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<document id="0E66188E44131296649E9AA02FCD7B4D" ID-CLB-Dataset="22035" ID-DOI="10.11646/zootaxa.4779.2.10" ID-GBIF-Dataset="612343eb-2f73-4265-b5dd-25ae1a5f14b1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3833453" ID-ZooBank="52571CFB-BA39-4B6D-83C5-2DC1BDAE4542" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589870053776" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2020" docId="03E61111276BFF861ADC2E89FC00FC73" docLanguage="en" docName="zootaxa.4779.2.10.pdf" docOrigin="Zootaxa 4779 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stenichnodes bouakensis Jałoszyński 2020, sp. n." docType="treatment" docVersion="6" lastPageNumber="294" masterDocId="FFDF6969276FFF831A4B2E10FFD9FFD4" masterDocTitle="Three new species and new records of Stenichnodes Franz (Coleoptera Staphylinidae: Scydmaeninae)" masterLastPageNumber="296" masterPageNumber="289" pageNumber="293" updateTime="1698840248207" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B8D28C246F879E3772A8FBB8CCAB12B2">Three new species and new records of Stenichnodes Franz (Coleoptera Staphylinidae: Scydmaeninae)</mods:title>
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<mods:namePart id="54785DC55A562CEE444553EBCA1986CD">Jałoszyński, Paweł</mods:namePart>
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<treatment id="03E61111276BFF861ADC2E89FC00FC73" ID-DOI="http://doi.org/10.5281/zenodo.3851253" ID-GBIF-Taxon="164184002" ID-Zenodo-Dep="3851253" LSID="urn:lsid:plazi:treatment:03E61111276BFF861ADC2E89FC00FC73" httpUri="http://treatment.plazi.org/id/03E61111276BFF861ADC2E89FC00FC73" lastPageId="5" lastPageNumber="294" pageId="4" pageNumber="293">
<subSubSection id="C355F38C276BFF871ADC2E89FDB4FF60" box="[151,621,153,180]" pageId="4" pageNumber="293" type="nomenclature">
<paragraph id="8BF0A007276BFF871ADC2E89FDB4FF60" blockId="4.[151,621,153,216]" box="[151,621,153,180]" pageId="4" pageNumber="293">
<heading id="D0B8176B276BFF871ADC2E89FDB4FF60" bold="true" box="[151,621,153,180]" fontSize="11" level="1" pageId="4" pageNumber="293" reason="1">
<emphasis id="B93B7C15276BFF871ADC2E89FDB4FF60" bold="true" box="[151,621,153,180]" pageId="4" pageNumber="293">
<taxonomicName id="4C4FDB84276BFF871ADC2E89FDFAFF60" ID-CoL="8Y9T3" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[151,547,153,180]" class="Insecta" family="Staphylinidae" genus="Stenichnodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="293" phylum="Arthropoda" rank="species" species="bouakensis" status="sp. nov.">
<emphasis id="B93B7C15276BFF871ADC2E89FEE1FF60" bold="true" box="[151,312,153,180]" italics="true" pageId="4" pageNumber="293">Stenichnodes</emphasis>
(s. str.)
<emphasis id="B93B7C15276BFF871BD72E89FDFAFF60" bold="true" box="[412,547,153,180]" italics="true" pageId="4" pageNumber="293">bouakensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A208C16E276BFF8718612E8AFDB4FF60" box="[554,621,154,180]" pageId="4" pageNumber="293" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C355F38C276BFF871ADC2EADFE86FF0C" box="[151,351,189,216]" pageId="4" pageNumber="293" type="description">
<paragraph id="8BF0A007276BFF871ADC2EADFE86FF0C" blockId="4.[151,621,153,216]" box="[151,351,189,216]" pageId="4" pageNumber="293">
(
<figureCitation id="1374BC82276BFF871AD42EADFEDDFF0C" box="[159,260,189,216]" captionStart="FIGURES 16" captionStartId="2.[151,264,988,1013]" captionTargetBox="[156,1431,275,956]" captionTargetId="figure@2.[151,1436,270,961]" captionTargetPageId="2" captionText="FIGURES 16. Dorsal habitus (12, 4, 6) and elytral apices in male (3, 5). Stenichnodes cordicollis sp. n. (1); S. spinosus sp. n. (23); S. bouakensis sp. n. (45); S. ceylonensis (Franz), specimen from India (6)." figureDoi="http://doi.org/10.5281/zenodo.3833455" httpUri="https://zenodo.org/record/3833455/files/figure.png" pageId="4" pageNumber="293">Figs 45</figureCitation>
