180 lines
20 KiB
XML
180 lines
20 KiB
XML
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<mods:title id="4FE2ED67CE7092C0EE74C2E667689362">Additional Amphibians And Reptiles From The Phnom Samkos Wildlife Sanctuary In Northwestern Cardamom Mountains, Cambodia, With Comments On Their Taxonomy And The Discovery Of Three New Species</mods:title>
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<mods:namePart id="E61AC5142C26C681AFA7328543C2BA2D">Grismer, L. Lee</mods:namePart>
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<mods:namePart id="7E68083C3F8D7A7AD314B9E82B683F45">Neang, Thy</mods:namePart>
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<mods:namePart id="211FF4A1AEAC9327A2D6C599F256A383">Chav, Thou</mods:namePart>
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<mods:namePart id="549A4375AEA97DE9ADCF7D590D69EC48">Wood, Perry L.</mods:namePart>
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<mods:namePart id="35D4FD97B70C74CCEB07A6110D42257B">Oaks, Jamie R.</mods:namePart>
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<mods:title id="3C8462C1235D054D7FAC8F0BB5D33397">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="950CF9CC40092304C02263F2F68AEBDB">2008</mods:date>
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<mods:number id="8CABBD93F33C8EDBFB487189496B6783">56</mods:number>
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<mods:identifier id="BAE4A0C053791D24CFF11EB5AF771862" type="ISSN">2345-7600</mods:identifier>
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<treatment id="039E5D54FF94C52F6587FBC5A01FF84E" ID-DOI="http://doi.org/10.5281/zenodo.5415543" ID-Zenodo-Dep="5415543" LSID="urn:lsid:plazi:treatment:039E5D54FF94C52F6587FBC5A01FF84E" httpUri="http://treatment.plazi.org/id/039E5D54FF94C52F6587FBC5A01FF84E" lastPageNumber="168" pageId="7" pageNumber="168">
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<subSubSection id="C32DBFC9FF94C52F6587FBC5A703F973" box="[243,622,1053,1077]" pageId="7" pageNumber="168" type="nomenclature">
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<paragraph id="8B88EC42FF94C52F6587FBC5A703F973" blockId="7.[243,622,1053,1077]" box="[243,622,1053,1077]" pageId="7" pageNumber="168">
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<heading id="D0C05B2EFF94C52F6587FBC5A703F973" bold="true" box="[243,622,1053,1077]" centered="true" fontSize="10" level="2" pageId="7" pageNumber="168" reason="2">
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<emphasis id="B9433050FF94C52F6587FBC5A703F973" bold="true" box="[243,622,1053,1077]" pageId="7" pageNumber="168">
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<taxonomicName id="4C3797C1FF94C52F6587FBC5A76BF973" ID-CoL="4LD52" baseAuthorityName="Gravenhorst" baseAuthorityYear="1829" box="[243,518,1053,1077]" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="7" pageNumber="168" phylum="Chordata" rank="species" species="leucomystax">
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<emphasis id="B9433050FF94C52F6587FBC5A76BF973" bold="true" box="[243,518,1053,1077]" italics="true" pageId="7" pageNumber="168">Polypedates leucomystax</emphasis>
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</taxonomicName>
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complex
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C32DBFC9FF94C52F6505FB85A71AF9EA" pageId="7" pageNumber="168" type="materials_examined">
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<paragraph id="8B88EC42FF94C52F6505FB85A71AF9EA" blockId="7.[113,752,1117,1196]" pageId="7" pageNumber="168">
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<emphasis id="B9433050FF94C52F6505FB85A426F934" bold="true" box="[113,331,1117,1138]" italics="true" pageId="7" pageNumber="168">Material examined. –</emphasis>
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<materialsCitation id="3B5FE61FFF94C52F6420FB85A780F934" box="[340,749,1117,1138]" collectingDate="2006-08-04" collectionCode="LSUHC" pageId="7" pageNumber="168" specimenCode="LSUHC 7759-65" specimenCount="1">
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<specimenCode id="DB914439FF94C52F6420FB85A769F934" box="[340,516,1117,1138]" collectionCode="LSUHC" country="0" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="7" pageNumber="168">LSUHC 7759–65</specimenCode>
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: Pramaoy,
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<date id="FF89CA82FF94C52F6701FB85A780F934" box="[629,749,1117,1138]" pageId="7" pageNumber="168" value="2006-08-04">
