treatments-xml/data/03/9B/69/039B692C9A347E620DA4FBAEFA8016A0.xml

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<mods:title id="335FC8A2BA334E8282FBC3133AB6B701">Revision of the loach genus Claea Kottelat, 2010 (Teleostei: Nemacheilidae) in China, with a description of a new species from the Chang-Jiang basin</mods:title>
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<mods:namePart id="74AF09D8C8D508EEB75D21A093CB7914">Zhang, Chu-Yi</mods:namePart>
<mods:affiliation id="C72F2228E9D81574A30757D6E321A295">Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 430072, Hubei Province, P. R. China &amp; Shanghai Universities Key Laboratory of Marine Animal Taxonomy and Evolution, Shanghai Ocean University, Shanghai 201306, P. R. China</mods:affiliation>
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<mods:namePart id="CD77CAC1A78268605F08B7507CC6BAED">Luo, Pan</mods:namePart>
<mods:affiliation id="1FED717770B05649FC7946AD2A4E1BE7">Hubei Provincial Academy of Eco-environmental Sciences, Wuhan 430072, Hubei Province, P. R. China</mods:affiliation>
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<mods:namePart id="9D7F83AB7B680D5B8C67EB63291062F8">Huang, Feng</mods:namePart>
<mods:affiliation id="CD077CECF4D5B3129C6BBB2B725BDC91">Hubei Provincial Academy of Eco-environmental Sciences, Wuhan 430072, Hubei Province, P. R. China</mods:affiliation>
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<taxonomicName id="4C32A3B99A347E610DA4FBAEFCEA15FC" authority="(Chen, Sheraliev, Shu &amp; Peng, 2021)" baseAuthorityName="Chen, Sheraliev, Shu &amp; Peng" baseAuthorityYear="2021" box="[151,809,1127,1154]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="species" species="wulongensis">
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<emphasis id="B94604289A347E610DA4FBAEFEB315FF" bold="true" box="[151,368,1127,1154]" italics="true" pageId="13" pageNumber="417">Claea wulongensis</emphasis>
(
<bibRefCitation id="EFA3A5CB9A347E610C4CFBAEFCE115FC" author="Chen, S. &amp; Sheraliev, B. &amp; Shu, L. &amp; Peng, Z." box="[383,802,1127,1154]" pageId="13" pageNumber="417" pagination="179 - 192" refId="ref9787" refString="Chen, S., Sheraliev, B., Shu, L. &amp; Peng, Z. (2021) Triplophysa wulongensis, a new species of cave-dwelling loach (Teleostei, Nemacheilidae) from Chongqing, Southwest China. ZooKeys, 1026, 179 - 192. https: // doi. org / 10.3897 / zookeys. 1026.61570" type="journal article" year="2021">Chen, Sheraliev, Shu &amp; Peng, 2021</bibRefCitation>
)
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(
<figureCitation id="1309C4BF9A347E610DACFB42FEC315DB" box="[159,256,1163,1190]" captionStart="FIGURE 7" captionStartId="8.[151,250,1405,1429]" captionTargetBox="[151,1435,179,1380]" captionTargetId="figure-17@8.[151,1435,179,1380]" captionTargetPageId="8" captionText="FIGURE 7. Lateral view of body in Claea wulongensis: SWU 2019051309, holotype, 64.0 mm SL, Southwest China: Chongqing: Wulong County: Wu-Jiang." figureDoi="http://doi.org/10.5281/zenodo.14386540" httpUri="https://zenodo.org/record/14386540/files/figure.png" pageId="13" pageNumber="417">Figure 7</figureCitation>
)
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<treatmentCitation id="0A93FE2B9A347E610DA4FB19FC941595" LSID="urn:lsid:zoobank.org:act:C5034BEA-EC81-4BC1-ADA8-E45CB1699B46" author="Chen, S. &amp; Sheraliev, B. &amp; Shu, L. &amp; Peng, Z." box="[151,855,1232,1256]" httpUri="http://treatment.plazi.org/id/13E92C6531415CAA99E72A14E031B734" page="183" pageId="13" pageNumber="417" year="2021">
<taxonomicName id="4C32A3B99A347E610DA4FB19FC941595" authority="Chen, Sheraliev, Shu &amp; Peng, 2021: 183" authorityName="Chen, Sheraliev, Shu &amp; Peng" authorityPageNumber="183" authorityYear="2021" box="[151,855,1232,1256]" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="species" species="wulongensis">
<emphasis id="B94604289A347E610DA4FB19FE5A1595" box="[151,409,1232,1256]" italics="true" pageId="13" pageNumber="417">Triplophysa wulongensis</emphasis>
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(
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locality: 
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: 
<collectingRegion id="49F616D89A347E610960FB19FB0B1595" box="[1107,1224,1232,1256]" country="China" name="Chongqing" pageId="13" pageNumber="417">Chongqing</collectingRegion>
: 
<collectingCounty id="62ECA0B69A347E6109E5FB19FA411595" box="[1238,1410,1232,1256]" pageId="13" pageNumber="417">Wulong County</collectingCounty>
: a subterranean pool in 
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).
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</paragraph>
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<emphasis id="B94604289A347E610DA4FAFDFE5D1432" bold="true" box="[151,414,1332,1359]" pageId="13" pageNumber="417">Specimens examined.</emphasis>
<materialsCitation id="3B5AD2679A347E610C9FFAFCFDDB140E" collectionCode="SWU" country="China" county="Wulong County" location="subterranean pool in Furong Cave" municipality="Chongqing City" pageId="13" pageNumber="417" specimenCode="SWU2019051309" specimenCount="1" typeStatus="holotype">
<specimenCode id="DB9470419A347E610C9FFAFCFDBA142D" box="[428,633,1333,1360]" pageId="13" pageNumber="417">SWU2019051309</specimenCode>
, 
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, 64.0 mm SL; 
<collectingCountry id="F32598AA9A347E610E81FAFCFC3A1432" box="[946,1017,1333,1359]" name="China" pageId="13" pageNumber="417">China</collectingCountry>
: 
<collectingMunicipality id="6BE942409A347E610939FAFCFB091432" box="[1034,1226,1333,1359]" pageId="13" pageNumber="417">Chongqing City</collectingMunicipality>
: 
<collectingCounty id="62ECA0B69A347E6109E8FAFCFA5A1432" box="[1243,1433,1333,1359]" pageId="13" pageNumber="417">Wulong County</collectingCounty>
: 
<location id="8EED8EE19A347E610DA4FA90FDDB140E" LSID="urn:lsid:plazi:treatment:039B692C9A347E620DA4FBAEFA8016A0:8EED8EE19A347E610DA4FA90FDDB140E" box="[151,536,1369,1395]" country="China" county="Wulong County" municipality="Chongqing City" name="subterranean pool in Furong Cave" pageId="13" pageNumber="417">subterranean pool in Furong Cave</location>
</materialsCitation>
. 