,
<figureCitation id="1374BC82276BFF871B5B2EADFE8FFF0C" box="[272,342,189,216]" captionStart="FIGURES 1522" captionStartId="4.[151,264,1805,1830]" captionTargetBox="[159,1424,570,1770]" captionTargetId="figure@4.[151,1436,561,1779]" captionTargetPageId="4" captionText="FIGURES 1522. Aedeagus in ventral (15, 17, 19, 21) and lateral (16, 18, 20, 22) views. Stenichnodes bouakensis sp. n. (1518); S. ceylonensis (Franz), specimen from India (1922)." figureDoi="http://doi.org/10.5281/zenodo.3833459" httpUri="https://zenodo.org/record/3833459/files/figure.png" pageId="4" pageNumber="293">1518</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C355F38C276BFF871ADC2F16FCE1FEB3" pageId="4" pageNumber="293" type="materials_examined">
<paragraph id="8BF0A007276BFF871ADC2F16FCE1FEB3" blockId="4.[151,1437,261,540]" pageId="4" pageNumber="293">
<emphasis id="B93B7C15276BFF871ADC2F16FE9EFEF4" bold="true" box="[151,327,261,288]" pageId="4" pageNumber="293">Type material.</emphasis>
<materialsCitation id="3B27AA5A276BFF871B1B2F16FCE1FEB3" ID-GBIF-Occurrence="2620583306" collectionCode="MHNG" collectorName="P. Jaloszynski" country="Ivory Coast" location="IVORY COAST" pageId="4" pageNumber="293" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93B7C15276BFF871B1B2F16FD45FEF4" bold="true" box="[336,668,261,288]" pageId="4" pageNumber="293">
<typeStatus id="54F41EA5276BFF871B1B2F16FE18FEF4" box="[336,449,262,288]" pageId="4" pageNumber="293" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F358E097276BFF871B9B2F15FD4EFEF4" box="[464,663,261,288]" name="Ivory Coast" pageId="4" pageNumber="293">IVORY COAST</collectingCountry>
:
</emphasis>
<specimenCount id="9D496B8E276BFF8718EE2F16FD65FEF4" box="[677,700,262,288]" pageId="4" pageNumber="293" type="male"></specimenCount>
, two labels: “IVORY COAST Bouaké / env. Moronou tamisage / 30kmS Toumodi 4.X.80 / Mahnert - Perret” [white, printed], “
<taxonomicName id="4C4FDB84276BFF8719C82F3AFB96FE90" authorityName="Franz" authorityYear="1966" box="[899,1103,298,324]" class="Insecta" family="Staphylinidae" genus="Stenichnodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="293" phylum="Arthropoda" rank="genus">
<emphasis id="B93B7C15276BFF8719C82F3AFB96FE90" box="[899,1103,298,324]" italics="true" pageId="4" pageNumber="293">STENICHNODES</emphasis>
</taxonomicName>
(s. str.) /
<taxonomicName id="4C4FDB84276BFF871E832F39FA9DFE97" authorityName="Jałoszyński" authorityYear="2020" box="[1224,1348,297,323]" class="Insecta" family="Staphylinidae" genus="Stenichnodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="293" phylum="Arthropoda" rank="species" species="bouakensis">
<emphasis id="B93B7C15276BFF871E832F39FA9DFE97" box="[1224,1348,297,323]" italics="true" pageId="4" pageNumber="293">bouakensis</emphasis>
</taxonomicName>
m. /
<collectorName id="26BAC5D1276BFF871FC22F3AFEC2FEB3" pageId="4" pageNumber="293">P. Jałoszyński</collectorName>
, 2020 /
<typeStatus id="54F41EA5276BFF871B3F2F5DFDC7FEB3" box="[372,542,333,359]" pageId="4" pageNumber="293" type="holotype">HOLOTYPUS</typeStatus>
” [red, printed] (
<collectionCode id="ED5E38C2276BFF87189B2F5DFCF5FEB3" box="[720,812,333,359]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="4" pageNumber="293" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C355F38C276BFF871A8C2F61FF15FE00" pageId="4" pageNumber="293" type="diagnosis">
<paragraph id="8BF0A007276BFF871A8C2F61FF15FE00" blockId="4.[151,1437,261,540]" pageId="4" pageNumber="293">
<emphasis id="B93B7C15276BFF871A8C2F61FE9BFE58" bold="true" box="[199,322,369,396]" pageId="4" pageNumber="293">Diagnosis.</emphasis>
Elytral apex in male with small and shallow common subapical impression; protarsomere I in male unmodified; antennae relatively thick, antennomeres IX and X each slightly transverse; aedeagus lacking param- eres.
