 24. Agia Kyriaki; 25. Tigani; 26. Fteno; 27. Kounoupoi; 28. Koutsomytis; 29. Mikros &amp; Megalos Aderfos; 30. Syrna; 31. Stefania; 32. Mesonisi; 33. Megali Zafora; 34. Sochas; 35. Mikri Zafora; 36. Karavi North; 37. Ounio West; 38. Ounio East." figureDoi="http://doi.org/10.5281/zenodo.8135449" httpUri="https://zenodo.org/record/8135449/files/figure.png" pageId="16" pageNumber="17">8</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C345650DFFD4FF97FF4EFEAAFCF9FDDB" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BE03686FFD4FF97FF4EFEAAFE6FFE3D" blockId="16.[189,426,366,392]" box="[189,426,366,392]" pageId="16" pageNumber="17">
<heading id="D0A881EAFFD4FF97FF4EFEAAFE6FFE3D" bold="true" box="[189,426,366,392]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B92BEA94FFD4FF97FF4EFEAAFE6FFE3D" bold="true" box="[189,426,366,392]" pageId="16" pageNumber="17">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BE03686FFD4FF97FF4EFE59FBEEFE4B" blockId="16.[189,1399,412,510]" pageId="16" pageNumber="17">
<materialsCitation id="3B373CDBFFD4FF97FF4EFE59FBF1FE6E" ID-GBIF-Occurrence="4148095310" collectingDate="1989-04-21" collectionCode="NHMC" collectorName="M. Mylonas" country="Greece" latitude="36.0006" location="Karavia Nisia" longLatPrecision="6" longitude="26.4354" pageId="16" pageNumber="17" specimenCode="NHMC 50.6805" specimenCount="16" stateProvince="Dodecanese">
<collectingCountry id="F3487616FFD4FF97FF4EFE59FEEAFE02" box="[189,303,413,439]" name="Greece" pageId="16" pageNumber="17">GREECE</collectingCountry>
–
<emphasis id="B92BEA94FFD4FF97FEA0FE59FE23FE02" bold="true" box="[339,486,413,439]" pageId="16" pageNumber="17">
<collectingRegion id="499BF864FFD4FF97FEA0FE59FE23FE02" box="[339,486,413,439]" pageId="16" pageNumber="17">Dodecanese</collectingRegion>
</emphasis>
• 7 shells, 9 specs in 75% alcohol, 4 dissections;
<location id="8E80605DFFD4FF97FBB2FE59FB2DFE02" LSID="urn:lsid:plazi:treatment:03F68790FFD4FF97FE23FECCFE69FA2C:8E80605DFFD4FF97FBB2FE59FB2DFE02" box="[1089,1256,413,439]" country="Greece" latitude="36.0006" longLatPrecision="6" longitude="26.4354" name="Karavia Nisia" pageId="16" pageNumber="17" stateProvince="Dodecanese">Karavia Nisia</location>
, north islet;
<geoCoordinate id="EE6B5041FFD4FF97FF4EFE04FE8CFE6F" box="[189,329,448,474]" degrees="36.0006" direction="north" orientation="latitude" pageId="16" pageNumber="17" precision="5" value="36.0006">36.0006° N</geoCoordinate>
,
<geoCoordinate id="EE6B5041FFD4FF97FEA4FE04FE25FE6F" box="[343,480,448,475]" degrees="26.4354" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="5" value="26.4354">26.4354° E</geoCoordinate>
;
<date id="FFE11046FFD4FF97FE1CFE04FD4BFE6F" box="[495,654,448,474]" pageId="16" pageNumber="17" value="1989-04-21">
<collectingDate id="EFA5E9AEFFD4FF97FE1CFE04FD4BFE6F" box="[495,654,448,474]" pageId="16" pageNumber="17" value="1989-04-21">21 Apr. 1989</collectingDate>
</date>
;
<collectorName id="26AA5350FFD4FF97FD6FFE04FCEBFE6F" box="[668,814,448,474]" pageId="16" pageNumber="17">M. Mylonas</collectorName>
leg.;
<specimenCode id="DBF99EFDFFD4FF97FC87FE04FBF1FE6E" box="[884,1076,448,475]" collectionCode="NHMC" country="Myanmar" httpUri="http://grbio.org/cool/4819-0jtd" name="Natural History Museum, Rangoon" pageId="16" pageNumber="17">NHMC 50.6805</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B373CDBFFD4FF97FBA2FE04FBE2FE4B" ID-GBIF-Occurrence="4148095381" collectingDate="2018-05-16" collectionCode="NHMC" collectorName="P. Lymberakis" country="Greece" latitude="36.0006" location="Karavia Nisia" longLatPrecision="6" longitude="26.4354" pageId="16" pageNumber="17" specimenCode="NHMC 50.20961" specimenCount="6" stateProvince="Dodecanese">