<materialsCitation id="3B5AD2679A347E610F10FA90FE1114EA" collectionCode="SWU" country="China" county="Wulong County" location="subterranean pool in Furong Cave" municipality="Chongqing City" pageId="13" pageNumber="417" specimenCode="SWU2019051301-2019051308" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB9470419A347E610F10FA90FC491409" box="[547,906,1369,1396]" pageId="13" pageNumber="417">SWU2019051301–2019051308</specimenCode>
, 
<specimenCount id="9D3413B39A347E610EA6FA90FC0E140E" box="[917,973,1369,1395]" pageId="13" pageNumber="417" type="generic">eight</specimenCount>
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specimens, 49.0– 
<quantity id="4CCA75DF9A347E61083AFA90FAAE140E" box="[1289,1389,1369,1395]" metricMagnitude="-2" metricUnit="m" metricValue="6.72" pageId="13" pageNumber="417" unit="mm" value="67.2">67.2 mm</quantity>
SL; other data same as holotype
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.
</paragraph>
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<subSubSection id="C3288BB19A347E610DF4FA68FCDB17EE" pageId="13" pageNumber="417" type="diagnosis">
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<emphasis id="B94604289A347E610DF4FA68FE8114C6" bold="true" box="[199,322,1441,1467]" pageId="13" pageNumber="417">Diagnosis.</emphasis>
<taxonomicName id="4C32A3B99A347E610C7BFA68FDD814C6" baseAuthorityName="Chen, Sheraliev, Shu &amp; Peng" baseAuthorityYear="2021" box="[328,539,1441,1467]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="species" species="wulongensis">
<emphasis id="B94604289A347E610C7BFA68FDD814C6" box="[328,539,1441,1467]" italics="true" pageId="13" pageNumber="417">Claea wulongensis</emphasis>
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is distinguished from two generic species by the presence of a wider interorbital space (width 38.5–43.1% of HL vs. 20.0– 
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<taxonomicName id="4C32A3B99A347E610FFEFA0CFCFB14A2" baseAuthorityName="Sauvage" baseAuthorityYear="1874" box="[717,824,1477,1503]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="species" species="dabryi">
<emphasis id="B94604289A347E610FFEFA0CFCFB14A2" box="[717,824,1477,1503]" italics="true" pageId="13" pageNumber="417">C. dabryi</emphasis>
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, and 
<quantity id="4CCA75DF9A347E610E4AFA0CFBCB14A2" box="[889,1032,1477,1503]" metricMagnitude="-1" metricUnit="m" metricValue="8.4582" metricValueMax="9.2202" metricValueMin="7.6962" pageId="13" pageNumber="417" unit="in" value="33.3" valueMax="36.3" valueMin="30.3">30.3–36.3 in</quantity>
<taxonomicName id="4C32A3B99A347E610923FA0CFB6B14A2" authorityName="Zhang &amp; Luo &amp; Huang &amp; Zhang" authorityYear="2024" box="[1040,1192,1477,1503]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="species" species="minibarba">
<emphasis id="B94604289A347E610923FA0CFB6B14A2" box="[1040,1192,1477,1503]" italics="true" pageId="13" pageNumber="417">C. minibarba</emphasis>
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; 
<figureCitation id="1309C4BF9A347E61098BFA0CFAE414A2" box="[1208,1319,1477,1503]" captionStart="FIGURE 3" captionStartId="6.[151,250,1941,1965]" captionTargetBox="[160,1427,380,1921]" captionTargetId="figure-53@6.[151,1436,371,1930]" captionTargetPageId="6" captionText="FIGURE 3. Relationship between interorbital width and head length (a); body depth and standard length(b); outer rostral-barbel length and head length (c); maxillary-barbel length and head length (d); caudal-peduncle depth and standard length (e) for Claea dabryi (blue square), C. minibarba sp. nov. (red circle), and C. wulongensis (green triangle)." figureDoi="http://doi.org/10.5281/zenodo.14386530" httpUri="https://zenodo.org/record/14386530/files/figure.png" pageId="13" pageNumber="417">Figure 3a</figureCitation>
). It, along with 
<taxonomicName id="4C32A3B99A347E610DE0FA20FEAF177E" authorityName="Zhang &amp; Luo &amp; Huang &amp; Zhang" authorityYear="2024" box="[211,364,1513,1539]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="species" species="minibarba">
<emphasis id="B94604289A347E610DE0FA20FEAF177E" box="[211,364,1513,1539]" italics="true" pageId="13" pageNumber="417">C. minibarba</emphasis>
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, is distinct from 
<taxonomicName id="4C32A3B99A347E610F05FA20FD60177E" baseAuthorityName="Sauvage" baseAuthorityYear="1874" box="[566,675,1513,1539]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="species" species="dabryi">
<emphasis id="B94604289A347E610F05FA20FD60177E" box="[566,675,1513,1539]" italics="true" pageId="13" pageNumber="417">C. dabryi</emphasis>
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in having shorter outer rostral barbels extending away from (vs. beyond) the anterior margin of eye and shorter maxillary barbels extending to the anterior margin of eye (vs. beyond the posterior edge of eye). This species is further distinct from 
<taxonomicName id="4C32A3B99A347E610E51F9F8FC3A1736" authorityName="Zhang &amp; Luo &amp; Huang &amp; Zhang" authorityYear="2024" box="[866,1017,1585,1611]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="species" species="minibarba">
<emphasis id="B94604289A347E610E51F9F8FC3A1736" box="[866,1017,1585,1611]" italics="true" pageId="13" pageNumber="417">C. minibarba</emphasis>
</taxonomicName>
in having a lower count of vertebrae (4+38–39 vs. 4+41–43), a shallower body (depth 9.3–13.6% of SLvs. 14.7–17.6; 
<figureCitation id="1309C4BF9A347E61092DF99CFB481712" box="[1054,1163,1621,1647]" captionStart="FIGURE 3" captionStartId="6.[151,250,1941,1965]" captionTargetBox="[160,1427,380,1921]" captionTargetId="figure-53@6.[151,1436,371,1930]" captionTargetPageId="6" captionText="FIGURE 3. Relationship between interorbital width and head length (a); body depth and standard length(b); outer rostral-barbel length and head length (c); maxillary-barbel length and head length (d); caudal-peduncle depth and standard length (e) for Claea dabryi (blue square), C. minibarba sp. nov. (red circle), and C. wulongensis (green triangle)." figureDoi="http://doi.org/10.5281/zenodo.14386530" httpUri="https://zenodo.org/record/14386530/files/figure.png" pageId="13" pageNumber="417">Figure 3b</figureCitation>
), and a shallower caudal peduncle (depth 7.6–9.4% of SL vs. 9.5–11.7; 
<figureCitation id="1309C4BF9A347E610FAEF9B0FCC917EE" box="[669,778,1657,1683]" captionStart="FIGURE 3" captionStartId="6.[151,250,1941,1965]" captionTargetBox="[160,1427,380,1921]" captionTargetId="figure-53@6.[151,1436,371,1930]" captionTargetPageId="6" captionText="FIGURE 3. Relationship between interorbital width and head length (a); body depth and standard length(b); outer rostral-barbel length and head length (c); maxillary-barbel length and head length (d); caudal-peduncle depth and standard length (e) for Claea dabryi (blue square), C. minibarba sp. nov. (red circle), and C. wulongensis (green triangle)." figureDoi="http://doi.org/10.5281/zenodo.14386530" httpUri="https://zenodo.org/record/14386530/files/figure.png" pageId="13" pageNumber="417">Figure 3e</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3288BB19A347E610DF4F954FACB17CA" box="[199,1288,1693,1719]" pageId="13" pageNumber="417" type="distribution">
<paragraph id="8B8DD83A9A347E610DF4F954FACB17CA" blockId="13.[151,1437,1332,2043]" box="[199,1288,1693,1719]" pageId="13" pageNumber="417">
<emphasis id="B94604289A347E610DF4F954FEA217CA" bold="true" box="[199,353,1693,1719]" pageId="13" pageNumber="417">Distribution.</emphasis>
Known only from a pool in Furong Cave, connected to the Wu-Jiang near Wulong.