</paragraph>
</subSubSection>
<subSubSection id="C355F38C276BFF861A8C2FCDFE47FD27" lastPageId="5" lastPageNumber="294" pageId="4" pageNumber="293" type="description">
<paragraph id="8BF0A007276BFF871A8C2FCDFE42FDC8" blockId="4.[151,1437,261,540]" pageId="4" pageNumber="293">
<emphasis id="B93B7C15276BFF871A8C2FCDFE80FE2C" bold="true" box="[199,345,477,504]" pageId="4" pageNumber="293">Description.</emphasis>
Body of male (
<figureCitation id="1374BC82276BFF87184E2FCDFD91FE23" box="[517,584,477,503]" captionStart="FIGURES 16" captionStartId="2.[151,264,988,1013]" captionTargetBox="[156,1431,275,956]" captionTargetId="figure@2.[151,1436,270,961]" captionTargetPageId="2" captionText="FIGURES 16. Dorsal habitus (12, 4, 6) and elytral apices in male (3, 5). Stenichnodes cordicollis sp. n. (1); S. spinosus sp. n. (23); S. bouakensis sp. n. (45); S. ceylonensis (Franz), specimen from India (6)." figureDoi="http://doi.org/10.5281/zenodo.3833455" httpUri="https://zenodo.org/record/3833455/files/figure.png" pageId="4" pageNumber="293">Fig. 4</figureCitation>
) elongate and slender, strongly convex, light brown, covered with yellowish vestiture; BL
<quantity id="4CB70DE2276BFF871B7B2C11FE4EFDC8" box="[304,407,513,540]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="4" pageNumber="293" unit="mm" value="0.85">0.85 mm</quantity>
.
</paragraph>
<caption id="DF30F08F276BFF871ADC291DFCCAF89E" ID-DOI="http://doi.org/10.5281/zenodo.3833459" ID-Zenodo-Dep="3833459" httpUri="https://zenodo.org/record/3833459/files/figure.png" pageId="4" pageNumber="293" startId="4.[151,264,1805,1830]" targetBox="[159,1424,570,1770]" targetPageId="4">
<paragraph id="8BF0A007276BFF871ADC291DFCCAF89E" blockId="4.[151,1436,1805,1866]" pageId="4" pageNumber="293">
<emphasis id="B93B7C15276BFF871ADC291DFE8EF8F2" bold="true" box="[151,343,1805,1830]" pageId="4" pageNumber="293">FIGURES 1522.</emphasis>
Aedeagus in ventral (15, 17, 19, 21) and lateral (16, 18, 20, 22) views.
<taxonomicName id="4C4FDB84276BFF871E1C291DFA8EF8F1" authority="Jałoszyński, 2020" authorityName="Jałoszyński" authorityYear="2020" box="[1111,1367,1805,1829]" class="Insecta" family="Staphylinidae" genus="Stenichnodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="293" phylum="Arthropoda" rank="species" species="bouakensis" status="sp. nov.">
<emphasis id="B93B7C15276BFF871E1C291DFA8EF8F1" box="[1111,1367,1805,1829]" italics="true" pageId="4" pageNumber="293">Stenichnodes bouakensis</emphasis>
</taxonomicName>
<emphasis id="B93B7C15276BFF871F2B291EFA45F8F2" bold="true" box="[1376,1436,1806,1830]" pageId="4" pageNumber="293">
<taxonomicNameLabel id="A208C16E276BFF871F2B291EFA45F8F2" box="[1376,1436,1806,1830]" pageId="4" pageNumber="293" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(1518);
<taxonomicName id="4C4FDB84276BFF871ABD2922FE01F89E" authority="(Franz)" baseAuthorityName="Franz" baseAuthorityYear="1982" box="[246,472,1841,1866]" class="Insecta" family="Staphylinidae" genus="Stenichnodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="293" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="B93B7C15276BFF871ABD2922FE5CF89D" box="[246,389,1841,1865]" italics="true" pageId="4" pageNumber="293">S. ceylonensis</emphasis>
(Franz)
</taxonomicName>
, specimen from India (1922).