6 shells; same collection data as for preceding;
<date id="FFE11046FFD4FF97FE31FE27FDA4FE48" box="[450,609,483,510]" pageId="16" pageNumber="17" value="2018-05-16">
<collectingDate id="EFA5E9AEFFD4FF97FE31FE27FDA4FE48" box="[450,609,483,510]" pageId="16" pageNumber="17" value="2018-05-16">16 May 2018</collectingDate>
</date>
;
<collectorName id="26AA5350FFD4FF97FD9DFE20FCD1FE48" box="[622,788,483,510]" pageId="16" pageNumber="17">P. Lymberakis</collectorName>
leg.;
<specimenCode id="DBF99EFDFFD4FF97FCA5FE27FBE2FE4B" box="[854,1063,483,510]" collectionCode="NHMC" country="Myanmar" httpUri="http://grbio.org/cool/4819-0jtd" name="Natural History Museum, Rangoon" pageId="16" pageNumber="17">NHMC 50.20961</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE03686FFD4FF97FF4EFDE1FEACFD8B" blockId="16.[189,361,548,575]" box="[189,361,548,575]" pageId="16" pageNumber="17">
<heading id="D0A881EAFFD4FF97FF4EFDE1FEACFD8B" bold="true" box="[189,361,548,575]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B92BEA94FFD4FF97FF4EFDE1FEACFD8B" bold="true" box="[189,361,548,575]" pageId="16" pageNumber="17">Locus typicus</emphasis>
</heading>
</paragraph>
<paragraph id="8BE03686FFD4FF97FF4EFD97FCF9FDDB" blockId="16.[189,828,595,622]" box="[189,828,595,622]" pageId="16" pageNumber="17">
<collectingCountry id="F3487616FFD4FF97FF4EFD97FED6FDD8" box="[189,275,595,621]" name="Greece" pageId="16" pageNumber="17">Greece</collectingCountry>
, Karavia Nisia, northern larger islet of the two.
</paragraph>
</subSubSection>
<subSubSection id="C345650DFFD4FF97FF4EFD50FAC7FBE2" pageId="16" pageNumber="17" type="description">
<paragraph id="8BE03686FFD4FF97FF4EFD50FE8BFD1B" blockId="16.[189,334,660,686]" box="[189,334,660,686]" pageId="16" pageNumber="17">
<heading id="D0A881EAFFD4FF97FF4EFD50FE8BFD1B" bold="true" box="[189,334,660,686]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B92BEA94FFD4FF97FF4EFD50FE8BFD1B" bold="true" box="[189,334,660,686]" pageId="16" pageNumber="17">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BE03686FFD4FF97FF4EFD07FCBBFD68" blockId="16.[189,894,707,734]" box="[189,894,707,734]" pageId="16" pageNumber="17">
<smallCapsWord id="8D06A05AFFD4FF97FF4EFD07FEC7FD69" baselines="726,727" box="[189,258,707,733]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="16" pageNumber="17">SHELL</smallCapsWord>
. Shell description is given by
<bibRefCitation id="EFCE4B77FFD4FF97FD94FD07FCBFFD68" author="Bank R. A. &amp; Gittenberger E. &amp; Neubert E." box="[615,890,707,734]" pageId="16" pageNumber="17" pagination="67 - 136" refId="ref11778" refString="Bank R. A., Gittenberger E. &amp; Neubert E. 2013. Radiation of an eastern Mediterranean landsnail genus: revision of the taxa belonging to Metafruticicola von Ihering, 1892 (Gastropoda, Pulmonata: Hygromiidae). Archiv fur Molluskenkunde 142 (1): 67 - 136. https: // doi. org / 10.1127 / arch. moll / 1869 - 0963 / 142 / 067 - 136" type="journal article" year="2013">
Bank
<emphasis id="B92BEA94FFD4FF97FD5FFD00FD1AFD68" box="[684,735,707,733]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
. (2013: 119)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BE03686FFD4FF97FF4EFCC1FBC7FCF7" blockId="16.[187,1399,772,834]" pageId="16" pageNumber="17">
<smallCapsWord id="8D06A05AFFD4FF97FF4EFCC1FEC7FCA8" baselines="792,792" box="[189,258,773,799]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="16" pageNumber="17">SHELL</smallCapsWord>
<smallCapsWord id="8D06A05AFFD4FF97FEFBFCCEFE4DFCA8" baselines="792" box="[264,392,778,797]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="dimensions" pageId="16" pageNumber="17">DIMENSIONS</smallCapsWord>