</paragraph>
</subSubSection>
<subSubSection id="C3288BB19A347E620DF4F908FA8016A0" lastPageId="14" lastPageNumber="418" pageId="13" pageNumber="417" type="discussion">
<paragraph id="8B8DD83A9A347E620DF4F908FB0C1184" blockId="13.[151,1437,1332,2043]" lastBlockId="14.[151,1437,151,2013]" lastPageId="14" lastPageNumber="418" pageId="13" pageNumber="417">
<emphasis id="B94604289A347E610DF4F908FE8F17A6" bold="true" box="[199,332,1729,1755]" pageId="13" pageNumber="417">Discussion.</emphasis>
Despite the generic recognition of 
<taxonomicName id="4C32A3B99A347E610FE5F908FCDA17A6" authorityName="Kottelat" authorityYear="2011" box="[726,793,1729,1755]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E610FE5F908FCDA17A6" box="[726,793,1729,1755]" italics="true" pageId="13" pageNumber="417">Claea</emphasis>
</taxonomicName>
in the 
<taxonomicName id="4C32A3B99A347E610E56F908FBCE17A6" box="[869,1037,1729,1755]" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
(
<bibRefCitation id="EFA3A5CB9A347E610928F908FB0617A6" author="Kottelat, M." box="[1051,1221,1729,1755]" pageId="13" pageNumber="417" pagination="1 - 199" refId="ref10940" refString="Kottelat, M. (2012 a) Conspectus cobitidum: An inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). The Raffles Bulletin of Zoology, 26, 1 - 199." type="journal article" year="2012">Kottelat 2012a</bibRefCitation>
; 
<bibRefCitation id="EFA3A5CB9A347E6109E3F908FF0C1782" author="Tan, M. &amp; Armbruster, J. W." pageId="13" pageNumber="417" pagination="6 - 39" refId="ref12378" refString="Tan, M. &amp; Armbruster, J. W. (2018) Phylogenetic classification of extant genera of fishes of the order Cypriniformes (Teleostei: Ostariophysi). Zootaxa, 4476 (1), 6 - 39. https: // doi. org / 10.11646 / zootaxa. 4476.1.4" type="journal article" year="2018">Tan &amp; Armbruster 2018</bibRefCitation>
), its phylogenetic position within this family remains undetermined. 
<taxonomicName id="4C32A3B99A347E610ED8F92CFBF51782" authorityName="Sauvage" authorityYear="1874" box="[1003,1078,1765,1791]" family="Nemacheilidae" genus="Oreias" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="419" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E610ED8F92CFBF51782" box="[1003,1078,1765,1791]" italics="true" pageId="13" pageNumber="417">Oreias</emphasis>
</taxonomicName>
(so far replaced by 
<taxonomicName id="4C32A3B99A347E61082FF92CFA9C1782" authorityName="Kottelat" authorityYear="2011" box="[1308,1375,1765,1791]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E61082FF92CFA9C1782" box="[1308,1375,1765,1791]" italics="true" pageId="13" pageNumber="417">Claea</emphasis>
</taxonomicName>
) was rendered synonymous to 
<taxonomicName id="4C32A3B99A347E610C8DF8C0FD7A165E" authority="(Zhu 1989)" baseAuthorityName="Zhu" baseAuthorityYear="1989" box="[446,697,1801,1827]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E610C8DF8C0FDEB165E" box="[446,552,1801,1827]" italics="true" pageId="13" pageNumber="417">Schistura</emphasis>
(
<bibRefCitation id="EFA3A5CB9A347E610F09F8C0FD73165E" author="Zhu, S. Q." box="[570,688,1801,1827]" pageId="13" pageNumber="417" pagination="55 - 56" refId="ref13528" refString="Zhu, S. Q. (1989) The loaches of the subfamily Nemacheilinae in China (Cypriniformes: Cobitidae) Jiangsu Science and Technology Publishing House, Nanjing, pp. 55 - 56." type="book chapter" year="1989">Zhu 1989</bibRefCitation>
)
</taxonomicName>
; however, this synonymization was refuted as its 
<typeStatus id="548966989A347E610823F8C0FA83165E" box="[1296,1344,1801,1827]" pageId="13" pageNumber="417">type</typeStatus>
species possesses has no pelvic axillary lobe, a character diagnostic for 
<taxonomicName id="4C32A3B99A347E610E7CF8E4FB43163A" authority="(Prokofiev 2009)" baseAuthorityName="Prokofiev" baseAuthorityYear="2009" box="[847,1152,1837,1863]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E610E7CF8E4FC7A163A" box="[847,953,1837,1863]" italics="true" pageId="13" pageNumber="417">Schistura</emphasis>
(
<bibRefCitation id="EFA3A5CB9A347E610EF6F8E4FBBB163A" author="Prokofiev, A. M." box="[965,1144,1837,1863]" pageId="13" pageNumber="417" pagination="874 - 898" refId="ref11855" refString="Prokofiev, A. M. (2009) Problems of the classification and phylogeny of Nemacheiline loaches of the group lacking the preethmoid I (Cypriniformes: Balitoridae: Nemacheilinae). Journal of Ichthyology, 49 (10), 874 - 898. https: // doi. org / 10.1134 / S 0032945209100051" type="journal article" year="2009">Prokofiev 2009</bibRefCitation>
)
</taxonomicName>
. In the revision of loaches of the world by 
<bibRefCitation id="EFA3A5CB9A347E610C7AF898FD631611" author="Banarescu, P. M. &amp; Nalbant, T. T." box="[329,672,1873,1900]" pageId="13" pageNumber="417" pagination="429 - 496" refId="ref9579" refString="Banarescu, P. M. &amp; Nalbant, T. T. (1995) A generical classification of Nemacheilinae with description of two new genera (Teleostei: Cypriniformes: Cobitidae). Travaux du Museum d'Histoire Naturelle &quot; Grigore Antipa &quot;, 35, 429 - 496." type="journal article" year="1995">Bănărescu and Nalbant (1995)</bibRefCitation>
, 
<taxonomicName id="4C32A3B99A347E610F9FF898FD341616" authorityName="Sauvage" authorityYear="1874" box="[684,759,1873,1899]" family="Nemacheilidae" genus="Oreias" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="419" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E610F9FF898FD341616" box="[684,759,1873,1899]" italics="true" pageId="13" pageNumber="417">Oreias</emphasis>