</paragraph>
</caption>
<paragraph id="8BF0A007276BFF861A8C2968FB43FECB" blockId="4.[151,1437,1911,2010]" lastBlockId="5.[151,1437,153,935]" lastPageId="5" lastPageNumber="294" pageId="4" pageNumber="293">
Head broadest at large, strongly convex and finely faceted eyes, HL
<quantity id="4CB70DE2276BFF8719F52967FBFAF846" box="[958,1059,1911,1938]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="293" unit="mm" value="0.15">0.15 mm</quantity>
, HW
<quantity id="4CB70DE2276BFF871E292968FB10F846" box="[1122,1225,1912,1938]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="4" pageNumber="293" unit="mm" value="0.18">0.18 mm</quantity>
; tempora vestigial; vertex weakly convex and anteriorly confluent with frons; frons slightly flattened, supraantennal tubercles indistinct. Each eye bean-shaped, transverse in relation to the long body axis, with narrow posteromedian emargination. Vertex and frons with fine, inconspicuous punctures; setae fine, nearly recumbent, barely discernible under magnification 100 ×. Antennae relatively robust, with slightly flattened, trimerous clubs, AnL
<quantity id="4CB70DE2276AFF861EC62EADFB2CFF03" box="[1165,1269,189,215]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="5" pageNumber="294" unit="mm" value="0.28">0.28 mm</quantity>
; antennomeres III strongly elongate, III slightly transverse, IVVII each about as long as broad, VIIIX each slightly transverse, XI about 1.2 × as long as broad, much shorter than IX and X combined, with rounded apex.
</paragraph>
<paragraph id="8BF0A007276AFF861A8C2F39FD14FE7B" blockId="5.[151,1437,153,935]" pageId="5" pageNumber="294">
Pronotum heart-shaped, broadest near anterior third; PL
<quantity id="4CB70DE2276AFF8619732F39FC44FE97" box="[824,925,297,323]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="294" unit="mm" value="0.23">0.23 mm</quantity>
, PW
<quantity id="4CB70DE2276AFF86199C2F39FBE7FE97" box="[983,1086,297,323]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="294" unit="mm" value="0.2">0.20 mm</quantity>
. Anterior and posterior margins weakly rounded, sides in anterior half strongly rounded; posteriorly nearly straight and not constricted; base with shallow but distinct transverse groove indistinctly deepened at each end. Punctures on pronotal disc inconspicuous, fine and sparse; setae sparse, short and recumbent.
</paragraph>
<paragraph id="8BF0A007276AFF861A8C2FA9FBAAFDEB" blockId="5.[151,1437,153,935]" pageId="5" pageNumber="294">
Elytra together oval, broadest distinctly in front of middle; EL
<quantity id="4CB70DE2276AFF8619C62FA9FC2DFE07" box="[909,1012,441,467]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="294" unit="mm" value="0.48">0.48 mm</quantity>
, EW
<quantity id="4CB70DE2276AFF861E782FA8FB43FE07" box="[1075,1178,440,467]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="294" unit="mm" value="0.35">0.35 mm</quantity>
, EI 1.36; humeral calli distinct, elongate; basal impressions shallow but distinct; apices separately rounded, each with small semicircular subapical impression, together forming common round elytral pit (
<figureCitation id="1374BC82276AFF8619302C11FC66FDCF" box="[891,959,512,539]" captionStart="FIGURES 16" captionStartId="2.[151,264,988,1013]" captionTargetBox="[156,1431,275,956]" captionTargetId="figure@2.[151,1436,270,961]" captionTargetPageId="2" captionText="FIGURES 16. Dorsal habitus (12, 4, 6) and elytral apices in male (3, 5). Stenichnodes cordicollis sp. n. (1); S. spinosus sp. n. (23); S. bouakensis sp. n. (45); S. ceylonensis (Franz), specimen from India (6)." figureDoi="http://doi.org/10.5281/zenodo.3833455" httpUri="https://zenodo.org/record/3833455/files/figure.png" pageId="5" pageNumber="294">Fig. 5</figureCitation>
). Punctures on elytra slightly more distinct than those on pronotum, but superficial and inconspicuous; setae sparse, short, suberect.