(n = 13). H
<quantity id="4CA79B63FFD4FF97FDE1FCC1FD7EFCAA" box="[530,699,773,799]" metricMagnitude="-3" metricUnit="m" metricValue="9.004999999999999" metricValueMax="9.83" metricValueMin="8.18" pageId="16" pageNumber="17" unit="mm" value="9.004999999999999" valueMax="9.83" valueMin="8.18">8.18–9.83 mm</quantity>
(mean =
<quantity id="4CA79B63FFD4FF97FCD5FCC1FC57FCAA" box="[806,914,773,799]" metricMagnitude="-3" metricUnit="m" metricValue="9.23" pageId="16" pageNumber="17" unit="mm" value="9.23">9.23 mm</quantity>
); D
<quantity id="4CA79B63FFD4FF97FC4CFCC0FB40FCAA" box="[959,1157,772,799]" metricMagnitude="-2" metricUnit="m" metricValue="1.496" metricValueMax="1.565" metricValueMin="1.427" pageId="16" pageNumber="17" unit="mm" value="14.96" valueMax="15.65" valueMin="14.27">14.27–15.65 mm</quantity>
(mean =
<quantity id="4CA79B63FFD4FF97FB1CFCC1FAAFFCAA" box="[1263,1386,773,799]" metricMagnitude="-2" metricUnit="m" metricValue="1.498" pageId="16" pageNumber="17" unit="mm" value="14.98">14.98 mm</quantity>
); W 54/s–56/8 (mean = 55/8). All the shells with two growth interruptions.
</paragraph>
<paragraph id="8BE03686FFD4FF97FF4EFCADFAC7FBE2" blockId="16.[188,1400,873,1111]" pageId="16" pageNumber="17">
<smallCapsWord id="8D06A05AFFD4FF97FF4EFCADFEF4FC37" baselines="892,893" box="[189,305,873,899]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="16" pageNumber="17">GENITALIA</smallCapsWord>
(
<figureCitation id="13642A03FFD4FF97FEBBFCADFE08FC36" box="[328,461,873,900]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[262,1325,265,1852]" captionTargetId="figure-12@17.[262,1325,265,1852]" captionTargetPageId="17" captionText="Fig. 7. Metafruticicola (Westerlundia) coartata. A. M. (Westerlundia) c. coartata Fuchs &amp; Käufel, 1936, Greece, Cyclades, Makra Islet (NHMC 50.4056). Shell. B–D. M. (Westerlundia) c. gemina Fuchs &amp; Käufel, 1936, Greece, Dodecanese, Karavia Nisia, north Islet (NHMC 50.6805). B. Shell. C. General view of genitalia. D. View of penial papilla. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.8135445" httpUri="https://zenodo.org/record/8135445/files/figure.png" pageId="16" pageNumber="17">Fig. 7C–D</figureCitation>
). The reproductive system and its inner structures are presented for the first time. Flagellum at least twice the length of the epiphallus. Epiphallus evenly thick. Penial retractor is attached near the junction of the epiphallus with the penis. Penis fusiform, around the same length as the epiphallus and with less distinct boundaries with the latter. Penial papilla semi-ellipsoid with smooth surface and a vertical groove in the center; the epiphallic pore lies at its base. Inner penial surface has some prominent vertical ridges (
<figureCitation id="13642A03FFD4FF97FDB3FBDDFD5AFB86" box="[576,671,1049,1075]" captionStart="Fig" captionStartId="17.[189,232,1892,1918]" captionTargetBox="[262,1325,265,1852]" captionTargetId="figure-12@17.[262,1325,265,1852]" captionTargetPageId="17" captionText="Fig. 7. Metafruticicola (Westerlundia) coartata. A. M. (Westerlundia) c. coartata Fuchs &amp; Käufel, 1936, Greece, Cyclades, Makra Islet (NHMC 50.4056). Shell. B–D. M. (Westerlundia) c. gemina Fuchs &amp; Käufel, 1936, Greece, Dodecanese, Karavia Nisia, north Islet (NHMC 50.6805). B. Shell. C. General view of genitalia. D. View of penial papilla. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.8135445" httpUri="https://zenodo.org/record/8135445/files/figure.png" pageId="16" pageNumber="17">Fig. 7D</figureCitation>
). Vas deferens slender. Vagina short and slender with distinct boundaries to the thick base of the duct of the gametolytic gland. Gametolytic gland is ovate.
</paragraph>
</subSubSection>
<subSubSection id="C345650DFFD4FF97FF4EFBBAFE69FA2C" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BE03686FFD4FF97FF4EFBBAFEEAFB2D" blockId="16.[189,303,1150,1176]" box="[189,303,1150,1176]" pageId="16" pageNumber="17">
<heading id="D0A881EAFFD4FF97FF4EFBBAFEEAFB2D" bold="true" box="[189,303,1150,1176]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
<emphasis id="B92BEA94FFD4FF97FF4EFBBAFEEAFB2D" bold="true" box="[189,303,1150,1176]" pageId="16" pageNumber="17">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BE03686FFD4FF97FF4EFB68FE69FA2C" blockId="16.[188,1399,1196,1434]" pageId="16" pageNumber="17">
The shell measurements provided here are slightly different than those given by
<bibRefCitation id="EFCE4B77FFD4FF97FBADFB68FAF1FB73" author="Bank R. A. &amp; Gittenberger E. &amp; Neubert E." box="[1118,1332,1196,1223]" pageId="16" pageNumber="17" pagination="67 - 136" refId="ref11778" refString="Bank R. A., Gittenberger E. &amp; Neubert E. 2013. Radiation of an eastern Mediterranean landsnail genus: revision of the taxa belonging to Metafruticicola von Ihering, 1892 (Gastropoda, Pulmonata: Hygromiidae). Archiv fur Molluskenkunde 142 (1): 67 - 136. https: // doi. org / 10.1127 / arch. moll / 1869 - 0963 / 142 / 067 - 136" type="journal article" year="2013">
Bank
<emphasis id="B92BEA94FFD4FF97FB52FB69FB11FB73" box="[1185,1236,1196,1222]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
. (2013)
</bibRefCitation>
. Both subspecies of
<taxonomicName id="4C5F4D05FFD4FF97FE94FB14FE36FB5C" authorityName="Fuchs &amp; Kaufel" authorityYear="1936" box="[359,499,1232,1257]" class="Gastropoda" family="Hygromiidae" genus="Metafruticicola" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="coartata">
<emphasis id="B92BEA94FFD4FF97FE94FB14FE36FB5C" box="[359,499,1232,1257]" italics="true" pageId="16" pageNumber="17">M. coartata</emphasis>
</taxonomicName>
share similar genitalia, revealing a close relationship between them. Penial papillae of the animals from Karavi north islet are semi-ellipsoid and almost identical to those of
<emphasis id="B92BEA94FFD4FF97FAC8FB37FEE5FA85" italics="true" pageId="16" pageNumber="17">
M. c.
<taxonomicName id="4C5F4D05FFD4FF97FF4EFAD3FEE5FA85" authorityName="Fuchs &amp; Kaufel" authorityYear="1936" box="[189,288,1303,1328]" class="Gastropoda" family="Hygromiidae" genus="Metafruticicola" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="coartata">coartata</taxonomicName>
</emphasis>
from Sochas and Mikri Zafora islets, even though they belong to different subspecies. All three islets belong to the same island group and are part of the Syrna archipelago. In general, penial papillae of the westernmost islets of
<taxonomicName id="4C5F4D05FFD4FF97FDE2FA99FD5EFAC3" authorityName="Fuchs &amp; Kaufel" authorityYear="1936" box="[529,667,1373,1398]" class="Gastropoda" family="Hygromiidae" genus="Metafruticicola" kingdom="Animalia" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="coartata">
<emphasis id="B92BEA94FFD4FF97FDE2FA99FD5EFAC3" box="[529,667,1373,1398]" italics="true" pageId="16" pageNumber="17">M. coartata</emphasis>
</taxonomicName>
’s distribution (Christiani, Askania, Fteno and Makra) show a more bilobed shape.
</paragraph>
</subSubSection>
</treatment>
</document>
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