</taxonomicName>
was considered to bear a closer relationship with 
<taxonomicName id="4C32A3B99A347E610816F898FA561616" authorityName="Nichols" authorityYear="1925" box="[1317,1429,1873,1899]" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E610816F898FA561616" box="[1317,1429,1873,1899]" italics="true" pageId="13" pageNumber="417">Homatula</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C32A3B99A347E610DA4F8BCFD3F16F2" authority="McClelland, Nun Banarescu &amp; Nalbant" authorityName="McClelland, Nun Banarescu &amp; Nalbant" box="[151,764,1909,1935]" family="Nemacheilidae" genus="Longischistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E610DA4F8BCFEFC16F2" box="[151,319,1909,1935]" italics="true" pageId="13" pageNumber="417">Longischistura</emphasis>
McClelland, 
<emphasis id="B94604289A347E610CE4F8BFFDC516F2" box="[471,518,1910,1935]" italics="true" pageId="13" pageNumber="417">Nun</emphasis>
Bănărescu &amp; Nalbant
</taxonomicName>
, 
<taxonomicName id="4C32A3B99A347E610E3BF8BCFC2916F2" authority="Bleeker" authorityName="Bleeker" authorityYear="1863" box="[776,1002,1909,1935]" family="Nemacheilidae" genus="Paracobitis" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E610E3BF8BCFC4816F2" box="[776,907,1909,1935]" italics="true" pageId="13" pageNumber="417">Paracobitis</emphasis>
Bleeker
</taxonomicName>
, 
<taxonomicName id="4C32A3B99A347E610EC7F8BCFA5A16F2" authority="Banarescu &amp; Nalbant" authorityName="Banarescu &amp; Nalbant" box="[1012,1433,1909,1935]" family="Nemacheilidae" genus="Physoschistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E610EC7F8BCFB5C16F2" box="[1012,1183,1909,1935]" italics="true" pageId="13" pageNumber="417">Physoschistura</emphasis>
Bănărescu &amp; Nalbant
</taxonomicName>
, 
<taxonomicName id="4C32A3B99A347E610DA4F850FEC216CE" baseAuthorityName="Prokofiev" baseAuthorityYear="2009" box="[151,257,1945,1971]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E610DA4F850FEC216CE" box="[151,257,1945,1971]" italics="true" pageId="13" pageNumber="417">Schistura</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C32A3B99A347E610C73F850FE3D16CE" authority="Kottelat" authorityName="Kottelat" authorityYear="1990" box="[320,510,1945,1971]" family="Nemacheilidae" genus="Sectoria" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E610C73F850FE5E16CE" box="[320,413,1945,1971]" italics="true" pageId="13" pageNumber="417">Sectoria</emphasis>
Kottelat
</taxonomicName>
for having a similar body color pattern, a processus dentiformis on the upper jaw, and no sexual dimorphism in breeding tubercles. Moreover, Liu 
<emphasis id="B94604289A347E610E5DF877FC6316AA" box="[878,928,1981,2007]" italics="true" pageId="13" pageNumber="417">et al</emphasis>
.’s (2012) molecular phylogenetic analysis of the 
<taxonomicName id="4C32A3B99A347E610D8CF828FEA41686" box="[191,359,2017,2043]" family="Nemacheilidae" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="family">Nemacheilidae</taxonomicName>
revealed that the 
<typeStatus id="548966989A347E610F1BF828FD9B1686" box="[552,600,2017,2043]" pageId="13" pageNumber="417">type</typeStatus>
species of 
<taxonomicName id="4C32A3B99A347E610FE2F828FCD71686" authorityName="Kottelat" authorityYear="2011" box="[721,788,2017,2043]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E610FE2F828FCD71686" box="[721,788,2017,2043]" italics="true" pageId="13" pageNumber="417">Claea</emphasis>
</taxonomicName>
(under the name of 
<taxonomicName id="4C32A3B99A347E610EDCF828FB9A1686" baseAuthorityName="Prokofiev" baseAuthorityYear="2009" box="[1007,1113,2017,2043]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="417" phylum="Chordata" rank="genus">
<emphasis id="B94604289A347E610EDCF828FB9A1686" box="[1007,1113,2017,2043]" italics="true" pageId="13" pageNumber="417">Schistura</emphasis>
</taxonomicName>
) was clustered with analysed species of 
<taxonomicName id="4C32A3B99A377E620C3FFF5EFE5111CC" authorityName="Rendahl" authorityYear="1933" box="[268,402,151,177]" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620C3FFF5EFE5111CC" box="[268,402,151,177]" italics="true" pageId="14" pageNumber="418">Triplophysa</emphasis>
</taxonomicName>
. The same finding was also repeated in Chen 
<emphasis id="B94604289A377E620EBEFF5EFC7D11CC" box="[909,958,151,177]" italics="true" pageId="14" pageNumber="418">et al</emphasis>
.’s (2019) molecular phylogenetic analysis. However, their species identification of 
<taxonomicName id="4C32A3B99A377E620F69FF72FD5E11A8" authorityName="Kottelat" authorityYear="2011" box="[602,669,187,213]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620F69FF72FD5E11A8" box="[602,669,187,213]" italics="true" pageId="14" pageNumber="418">Claea</emphasis>
</taxonomicName>
was not free from question. Therefore, their molecular evidence for the closer relationship of this genus with 
<taxonomicName id="4C32A3B99A377E620F57FF16FD291184" authorityName="Rendahl" authorityYear="1933" box="[612,746,223,249]" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620F57FF16FD291184" box="[612,746,223,249]" italics="true" pageId="14" pageNumber="418">Triplophysa</emphasis>
</taxonomicName>
was not conclusive (see elobration below).