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<collectingDate id="EFCD336AFF94C52F6701FB85A780F934" box="[629,749,1117,1138]" pageId="7" pageNumber="168" value="2006-08-04">4 Aug.2006</collectingDate>
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</date>
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</materialsCitation>
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;
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<materialsCitation id="3B5FE61FFF94C52F6505FBA2A74DF9C9" box="[113,544,1146,1167]" collectingDate="2006-08-05" collectionCode="LSUHC" pageId="7" pageNumber="168" specimenCode="LSUHC 7817" specimenCount="1">
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<specimenCode id="DB914439FF94C52F6505FBA2A594F9C9" box="[113,249,1146,1167]" collectionCode="LSUHC" country="0" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="7" pageNumber="168">LSUHC 7817</specimenCode>
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: Che Teal Chrum,
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<date id="FF89CA82FF94C52F64DCFBA2A74DF9C9" box="[424,544,1146,1167]" pageId="7" pageNumber="168" value="2006-08-05">
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<collectingDate id="EFCD336AFF94C52F64DCFBA2A74DF9C9" box="[424,544,1146,1167]" pageId="7" pageNumber="168" value="2006-08-05">5 Aug. 2006</collectingDate>
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</date>
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</materialsCitation>
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;
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<materialsCitation id="3B5FE61FFF94C52F675CFBA2A46CF9EA" collectingDate="2006-08-06" collectionCode="LSUHC" pageId="7" pageNumber="168" specimenCode="LSUHC 7847" specimenCount="1">
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<specimenCode id="DB914439FF94C52F675CFBA2A7C2F9C9" box="[552,687,1146,1167]" collectionCode="LSUHC" country="0" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="7" pageNumber="168">LSUHC 7847</specimenCode>
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: Camp 1,
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<date id="FF89CA82FF94C52F65FFFB4FA46CF9EA" box="[139,257,1175,1196]" pageId="7" pageNumber="168" value="2006-08-06">
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<collectingDate id="EFCD336AFF94C52F65FFFB4FA46CF9EA" box="[139,257,1175,1196]" pageId="7" pageNumber="168" value="2006-08-06">6 Aug.2006</collectingDate>
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</date>
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</materialsCitation>
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;
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<materialsCitation id="3B5FE61FFF94C52F6479FB4FA71EF9EA" box="[269,627,1175,1196]" collectingDate="2006-08-09" collectionCode="LSUHC" pageId="7" pageNumber="168" specimenCode="LSUHC 7899" specimenCount="1">
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<specimenCode id="DB914439FF94C52F6479FB4FA4FAF9EA" box="[269,407,1175,1196]" collectionCode="LSUHC" country="0" httpUri="http://grbio.org/cool/gc15-4qsc" name="La Sierra University, Herpetological Collection" pageId="7" pageNumber="168">LSUHC 7899</specimenCode>
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: Camp 2,
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<date id="FF89CA82FF94C52F6489FB4FA71EF9EA" box="[509,627,1175,1196]" pageId="7" pageNumber="168" value="2006-08-09">
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<collectingDate id="EFCD336AFF94C52F6489FB4FA71EF9EA" box="[509,627,1175,1196]" pageId="7" pageNumber="168" value="2006-08-09">9 Aug.2006</collectingDate>
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</date>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32DBFC9FF94C52F6505FB0CA01FF84E" pageId="7" pageNumber="168" type="discussion">
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<paragraph id="8B88EC42FF94C52F6505FB0CA00AFF23" blockId="7.[113,753,1236,1547]" lastBlockId="7.[813,1452,192,613]" pageId="7" pageNumber="168">
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<emphasis id="B9433050FF94C52F6505FB0CA59EF9AA" bold="true" box="[113,243,1236,1260]" italics="true" pageId="7" pageNumber="168">Remarks. –</emphasis>
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The