</paragraph>
<paragraph id="8BF0A007276AFF861A8C2C59FE31FDB7" blockId="5.[151,1437,153,935]" box="[199,488,585,611]" pageId="5" pageNumber="294">Hind wings present, long.</paragraph>
<paragraph id="8BF0A007276AFF861A8C2C7DFD8AFD53" blockId="5.[151,1437,153,935]" box="[199,595,621,647]" pageId="5" pageNumber="294">Legs long and slender, unmodified.</paragraph>
<paragraph id="8BF0A007276AFF861A8C2C81FB9BFD1B" blockId="5.[151,1437,153,935]" pageId="5" pageNumber="294">
Aedeagus (
<figureCitation id="1374BC82276AFF861B0F2C81FE1BFD7F" box="[324,450,656,683]" captionStart="FIGURES 1522" captionStartId="4.[151,264,1805,1830]" captionTargetBox="[159,1424,570,1770]" captionTargetId="figure@4.[151,1436,561,1779]" captionTargetPageId="4" captionText="FIGURES 1522. Aedeagus in ventral (15, 17, 19, 21) and lateral (16, 18, 20, 22) views. Stenichnodes bouakensis sp. n. (1518); S. ceylonensis (Franz), specimen from India (1922)." figureDoi="http://doi.org/10.5281/zenodo.3833459" httpUri="https://zenodo.org/record/3833459/files/figure.png" pageId="5" pageNumber="294">Figs 1518</figureCitation>
) elongate and with relatively thin walls, in ventral view median lobe slightly asymmetrical, with a group of broad and short apical projections; parameres absent; AeL 0.20.
</paragraph>
<paragraph id="8BF0A007276AFF861A8C2CC9FE47FD27" blockId="5.[151,1437,153,935]" box="[199,414,729,755]" pageId="5" pageNumber="294">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C355F38C276AFF861A8C2CECFD95FCC3" box="[199,588,764,791]" pageId="5" pageNumber="294" type="distribution">
<paragraph id="8BF0A007276AFF861A8C2CECFD95FCC3" blockId="5.[151,1437,153,935]" box="[199,588,764,791]" pageId="5" pageNumber="294">
<emphasis id="B93B7C15276AFF861A8C2CECFEB8FCC3" bold="true" box="[199,353,764,791]" pageId="5" pageNumber="294">Distribution.</emphasis>
Central
<collectingCountry id="F358E097276AFF861B8A2CEDFD91FCC3" box="[449,584,765,791]" name="Ivory Coast" pageId="5" pageNumber="294">Ivory Coast</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C355F38C276AFF861A8C2D31FCFFFCEF" box="[199,806,801,827]" pageId="5" pageNumber="294" type="etymology">
<paragraph id="8BF0A007276AFF861A8C2D31FCFFFCEF" blockId="5.[151,1437,153,935]" box="[199,806,801,827]" pageId="5" pageNumber="294">
<emphasis id="B93B7C15276AFF861A8C2D31FE97FCEF" bold="true" box="[199,334,801,827]" pageId="5" pageNumber="294">Etymology.</emphasis>
After Bouaké Department of
<collectingCountry id="F358E097276AFF8618D02D31FCFBFCEF" box="[667,802,801,827]" name="Ivory Coast" pageId="5" pageNumber="294">Ivory Coast</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C355F38C276AFF861A8C2D55FC00FC73" pageId="5" pageNumber="294" type="discussion">
<paragraph id="8BF0A007276AFF861A8C2D55FC00FC73" blockId="5.[151,1437,153,935]" pageId="5" pageNumber="294">
<emphasis id="B93B7C15276AFF861A8C2D55FEE2FC8B" bold="true" box="[199,315,837,863]" pageId="5" pageNumber="294">Remarks.</emphasis>
<taxonomicName id="4C4FDB84276AFF861B0D2D55FDB9FC8B" authorityName="Jałoszyński" authorityYear="2020" box="[326,608,837,863]" class="Insecta" family="Staphylinidae" genus="Stenichnodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="bouakensis">
<emphasis id="B93B7C15276AFF861B0D2D55FDB9FC8B" box="[326,608,837,863]" italics="true" pageId="5" pageNumber="294">Stenichnodes bouakensis</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C4FDB84276AFF8618AD2D55FCBCFC8B" authorityName="Jałoszyński" authorityYear="2020" box="[742,869,837,863]" class="Insecta" family="Staphylinidae" genus="Stenichnodes" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="294" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B93B7C15276AFF8618AD2D55FCBCFC8B" box="[742,869,837,863]" italics="true" pageId="5" pageNumber="294">S. spinosus</emphasis>
</taxonomicName>
in body form, measurements and proportions of body parts, and even in the shape and structure of the aedeagus. The differences in elytral modifications of males are profound, and can be used to unambiguously distinguish these species.
</paragraph>
</subSubSection>
</treatment>
</document>