</paragraph>
<paragraph id="8B8DD83A9A377E620DF4FECAFBA613F8" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="418">
Our molecular phylogenetic analysis unveiled that 
<taxonomicName id="4C32A3B99A377E620FC8FECAFCFD1060" authorityName="Kottelat" authorityYear="2011" box="[763,830,259,285]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620FC8FECAFCFD1060" box="[763,830,259,285]" italics="true" pageId="14" pageNumber="418">Claea</emphasis>
</taxonomicName>
was more closely allied to 
<taxonomicName id="4C32A3B99A377E620959FECAFB331060" authorityName="Rendahl" authorityYear="1933" box="[1130,1264,259,285]" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620959FECAFB331060" box="[1130,1264,259,285]" italics="true" pageId="14" pageNumber="418">Triplophysa</emphasis>
</taxonomicName>
than 
<taxonomicName id="4C32A3B99A377E62081FFECAFA5F1060" authorityName="Nichols" authorityYear="1925" box="[1324,1436,259,285]" family="Nemacheilidae" genus="Homatula" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E62081FFECAFA5F1060" box="[1324,1436,259,285]" italics="true" pageId="14" pageNumber="418">Homatula</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C32A3B99A377E620D89FEEEFEE7103C" baseAuthorityName="Prokofiev" baseAuthorityYear="2009" box="[186,292,295,321]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620D89FEEEFEE7103C" box="[186,292,295,321]" italics="true" pageId="14" pageNumber="418">Schistura</emphasis>
</taxonomicName>
in the cyt 
<emphasis id="B94604289A377E620C99FEEEFE7B103C" box="[426,440,295,321]" italics="true" pageId="14" pageNumber="418">b</emphasis>
gene-based BI tree (
<figureCitation id="1309C4BF9A377E620F83FEEEFCD0103C" box="[688,787,295,321]" captionStart="FIGURE 1" captionStartId="4.[151,250,1662,1686]" captionTargetBox="[151,1427,556,1626]" captionTargetId="figure-183@4.[151,1436,546,1636]" captionTargetPageId="4" captionText="FIGURE 1. Bayesian Inference tree generated from the cyt b gene for sampled species of the Nemacheilidae. Nodal numbers are posterior probability values greater than 50%." figureDoi="http://doi.org/10.5281/zenodo.14386524" httpUri="https://zenodo.org/record/14386524/files/figure.png" pageId="14" pageNumber="418">Figure 1</figureCitation>
). In Liu 
<emphasis id="B94604289A377E620EB7FEEEFC79103C" box="[900,954,295,321]" italics="true" pageId="14" pageNumber="418">et al</emphasis>
.’s (2012) phylogenetic analysis based on concatenated mitochondrial and nuclear sequences, 
<taxonomicName id="4C32A3B99A377E620FECFE82FC211018" authorityName="Prokofiev" authorityYear="2002" box="[735,994,331,357]" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="gundriseri">
<emphasis id="B94604289A377E620FECFE82FC211018" box="[735,994,331,357]" italics="true" pageId="14" pageNumber="418">Triplophysa gundriseri</emphasis>
</taxonomicName>
was demonstrated to be closely related to 
<taxonomicName id="4C32A3B99A377E620D87FEA6FEAD10F4" baseAuthorityName="Sauvage" baseAuthorityYear="1874" box="[180,366,367,393]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="dabryi">
<emphasis id="B94604289A377E620D87FEA6FEAD10F4" box="[180,366,367,393]" italics="true" pageId="14" pageNumber="418">Schistura dabryi</emphasis>
</taxonomicName>
from the Tuo-Jiang of the upper Chang-Jiang basin. The recognition of their samples under the name of 
<taxonomicName id="4C32A3B99A377E620DCAFE5AFE9D10D0" baseAuthorityName="Sauvage" baseAuthorityYear="1874" box="[249,350,403,429]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="dabryi">
<emphasis id="B94604289A377E620DCAFE5AFE9D10D0" box="[249,350,403,429]" italics="true" pageId="14" pageNumber="418">S. dabryi</emphasis>
</taxonomicName>
was unlikely under molecular scrutiny of our analysis due to no use of the cyt 
<emphasis id="B94604289A377E6209E0FE5AFB2210D0" box="[1235,1249,403,429]" italics="true" pageId="14" pageNumber="418">b</emphasis>
gene as markers of their phylogenetic inference. The species, identified by 
<bibRefCitation id="EFA3A5CB9A377E620E05FE7EFC3710AC" author="Liu, S. &amp; Mayden, R. L. &amp; Zhang, J. &amp; Yu, D. &amp; Tang, Q. &amp; Deng, X. &amp; Liu, H." box="[822,1012,439,465]" pageId="14" pageNumber="418" pagination="60 - 72" refId="ref11504" refString="Liu, S., Mayden, R. L., Zhang, J., Yu, D., Tang, Q., Deng, X. &amp; Liu, H. (2012) Phylogenetic relationships of the Cobitoidea (Teleostei: Cypriniformes) inferred from mitochondrial and nuclear genes with analyses of gene evolution. Gene, 508 (1), 60 - 72. https: // doi. org / 10.1016 / j. gene. 2012.07.040" type="journal article" year="2012">
Liu 
<emphasis id="B94604289A377E620E55FE7EFC5A10AC" box="[870,921,439,465]" italics="true" pageId="14" pageNumber="418">et al</emphasis>
. (2012)
</bibRefCitation>
as 
<taxonomicName id="4C32A3B99A377E62092DFE7EFB4610AC" baseAuthorityName="Sauvage" baseAuthorityYear="1874" box="[1054,1157,439,465]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="dabryi">
<emphasis id="B94604289A377E62092DFE7EFB4610AC" box="[1054,1157,439,465]" italics="true" pageId="14" pageNumber="418">S. dabryi</emphasis>
</taxonomicName>
, may be a misidentified member of 
<taxonomicName id="4C32A3B99A377E620C13FE12FE651088" authorityName="Rendahl" authorityYear="1933" box="[288,422,475,501]" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620C13FE12FE651088" box="[288,422,475,501]" italics="true" pageId="14" pageNumber="418">Triplophysa</emphasis>
</taxonomicName>
, given its closer relationship with 
<taxonomicName id="4C32A3B99A377E620E0DFE12FC111088" authorityName="Prokofiev" authorityYear="2002" box="[830,978,475,501]" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="gundriseri">
<emphasis id="B94604289A377E620E0DFE12FC111088" box="[830,978,475,501]" italics="true" pageId="14" pageNumber="418">T. gundriseri</emphasis>
</taxonomicName>
. Currently, 
<taxonomicName id="4C32A3B99A377E62096DFE12FB311088" authorityName="Prokofiev" authorityYear="2002" box="[1118,1266,475,501]" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="gundriseri">
<emphasis id="B94604289A377E62096DFE12FB311088" box="[1118,1266,475,501]" italics="true" pageId="14" pageNumber="418">T. gundriseri</emphasis>
</taxonomicName>
is confined to the Tesiin River and Lake Sangiin-Nurr of 
<collectingCountry id="F32598AA9A377E620FB9FE36FD341364" box="[650,759,511,537]" name="Mongolia" pageId="14" pageNumber="418">Mongolia</collectingCountry>
and the Uvs-Nuur Lake basin of 
<collectingRegion id="49F616D89A377E6209B3FE36FB7A1364" box="[1152,1209,511,537]" country="Russia" name="Tyva" pageId="14" pageNumber="418">Tuva</collectingRegion>
Republic of 
<collectingCountry id="F32598AA9A377E620861FE36FA5E1364" box="[1362,1437,511,537]" name="Russia" pageId="14" pageNumber="418">Russia</collectingCountry>
(Kottelat 2006). It is here posited to be more closely allied to species of Sbuclade 2 than Subclade 1. Despite the only use of the cyt 
<emphasis id="B94604289A377E620C5EFD8EFEB8131C" box="[365,379,583,609]" italics="true" pageId="14" pageNumber="418">b</emphasis>
gene, more species of 
<taxonomicName id="4C32A3B99A377E620F48FD8EFCC2131C" authorityName="Rendahl" authorityYear="1933" box="[635,769,583,609]" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620F48FD8EFCC2131C" box="[635,769,583,609]" italics="true" pageId="14" pageNumber="418">Triplophysa</emphasis>
</taxonomicName>
were sampled in our molecular analysis than in two studies aforementioned. Further research will confirm that the result of our analysis is reliable.
</paragraph>
<paragraph id="8B8DD83A9A377E620DF4FD46FB261244" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="418">
It is worth mentioning that sampled species of 
<taxonomicName id="4C32A3B99A377E620FC7FD46FCB913D4" authorityName="Rendahl" authorityYear="1933" box="[756,890,655,681]" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620FC7FD46FCB913D4" box="[756,890,655,681]" italics="true" pageId="14" pageNumber="418">Triplophysa</emphasis>
</taxonomicName>
, detected to nest with species of 
<taxonomicName id="4C32A3B99A377E62083BFD46FA8813D4" authorityName="Kottelat" authorityYear="2011" box="[1288,1355,655,681]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E62083BFD46FA8813D4" box="[1288,1355,655,681]" italics="true" pageId="14" pageNumber="418">Claea</emphasis>
</taxonomicName>
in our molecular phylogenetic analysis (
<figureCitation id="1309C4BF9A377E620F2DFD7AFD4113B0" box="[542,642,691,717]" captionStart="FIGURE 1" captionStartId="4.[151,250,1662,1686]" captionTargetBox="[151,1427,556,1626]" captionTargetId="figure-183@4.[151,1436,546,1636]" captionTargetPageId="4" captionText="FIGURE 1. Bayesian Inference tree generated from the cyt b gene for sampled species of the Nemacheilidae. Nodal numbers are posterior probability values greater than 50%." figureDoi="http://doi.org/10.5281/zenodo.14386524" httpUri="https://zenodo.org/record/14386524/files/figure.png" pageId="14" pageNumber="418">Figure 1</figureCitation>
), are troglobitic and found in Southwest 
<collectingCountry id="F32598AA9A377E620947FD7AFB7413B0" box="[1140,1207,691,717]" name="China" pageId="14" pageNumber="418">China</collectingCountry>
(primarily 
<collectingRegion id="49F616D89A377E620873FD7AFA5A13B0" box="[1344,1433,691,717]" country="China" name="Yunnan" pageId="14" pageNumber="418">Yunnan</collectingRegion>
, 
<collectingRegion id="49F616D89A377E620DA4FD1EFF34138C" box="[151,247,727,753]" country="China" name="Guangxi" pageId="14" pageNumber="418">Guangxi</collectingRegion>
and 
<collectingRegion id="49F616D89A377E620C03FD1EFE53138C" box="[304,400,727,753]" country="China" name="Guizhou" pageId="14" pageNumber="418">Guizhou</collectingRegion>
Provinces), with the largest continuous distribution of karst landforms in 
<collectingCountry id="F32598AA9A377E6209EBFD1EFADD138C" box="[1240,1310,727,753]" name="China" pageId="14" pageNumber="418">China</collectingCountry>
, or even in the world. Habitats of these species are quite distinct from those of epigean congeneric species. Whether or not these subterranean species represent a distinct genus from 
<taxonomicName id="4C32A3B99A377E620FD6FCD6FCA81244" authorityName="Rendahl" authorityYear="1933" box="[741,875,799,825]" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620FD6FCD6FCA81244" box="[741,875,799,825]" italics="true" pageId="14" pageNumber="418">Triplophysa</emphasis>
</taxonomicName>
require further in-depth research.
</paragraph>
<paragraph id="8B8DD83A9A377E620DF4FC8AFAD714BC" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="418">
The type locality of 
<taxonomicName id="4C32A3B99A377E620C98FC8AFDD61220" baseAuthorityName="Sauvage" baseAuthorityYear="1874" box="[427,533,835,861]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="dabryi">
<emphasis id="B94604289A377E620C98FC8AFDD61220" box="[427,533,835,861]" italics="true" pageId="14" pageNumber="418">C. dabryi</emphasis>
</taxonomicName>
is Dengchigou, Baoxing County, 
<collectingRegion id="49F616D89A377E620EA1FC8AFB991220" box="[914,1114,835,861]" country="China" name="Sichuan" pageId="14" pageNumber="418">Sichuan Province</collectingRegion>
, in the Qingyi-Jiang flowing into the Min-Jiang of the upper Chang-Jiang basin (
<bibRefCitation id="EFA3A5CB9A377E620FCEFCAEFC6D12FC" author="Kottelat, M." box="[765,942,871,897]" pageId="14" pageNumber="418" pagination="1 - 199" refId="ref10940" refString="Kottelat, M. (2012 a) Conspectus cobitidum: An inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). The Raffles Bulletin of Zoology, 26, 1 - 199." type="journal article" year="2012">Kottelat 2012a</bibRefCitation>
). The 
<typeStatus id="548966989A377E620EC8FCAEFB9E12FC" box="[1019,1117,871,897]" pageId="14" pageNumber="418" type="holotype">holotype</typeStatus>
(MNHN 6276, photograph examined) is broken or in bad condition, and thus of very limited taxonomic use for its identification. In light of Sauvage’s (1874) original description, 
<taxonomicName id="4C32A3B99A377E620F71FC66FD6912B4" baseAuthorityName="Sauvage" baseAuthorityYear="1874" box="[578,682,943,969]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="dabryi">
<emphasis id="B94604289A377E620F71FC66FD6912B4" box="[578,682,943,969]" italics="true" pageId="14" pageNumber="418">C. dabryi</emphasis>
</taxonomicName>
has small eyes located in the middle of head, a flat interorbital space, with its width nearly twice the diameter of eye; pelvic fins inserted slightly in front of the dorsal-fin orgin; anal-fin origin almost equidistant from the ventral fin and caudal-fin base; and upper portion of body yellowish, mottled with dark green blotches. These characters are shared with a specimen reported and figured by 
<bibRefCitation id="EFA3A5CB9A377E6209A4FBD2FA981548" author="Guo, Y. S. &amp; Sun, Z. Y. &amp; He, X. H. &amp; Jin, W. &amp; Chen, Y. L." box="[1175,1371,1051,1077]" pageId="14" pageNumber="418" pagination="505 - 507" refId="ref10389" refString="Guo, Y. S., Sun, Z. Y., He, X. H., Jin, W. &amp; Chen, Y. L. (2021) Coloured atlas of fishes in Sichuan. Science Press, Beijing, pp. 505 - 507." type="book chapter" year="2021">
Guo 
<emphasis id="B94604289A377E6209FCFBD2FAC21548" box="[1231,1281,1051,1077]" italics="true" pageId="14" pageNumber="418">et al</emphasis>
. (2021)
</bibRefCitation>
as 
<taxonomicName id="4C32A3B99A377E6208B0FBD2FF231524" baseAuthorityName="Sauvage" baseAuthorityYear="1874" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="dabryi">
<emphasis id="B94604289A377E6208B0FBD2FF231524" italics="true" pageId="14" pageNumber="418">C. dabryi</emphasis>
</taxonomicName>
from the Dachuan-He (a part of the Qingyi-Jiang) in Lushan County, about 
<quantity id="4CCA75DF9A377E620918FBF6FBB21524" box="[1067,1137,1087,1113]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="418" unit="km" value="20.0">20 km</quantity>
from Baoxing County (the type locality of the species) of 
<collectingRegion id="49F616D89A377E620CCBFBAAFD021500" box="[504,705,1123,1149]" country="China" name="Sichuan" pageId="14" pageNumber="418">Sichuan Province</collectingRegion>
, and can thus be considered as topotypical; two pictures (Figure 150–1 and 150–2 on Page 505) of the specimen exhibit that 
<taxonomicName id="4C32A3B99A377E620E03FB4EFC5A15DC" baseAuthorityName="Sauvage" baseAuthorityYear="1874" box="[816,921,1159,1185]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="dabryi">
<emphasis id="B94604289A377E620E03FB4EFC5A15DC" box="[816,921,1159,1185]" italics="true" pageId="14" pageNumber="418">C. dabryi</emphasis>
</taxonomicName>
has pelvic fins inserted slightly in front of the dorsal fin, no pelvic axillary lobe, no nasal barbels, and no sexual dimorphism in breeding tubercles, and (anterior and posterior) nostrils set closely. A median processus dentiformis on the upper jaw is illustrated in the other picture (Figure 150–7 on Page 507) for this species. Guo 
<emphasis id="B94604289A377E620FF1FB3AFD311470" box="[706,754,1267,1293]" italics="true" pageId="14" pageNumber="418">et al</emphasis>
.’s (2021) description, predicated on 
<specimenCount id="9D3413B39A377E6209B8FB3AFAE01470" box="[1163,1315,1267,1293]" count="20" pageId="14" pageNumber="418" type="generic">20 specimens</specimenCount>
(including those collected from Baoxing County, the type locality of the species) of the upper Chang-Jiang basin, recorded this species with a depressed head (depth 44.4–55.6% of HL), a shallow body (depth 13.2–16.7% of SL), long maxillary barbels (length 30.4–35.0% of HL), long outer rostral barbels (length 32.6–35.0% of HL), and 4+38-42 vertebrae. Despite no accessibility to topotypical specimens, primary diagnostic characters provided in this study for 
<taxonomicName id="4C32A3B99A377E620800FA4AFA5F14E0" baseAuthorityName="Sauvage" baseAuthorityYear="1874" box="[1331,1436,1411,1437]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="dabryi">
<emphasis id="B94604289A377E620800FA4AFA5F14E0" box="[1331,1436,1411,1437]" italics="true" pageId="14" pageNumber="418">C. dabryi</emphasis>
</taxonomicName>
largely agree with its original description, and also the data given by 
<bibRefCitation id="EFA3A5CB9A377E620EAFFA6EFB9E14BC" author="Guo, Y. S. &amp; Sun, Z. Y. &amp; He, X. H. &amp; Jin, W. &amp; Chen, Y. L." box="[924,1117,1447,1473]" pageId="14" pageNumber="418" pagination="505 - 507" refId="ref10389" refString="Guo, Y. S., Sun, Z. Y., He, X. H., Jin, W. &amp; Chen, Y. L. (2021) Coloured atlas of fishes in Sichuan. Science Press, Beijing, pp. 505 - 507." type="book chapter" year="2021">
Guo 
<emphasis id="B94604289A377E620EE7FA6EFBC614BC" box="[980,1029,1447,1473]" italics="true" pageId="14" pageNumber="418">et al</emphasis>
. (2021)
</bibRefCitation>
for this species.
</paragraph>
<paragraph id="8B8DD83A9A377E620DF4FA02FDB41630" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="418">
<taxonomicName id="4C32A3B99A377E620DF4FA02FE301498" authorityName="Chen, Lu &amp; Mao" authorityYear="2006" box="[199,499,1483,1509]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="niulanjiangensis">
<emphasis id="B94604289A377E620DF4FA02FE301498" box="[199,499,1483,1509]" italics="true" pageId="14" pageNumber="418">Schistura niulanjiangensis</emphasis>
</taxonomicName>
, a species originally described from the Niulan-Jiang discharging into the Jinsha-Jiang of the upper Chang-Jiang basin (
<bibRefCitation id="EFA3A5CB9A377E620F75FA26FCC61774" author="Chen, L. &amp; Lu, Z. &amp; Mao, W." box="[582,773,1519,1545]" pageId="14" pageNumber="418" pagination="54 - 55" refId="ref9746" refString="Chen, L., Lu, Z. &amp; Mao, W. (2006) A new species of Schistura discovered in Yunnan, China. Guizhou Agricultural Sciences, 34 (5), 54 - 55." type="journal article" year="2006">
Chen 
<emphasis id="B94604289A377E620FB9FA26FD781774" box="[650,699,1519,1545]" italics="true" pageId="14" pageNumber="418">et al</emphasis>
. 2006
</bibRefCitation>
), is excluded from 
<taxonomicName id="4C32A3B99A377E620EEFFA26FBDC1774" authorityName="Kottelat" authorityYear="2011" box="[988,1055,1519,1545]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620EEFFA26FBDC1774" box="[988,1055,1519,1545]" italics="true" pageId="14" pageNumber="418">Claea</emphasis>
</taxonomicName>
in this study. It was transferred to this genus, but without detailed explanation (
<bibRefCitation id="EFA3A5CB9A377E620FB7F9DAFCEC1750" author="Kottelat, M." box="[644,815,1555,1581]" pageId="14" pageNumber="418" pagination="1 - 199" refId="ref10940" refString="Kottelat, M. (2012 a) Conspectus cobitidum: An inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei). The Raffles Bulletin of Zoology, 26, 1 - 199." type="journal article" year="2012">Kottelat 2012a</bibRefCitation>
). Our photographic examination on the 
<typeStatus id="548966989A377E6209D1F9DAFA871750" box="[1250,1348,1555,1581]" pageId="14" pageNumber="418" type="holotype">holotype</typeStatus>
(
<figureCitation id="1309C4BF9A377E620862F9DAFF76172C" captionStart="FIGURE 12" captionStartId="15.[151,250,772,796]" captionTargetBox="[258,1330,179,746]" captionTargetId="figure-25@15.[258,1330,179,746]" captionTargetPageId="15" captionText="FIGURE 12. Lateral view of body in “Claea” niulanjiangensis (upper), holotype, Qujing Aquaculture Station 9206085, 84.0 mm SL, and Claea sp. (lower) from the Niulan-Jiang of the upper Chang-Jiang mainstream (Photographed by Wei-Ning Mao and Zong-Min Lu)." figureDoi="http://doi.org/10.5281/zenodo.14386553" httpUri="https://zenodo.org/record/14386553/files/figure.png" pageId="14" pageNumber="418">Figure 12</figureCitation>
, upper) indicated that this species, as stated in its original description, possesses no processus dentiformis on the upper jaw, pelvic axillary lobes present, and pelvic fins inserted slightly behind the dorsal-fin origin, three characters not placing it in 
<taxonomicName id="4C32A3B99A377E620C6CF9B6FE6117E4" authorityName="Kottelat" authorityYear="2011" box="[351,418,1663,1689]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620C6CF9B6FE6117E4" box="[351,418,1663,1689]" italics="true" pageId="14" pageNumber="418">Claea</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C32A3B99A377E620C87F9B6FD2117E4" authorityName="Chen, Lu &amp; Mao" authorityYear="2006" box="[436,738,1663,1689]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="niulanjiangensis">
<emphasis id="B94604289A377E620C87F9B6FD2117E4" box="[436,738,1663,1689]" italics="true" pageId="14" pageNumber="418">Schistura niulanjiangensis</emphasis>
</taxonomicName>
is indeed a species of 
<taxonomicName id="4C32A3B99A377E620EC5F9B6FBBF17E4" authorityName="Rendahl" authorityYear="1933" box="[1014,1148,1663,1689]" family="Nemacheilidae" genus="Triplophysa" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620EC5F9B6FBBF17E4" box="[1014,1148,1663,1689]" italics="true" pageId="14" pageNumber="418">Triplophysa</emphasis>
</taxonomicName>
in terms of unpublished molecular data (Feng Lin, personal communication). 
<specimenCount id="9D3413B39A377E620FC9F96AFC5117C0" box="[762,914,1699,1725]" count="1" pageId="14" pageNumber="418" type="generic" typeStatus="paratype">One paratype</specimenCount>
(
<figureCitation id="1309C4BF9A377E620E90F96AFBD617C0" box="[931,1045,1699,1725]" captionStart="FIGURE 12" captionStartId="15.[151,250,772,796]" captionTargetBox="[258,1330,179,746]" captionTargetId="figure-25@15.[258,1330,179,746]" captionTargetPageId="15" captionText="FIGURE 12. Lateral view of body in “Claea” niulanjiangensis (upper), holotype, Qujing Aquaculture Station 9206085, 84.0 mm SL, and Claea sp. (lower) from the Niulan-Jiang of the upper Chang-Jiang mainstream (Photographed by Wei-Ning Mao and Zong-Min Lu)." figureDoi="http://doi.org/10.5281/zenodo.14386553" httpUri="https://zenodo.org/record/14386553/files/figure.png" pageId="14" pageNumber="418">Figure 12</figureCitation>
, lower) of 
<taxonomicName id="4C32A3B99A377E6209A5F96AFAAD17C0" authorityName="Chen, Lu &amp; Mao" authorityYear="2006" box="[1174,1390,1699,1725]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="niulanjiangensis">
<emphasis id="B94604289A377E6209A5F96AFAAD17C0" box="[1174,1390,1699,1725]" italics="true" pageId="14" pageNumber="418">S. niulanjiangensis</emphasis>
</taxonomicName>
has pelvic fins inserted slightly in front of the dorsal-fin orgin as found in 
<taxonomicName id="4C32A3B99A377E620EFBF90EFBC8179C" authorityName="Kottelat" authorityYear="2011" box="[968,1035,1735,1761]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620EFBF90EFBC8179C" box="[968,1035,1735,1761]" italics="true" pageId="14" pageNumber="418">Claea</emphasis>
</taxonomicName>
, but not conforming to its original description. It can not be assigned to this genus due to no processus dentiformis on the upper jaw, and the presence of pelvic anxillary lobes. The generic classification and specific status of this 
<typeStatus id="548966989A377E620ECAF8C6FB9A1654" box="[1017,1113,1807,1833]" pageId="14" pageNumber="418" type="paratype">paratype</typeStatus>
need to be determined in the future when more specimens are available.
</paragraph>
<paragraph id="8B8DD83A9A377E620DF4F89EFA8016A0" blockId="14.[151,1437,151,2013]" pageId="14" pageNumber="418">
<taxonomicName id="4C32A3B99A377E620DF4F89EFEC9160C" authorityName="Kottelat" authorityYear="2011" box="[199,266,1879,1905]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620DF4F89EFEC9160C" box="[199,266,1879,1905]" italics="true" pageId="14" pageNumber="418">Claea</emphasis>
</taxonomicName>
has its representative in the Niulan-Jiang. 
<specimenCount id="9D3413B39A377E620FEFF89EFC58160C" box="[732,923,1879,1905]" count="3" pageId="14" pageNumber="418" type="generic">Three specimens</specimenCount>
, collected by 
<collectorName id="26C7BDEC9A377E620903F89EFB0E160C" box="[1072,1229,1879,1905]" pageId="14" pageNumber="418">Chun-Yun Lei</collectorName>
(
<collectingRegion id="49F616D89A377E6209E9F89EFAF1160C" box="[1242,1330,1879,1905]" country="China" name="Yunnan" pageId="14" pageNumber="418">Yunnan</collectingRegion>
<location id="8EED8EE19A377E620804F89EFE0016E8" LSID="urn:lsid:plazi:treatment:039B692C9A347E620DA4FBAEFA8016A0:8EED8EE19A377E620804F89EFE0016E8" name="Fisheries Science Research Institute" pageId="14" pageNumber="418" stateProvince="Yunnan">Fisheries Science Research Institute</location>
) from the 
<typeStatus id="548966989A377E620F08F8B2FDA816E8" box="[571,619,1915,1941]" pageId="14" pageNumber="418">type</typeStatus>
locality of 
<taxonomicName id="4C32A3B99A377E620FDCF8B2FC0616E8" authorityName="Chen, Lu &amp; Mao" authorityYear="2006" box="[751,965,1915,1941]" family="Nemacheilidae" genus="Schistura" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="species" species="niulanjiangensis">
<emphasis id="B94604289A377E620FDCF8B2FC0616E8" box="[751,965,1915,1941]" italics="true" pageId="14" pageNumber="418">S. niulanjiangensis</emphasis>
</taxonomicName>
, were initially recognized as this species. Our careful examination affirmed that these 
<specimenCount id="9D3413B39A377E620FB6F856FCFA16C4" box="[645,825,1951,1977]" count="3" pageId="14" pageNumber="418" type="generic">three specimens</specimenCount>
indeed represent an unidentified species of 
<taxonomicName id="4C32A3B99A377E620812F856FAA716C4" authorityName="Kottelat" authorityYear="2011" box="[1313,1380,1951,1977]" family="Nemacheilidae" genus="Claea" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="418" phylum="Chordata" rank="genus">
<emphasis id="B94604289A377E620812F856FAA716C4" box="[1313,1380,1951,1977]" italics="true" pageId="14" pageNumber="418">Claea</emphasis>
</taxonomicName>
. The formal description of this unidentified species, however, is hindered by the paucity of available specimens.
</paragraph>
</subSubSection>
</treatment>
</document>