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<taxonomicName id="4C3797C1FF94C52F6446FB0CA72AF9AD" baseAuthorityName="Gravenhorst" baseAuthorityYear="1829" box="[306,583,1236,1259]" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="7" pageNumber="168" phylum="Chordata" rank="species" species="leucomystax">
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<emphasis id="B9433050FF94C52F6446FB0CA72AF9AD" box="[306,583,1236,1259]" italics="true" pageId="7" pageNumber="168">Polypedates leucomystax</emphasis>
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</taxonomicName>
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complex is an incompletely understood species group with many distinct forms from various regions masquerading under a single name (
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<bibRefCitation id="EFA691B3FF94C52F650CFAECA455F80D" author="Inger, R" box="[120,312,1332,1355]" pageId="7" pageNumber="168" pagination="1 - 46" refId="ref10520" refString="Inger, R. F., N. Orlov, & I. Darevsky, 1999. Frogs of Vietnam: a report on new collections. Fieldiana: Zoology (New series), 92: 1 - 46." type="journal article" year="1999">Inger et al., 1999</bibRefCitation>
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; Narins, et al., 1998;
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<bibRefCitation id="EFA691B3FF94C52F6753FAECA781F80D" author="Orlov, N & Lathrop, R" box="[551,748,1332,1355]" pageId="7" pageNumber="168" pagination="17 - 44" refId="ref11063" refString="Orlov, N. L., A. Lathrop, R. W. Murphy, & H. T. Cuc, 2001. Frogs of the family Rhacophoridae (Anura: Amphibia) in northern Hoang Lien Mountains (Mount Fan Si Pan, Sa Pa District, Lao Cai Province), Vietnam. Russian Journal of Herpetology, 8: 17 - 44." type="journal article" year="2001">Orlov et al., 2001</bibRefCitation>
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||
;
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<bibRefCitation id="EFA691B3FF94C52F6505FA8CA43CF82D" author="Trepanier, T" box="[113,337,1364,1387]" pageId="7" pageNumber="168" pagination="102 - 106" refId="ref11826" refString="Trepanier, T. L., A. Lathrop, & R. W. Murphy, 1999. Rhacophorus leucomystax in Vietnam with acoustic analysis of courtship territorial calls. Asiatic Herpetological Research, 8: 102 - 106." type="journal article" year="1999">Trepanier et al., 1999</bibRefCitation>
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; Zhao & Adler 1993) if not more. Seven adult males (LSUHC 7759–65; SVL
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<quantity id="4CCF41A7FF94C52F675CFAACA7CAF8CD" box="[552,679,1396,1419]" metricMagnitude="-2" metricUnit="m" metricValue="5.45" metricValueMax="5.7" metricValueMin="5.2" pageId="7" pageNumber="168" unit="mm" value="54.5" valueMax="57.0" valueMin="52.0">52–57 mm</quantity>
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) from Pramaoy and an adult female (LSUHC 7817; SVL
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<quantity id="4CCF41A7FF94C52F67EFFA4CA784F8ED" box="[667,745,1428,1451]" metricMagnitude="-2" metricUnit="m" metricValue="7.9" pageId="7" pageNumber="168" unit="mm" value="79.0">79 mm</quantity>
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) from Che Teal Chrum match the description of
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<taxonomicName id="4C3797C1FF94C52F671AFA6CA41AF8AD" authority="leucomystax" class="Amphibia" family="Rhacophoridae" genus="Polypedates" kingdom="Animalia" order="Anura" pageId="7" pageNumber="168" phylum="Chordata" rank="subSpecies" species="leucomystax" subSpecies="leucomystax">
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<emphasis id="B9433050FF94C52F671AFA6CA41AF8AD" italics="true" pageId="7" pageNumber="168">Polypedates leucomystax leucomystax</emphasis>
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</taxonomicName>
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in
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<bibRefCitation id="EFA691B3FF94C52F64E1FA0CA749F8AD" author="Taylor, E" box="[405,548,1492,1515]" pageId="7" pageNumber="168" pagination="265 - 599" refId="ref11694" refString="Taylor, E. H., 1962. The amphibian fauna of Thailand. University of Science Kansas Bulletin, 43: 265 - 599." type="journal article" year="1962">Taylor (1962)</bibRefCitation>
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and
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<specimenCount id="9D3127CBFF94C52F6720FA0CA79DF8AD" box="[596,752,1492,1515]" pageId="7" pageNumber="168" type="generic">four specimens</specimenCount>
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from the northeastern Cardamoms (
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<bibRefCitation id="EFA691B3FF94C52F648FFA2CA781FB4D" author="Grismer, L & Chav T & Neang, T & Wood, Jr., J & Grismer, T & Youmans, A & Ponce, J" box="[507,748,1524,1547]" pageId="7" pageNumber="168" pagination="216 - 241" refId="ref10230" refString="Grismer, L. L., Chav T., Neang, T., P. L. Wood, Jr., J. L. Grismer, T. M. Youmans, A. Ponce, J. C. Daltry, & H. Kaiser. 2007 a. The herpetofauna of the Phnom Aural Wildelife Sanctuary and checklist of the herpetofauna of the Cardamom Mountains, Cambodia. Hamadryad 31: 216 - 241." type="journal article" year="2007">Grismer et al., 2007a</bibRefCitation>
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||
; LSUHC 7347–48, 7354–55) in that the skin of the head is fused to the frontoparietal and nasal bones; vocal sac present in males; interorbital space wider than upper eyelid; tympanum distinct; fingers with small (nearly absent in our sample) webbing; toes at least 2/3 webbed; digital discs smaller than tympanum; tibiotarsal articulation reaching from eye to end of snout; vomerine teeth present; and an hourglassshaped, dark marking on head and occiput reaching shoulders (only in LSUHC 7760–62, 7764–65; pattern obscured in LSUHC 7759, 7763). LSUHC 7817 from Che Teal Chrum has four, dark, dorsal stripes instead of an hourglass marking, thus matching the description of
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<emphasis id="B9433050FF94C52F61EDFDC9A026FF6E" box="[1177,1355,529,552]" italics="true" pageId="7" pageNumber="168">P. l. sexvirgatus</emphasis>
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||
(
|
||
<bibRefCitation id="EFA691B3FF94C52F6028FDC9A60AFF01" author="Taylor, E" pageId="7" pageNumber="168" pagination="265 - 599" refId="ref11694" refString="Taylor, E. H., 1962. The amphibian fauna of Thailand. University of Science Kansas Bulletin, 43: 265 - 599." type="journal article" year="1962">Taylor, 1962</bibRefCitation>
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||
). Similar variation was reported in populations of this complex from Phnom Aural (
|
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<bibRefCitation id="EFA691B3FF94C52F611AFD96A036FF23" author="Grismer, L & Chav T & Neang, T & Wood, Jr., J & Grismer, T & Youmans, A & Ponce, J" box="[1134,1371,590,613]" pageId="7" pageNumber="168" pagination="216 - 241" refId="ref10230" refString="Grismer, L. L., Chav T., Neang, T., P. L. Wood, Jr., J. L. Grismer, T. M. Youmans, A. Ponce, J. C. Daltry, & H. Kaiser. 2007 a. The herpetofauna of the Phnom Aural Wildelife Sanctuary and checklist of the herpetofauna of the Cardamom Mountains, Cambodia. Hamadryad 31: 216 - 241." type="journal article" year="2007">Grismer et al., 2007a</bibRefCitation>
|
||
).
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</paragraph>
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<paragraph id="8B88EC42FF94C52F6659FD54A06EFFB9" blockId="7.[813,1452,652,767]" pageId="7" pageNumber="168">Individuals of these two populations were collected at night while sitting on the leaves of low bushes next to roadside puddles in anthropogenically modified habitats. Many others were heard calling from nearby vegetation.</paragraph>
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<paragraph id="8B88EC42FF94C52F6659FCFDA19AFEF1" blockId="7.[813,1451,805,951]" pageId="7" pageNumber="168">
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LSUHC 7847 from Camp 1 (adult male, SVL
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||
<quantity id="4CCF41A7FF94C52F6025FCFDA0C9FE7A" box="[1361,1444,805,828]" metricMagnitude="-2" metricUnit="m" metricValue="4.9" pageId="7" pageNumber="168" unit="mm" value="49.0">49 mm</quantity>
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) matches
|
||
<bibRefCitation id="EFA691B3FF94C52F66FFFC9CA159FE1D" author="Taylor, E" box="[907,1076,836,859]" pageId="7" pageNumber="168" pagination="265 - 599" refId="ref11694" refString="Taylor, E. H., 1962. The amphibian fauna of Thailand. University of Science Kansas Bulletin, 43: 265 - 599." type="journal article" year="1962">Taylor’s (1962)</bibRefCitation>
|
||
description except that the dorsum has dark mottling and the flanks have dark stippling. This specimen was collected deep within undisturbed forest while sitting in vegetation along a small stream.
|
||
</paragraph>
|
||
<paragraph id="8B88EC42FF94C52F6659FC05A0F0F929" blockId="7.[813,1452,989,1135]" pageId="7" pageNumber="168">
|
||
LSUHC 7899 from Camp 2 is a large adult female (SVL
|
||
<quantity id="4CCF41A7FF94C52F60E5FC05A62EF955" metricMagnitude="1" metricUnit="m" metricValue="9.9" pageId="7" pageNumber="168" unit="m" value="99.0">99 m</quantity>
|
||
) generally matching
|
||
<bibRefCitation id="EFA691B3FF94C52F616FFC24A1D3F955" author="Taylor, E" box="[1051,1214,1020,1043]" pageId="7" pageNumber="168" pagination="265 - 599" refId="ref11694" refString="Taylor, E. H., 1962. The amphibian fauna of Thailand. University of Science Kansas Bulletin, 43: 265 - 599." type="journal article" year="1962">Taylor’s (1962)</bibRefCitation>
|
||
description except that the dorsum is cream coloured with a few isolated dark spots. This specimen was collected while sitting 1.5 m above a fastflowing, rocky stream coursing through secondary forest.
|
||
</paragraph>
|
||
<paragraph id="8B88EC42FF94C52F6659FB4DA01FF84E" blockId="7.[813,1452,1173,1288]" pageId="7" pageNumber="168">
|
||
This species complex has been reported from the central Cardamoms by
|
||
<bibRefCitation id="EFA691B3FF94C52F66A0FB6CA1F8F98D" author="Ohler, A" box="[980,1173,1204,1227]" pageId="7" pageNumber="168" pagination="465 - 481" refId="ref11012" refString="Ohler, A., S. R. Swann & J. C. Daltry, 2002. A recent survey of the amphibian fauna of the Cardamom Mountains, southwest Cambodia with descriptions of three new species. Raffles Bulletin of Zoology, 50: 465 - 481." type="journal article" year="2002">Ohler et al. (2002)</bibRefCitation>
|
||
and from the northeastern Cardamoms by
|
||
<bibRefCitation id="EFA691B3FF94C52F66ADFB0AA1A0F9AF" author="Grismer, L & Chav T & Neang, T & Wood, Jr., J & Grismer, T & Youmans, A & Ponce, J" box="[985,1229,1234,1257]" pageId="7" pageNumber="168" pagination="216 - 241" refId="ref10230" refString="Grismer, L. L., Chav T., Neang, T., P. L. Wood, Jr., J. L. Grismer, T. M. Youmans, A. Ponce, J. C. Daltry, & H. Kaiser. 2007 a. The herpetofauna of the Phnom Aural Wildelife Sanctuary and checklist of the herpetofauna of the Cardamom Mountains, Cambodia. Hamadryad 31: 216 - 241." type="journal article" year="2007">Grismer et al. (2007a)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFA691B3FF94C52F61AEFB0AA0C5F9AF" author="Ohler, A" box="[1242,1448,1234,1257]" pageId="7" pageNumber="168" pagination="465 - 481" refId="ref11012" refString="Ohler, A., S. R. Swann & J. C. Daltry, 2002. A recent survey of the amphibian fauna of the Cardamom Mountains, southwest Cambodia with descriptions of three new species. Raffles Bulletin of Zoology, 50: 465 - 481." type="journal article" year="2002">Ohler et al. (2002)</bibRefCitation>
|
||
, Stuart & Emmett (2006), and Swan & Daltry (2002).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |