169 lines
20 KiB
XML
169 lines
20 KiB
XML
<document id="7EE7D0D5A377C467BD67A623AACBC927" ID-CLB-Dataset="55978" ID-DOI="10.35929/RSZ.0064" ID-GBIF-Dataset="439e5b25-eb8c-4ffc-a7d3-0670028558d4" ID-ISSN="0035-418" ID-Zenodo-Dep="6461022" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1649936365219" checkinUser="carolina" docAuthor="Löbl, Ivan" docDate="2022" docId="038FEB7AFFE7FFBE6504683CFC04BBA9" docLanguage="en" docName="RevSuisseZool.129.1.103-118.pdf.imf" docOrigin="Revue suisse de Zoologie 129 (1)" docStyle="DocumentStyle:984CC6410E0DE5B23EAFEE8246BF4C6B.2:RevSuisseZool.2020-.journal_article.0cover" docStyleId="984CC6410E0DE5B23EAFEE8246BF4C6B" docStyleName="RevSuisseZool.2020-.journal_article.0cover" docStyleVersion="2" docTitle="Scaphobaeocera plana Löbl 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="115" masterDocId="FFB69302FFEBFFB2641F693CFFB2B961" masterDocTitle="Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas" masterLastPageNumber="118" masterPageNumber="103" pageNumber="115" updateTime="1699328943217" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8604DFA3A51022E062F9E9DFD7CD83E9">Supplement to the knowledge of the Scaphidiinae (Coleoptera: Staphylinidae) of the Moluccas</mods:title>
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</mods:titleInfo>
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<mods:name id="1F54949364BBFE9EE9F0166F6148802A" type="personal">
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<mods:namePart id="C406AE3821AC36371595B769EC6034D8">Löbl, Ivan</mods:namePart>
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<mods:typeOfResource id="C67BEF4AFE4256385B1852A5C92C2B48">text</mods:typeOfResource>
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<mods:title id="B33D804411AFAE446DD3CD0C4B5E96E6">Revue suisse de Zoologie</mods:title>
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<mods:part id="09E5D10640685F770E86D86FC3CE25A8">
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<mods:date id="92C17E5586F96090CE5F46BB7E69B86C">2022</mods:date>
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<mods:detail id="6A4B35E22067848C86A74DA5612A6CDC" type="pubDate">
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<mods:number id="97F196A18118F78D4BC0930FEDA1F3E9">2022-03-31</mods:number>
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<mods:detail id="37658DC1045D9C7C68FCDE6E2614CBF7" type="volume">
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<mods:number id="34C3BDDD655EC4800ED4BE902DC24FFE">129</mods:number>
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<mods:detail id="9B2F22057E0B9C64AD5394F4F614507C" type="issue">
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<mods:start id="0FDCA806D083937649E0002DBE578C29">103</mods:start>
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<mods:end id="0533D78A9B4EE420E20B5F8A0459DDD1">118</mods:end>
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<mods:classification id="DF0C5600B400D2BB18BEE26CFA5673C1">journal article</mods:classification>
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<mods:identifier id="BB733AFB49857A2F586D9741E0A45CE2" type="CLB-Dataset">55978</mods:identifier>
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<mods:identifier id="D090D380D669AE9C01997095955092A2" type="DOI">10.35929/RSZ.0064</mods:identifier>
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<mods:identifier id="44B043427A49A67D28F2EF48334D427F" type="GBIF-Dataset">439e5b25-eb8c-4ffc-a7d3-0670028558d4</mods:identifier>
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<mods:identifier id="1FF0104EB3A4A17DA7F0E19EDD076AB2" type="ISSN">0035-418</mods:identifier>
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<mods:identifier id="DDB30F541F2101ACE834B4F3568E9F48" type="Zenodo-Dep">6461022</mods:identifier>
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<treatment id="038FEB7AFFE7FFBE6504683CFC04BBA9" ID-DOI="http://doi.org/10.5281/zenodo.6461048" ID-GBIF-Taxon="194598637" ID-Zenodo-Dep="6461048" LSID="urn:lsid:plazi:treatment:038FEB7AFFE7FFBE6504683CFC04BBA9" httpUri="http://treatment.plazi.org/id/038FEB7AFFE7FFBE6504683CFC04BBA9" lastPageNumber="115" pageId="12" pageNumber="115">
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<subSubSection id="C33C09E7FFE7FFBE6504683CFDC3B879" box="[283,625,256,280]" pageId="12" pageNumber="115" type="nomenclature">
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<paragraph id="8B995A6CFFE7FFBE6504683CFDC3B879" blockId="12.[283,625,256,312]" box="[283,625,256,280]" pageId="12" pageNumber="115">
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<heading id="D0D1ED00FFE7FFBE6504683CFDC3B879" bold="true" box="[283,625,256,280]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="115" reason="2">
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<emphasis id="B952867EFFE7FFBE6504683CFDC3B879" bold="true" box="[283,625,256,280]" pageId="12" pageNumber="115">
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<taxonomicName id="4C2621EFFFE7FFBE6504683CFDA6B879" ID-CoL="9GD34" authority="Löbl, 2022" authorityName="Löbl" authorityYear="2022" box="[283,532,256,280]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="115" phylum="Arthropoda" rank="species" species="plana" status="sp. nov.">
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<emphasis id="B952867EFFE7FFBE6504683CFDA6B879" bold="true" box="[283,532,256,280]" italics="true" pageId="12" pageNumber="115">Scaphobaeocera plana</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2613B05FFE7FFBE6604683DFDC3B879" box="[539,625,257,280]" pageId="12" pageNumber="115" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C09E7FFE7FFBE6593681DFE4DB859" box="[396,511,289,312]" pageId="12" pageNumber="115" type="description">
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<paragraph id="8B995A6CFFE7FFBE6593681DFE4DB859" blockId="12.[283,625,256,312]" box="[396,511,289,312]" pageId="12" pageNumber="115">
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<figureCitation id="131D46E9FFE7FFBE6593681DFE4DB859" box="[396,511,289,312]" captionStart="Figs 23-29" captionStartId="13.[151,192,1925,1946]" captionTargetBox="[177,1384,257,1874]" captionTargetId="figure-4@13.[157,1440,254,1894]" captionTargetPageId="13" captionText="Figs 23-29. Scaphisoma donati sp. nov. (23) Parameres in ventral view, scale bar = 0.2 mm. Figs 24-27. Scaphobaeocera plana sp. nov. (24, 25) Aedeagus in dorsal and lateral views, scale = 0.1 mm. (26) Paramere in ventral view, scale = 0.05 mm. (27) Apical part of median lobe, in lateral view, scale = 0.05 mm. Figs 28, 29. Scaphoxium lobatum sp. nov. (28) Aedeagus in ventral view, scale = 0.1 mm. (29) Apex of median lobe with extruded internal sac and right paramere, in dorsal view, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6461034" httpUri="https://zenodo.org/record/6461034/files/figure.png" pageId="12" pageNumber="115">Figs 24-27</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C09E7FFE7FFBE6488686CFE46B885" pageId="12" pageNumber="115" type="materials_examined">
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<paragraph id="8B995A6CFFE7FFBE6488686CFEF8B8C4" blockId="12.[151,756,335,484]" pageId="12" pageNumber="115">
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<emphasis id="B952867EFFE7FFBE6488686CFE24B806" bold="true" box="[151,406,336,359]" pageId="12" pageNumber="115">Material examined:</emphasis>
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<materialsCitation id="3B4E5031FFE7FFBE65AB6873FEF8B8C4" ID-GBIF-Occurrence="3751563311" collectingDate="1981-09-04" collectionCode="MHNG, MALUK" collectorName="D. Agosti" country="India" location="Kai Besar" pageId="12" pageNumber="115" specimenCode="F911035" specimenCount="1" typeStatus="holotype">
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<emphasis id="B952867EFFE7FFBE65AB6873FDABB806" box="[436,537,335,359]" italics="true" pageId="12" pageNumber="115">
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<typeStatus id="549DE4CEFFE7FFBE65AB6873FDABB806" box="[436,537,335,359]" pageId="12" pageNumber="115" type="holotype">Holotype</typeStatus>
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</emphasis>
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;
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<collectionCode id="ED37C2A9FFE7FFBE6621686CFD27B806" box="[574,661,336,359]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="12" pageNumber="115" type="Museum">MHNG</collectionCode>
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;
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<specimenCount id="9D2091E5FFE7FFBE66A8686CFD5DB806" box="[695,751,336,359]" pageId="12" pageNumber="115" type="male">male</specimenCount>
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;
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<collectionCode id="ED37C2A9FFE7FFBE64886853FF4FB8E7" box="[151,253,367,390]" pageId="12" pageNumber="115">MALUK</collectionCode>
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:
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<location id="8EF90CB7FFE7FFBE65176853FEC6B8E7" LSID="urn:lsid:plazi:treatment:038FEB7AFFE7FFBE6504683CFC04BBA9:8EF90CB7FFE7FFBE65176853FEC6B8E7" box="[264,372,367,390]" country="India" name="Kai Besar" pageId="12" pageNumber="115">Kai Besar</location>
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,
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<location id="8EF90CB7FFE7FFBE659F6853FDDCB8E7" LSID="urn:lsid:plazi:treatment:038FEB7AFFE7FFBE6504683CFC04BBA9:8EF90CB7FFE7FFBE659F6853FDDCB8E7" box="[384,622,367,390]" country="India" name="G. Tukrau nr Bombay" pageId="12" pageNumber="115">G. Tukrau nr Bombay</location>
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, 4.IX.81,
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<collectorName id="26D33FBAFFE7FFBE66C56853FF6CB8C4" pageId="12" pageNumber="115">D. Agosti</collectorName>
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<specimenCode id="DB80F217FFE7FFBE64FA68B2FEF4B8C4" box="[229,326,398,421]" pageId="12" pageNumber="115">F911035</specimenCode>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8B995A6CFFE7FFBE64886892FE46B885" blockId="12.[151,756,335,484]" pageId="12" pageNumber="115">
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<materialsCitation id="3B4E5031FFE7FFBE64886892FE42B885" ID-GBIF-Occurrence="3751563303" collectingDate="1981-09-05" collectionCode="MHNG" collectorName=" D. Agosti" country="India" location="Bombay" pageId="12" pageNumber="115" specimenCode="F911046" specimenCount="1" typeStatus="paratype">
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<emphasis id="B952867EFFE7FFBE64886892FF48B8A4" box="[151,250,430,453]" italics="true" pageId="12" pageNumber="115">
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<typeStatus id="549DE4CEFFE7FFBE64886892FF48B8A4" box="[151,250,430,453]" pageId="12" pageNumber="115" type="paratype">Paratype</typeStatus>
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</emphasis>
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;
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<collectionCode id="ED37C2A9FFE7FFBE650F6892FED4B8A4" box="[272,358,430,453]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="12" pageNumber="115" type="Museum">MHNG</collectionCode>
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;
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<specimenCount id="9D2091E5FFE7FFBE65686892FE1DB8A4" box="[375,431,430,453]" pageId="12" pageNumber="115" type="male">male</specimenCount>
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; with the same data as the holotype but
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<date id="FF987CACFFE7FFBE653B68F1FEE9B885" box="[292,347,461,484]" pageId="12" pageNumber="115">5.IX.</date>
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and
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<specimenCode id="DB80F217FFE7FFBE659068F1FE42B885" box="[399,496,461,484]" pageId="12" pageNumber="115">F911046</specimenCode>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C09E7FFE7FFBE648868C0FE91BB53" pageId="12" pageNumber="115" type="etymology">
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<paragraph id="8B995A6CFFE7FFBE648868C0FE91BB53" blockId="12.[151,756,508,562]" pageId="12" pageNumber="115">
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<emphasis id="B952867EFFE7FFBE648868C0FEAEBB72" bold="true" box="[151,284,508,531]" pageId="12" pageNumber="115">Etymology:</emphasis>
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The species epithet is a Latin adjective meaning flat.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C09E7FFE7FFBE64886B76FF5EBAFD" pageId="12" pageNumber="115" type="diagnosis">
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<paragraph id="8B995A6CFFE7FFBE64886B76FF5EBAFD" blockId="12.[151,756,586,924]" pageId="12" pageNumber="115">
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<emphasis id="B952867EFFE7FFBE64886B76FEA2BB00" bold="true" box="[151,272,586,609]" pageId="12" pageNumber="115">Diagnosis:</emphasis>
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Body-length
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<quantity id="4CDEF789FFE7FFBE65A66B76FD91BB00" box="[441,547,586,609]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="115" unit="mm" value="0.9">0.90 mm</quantity>
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long, body light brown, not microsculptured and not iridescent; antennomere VIII exceeding half-length of antennomere VII, antennomere XI slightly longer than antennomere X and nearly 3 times as long as wide; elytra lacking parasutural stria; hypomeron lacking longitudinal striae; aedeagus as
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<figureCitation id="131D46E9FFE7FFBE65396A3BFE12BA7F" box="[294,416,775,798]" captionStart="Figs 23-29" captionStartId="13.[151,192,1925,1946]" captionTargetBox="[177,1384,257,1874]" captionTargetId="figure-4@13.[157,1440,254,1894]" captionTargetPageId="13" captionText="Figs 23-29. Scaphisoma donati sp. nov. (23) Parameres in ventral view, scale bar = 0.2 mm. Figs 24-27. Scaphobaeocera plana sp. nov. (24, 25) Aedeagus in dorsal and lateral views, scale = 0.1 mm. (26) Paramere in ventral view, scale = 0.05 mm. (27) Apical part of median lobe, in lateral view, scale = 0.05 mm. Figs 28, 29. Scaphoxium lobatum sp. nov. (28) Aedeagus in ventral view, scale = 0.1 mm. (29) Apex of median lobe with extruded internal sac and right paramere, in dorsal view, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6461034" httpUri="https://zenodo.org/record/6461034/files/figure.png" pageId="12" pageNumber="115">Figs 24-27</figureCitation>
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, with broad basal bulb, short, strongly sclerotized and bent apical process and short, bilobed apicodorsal plate, parameres expanded, notched mesally, internal sac with short, robust, incurved sclerite.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C09E7FFE7FFBE64886A8FFDF8B169" pageId="12" pageNumber="115" type="description">
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<paragraph id="8B995A6CFFE7FFBE64886A8FFE42BFDC" blockId="12.[151,757,947,1977]" pageId="12" pageNumber="115">
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<emphasis id="B952867EFFE7FFBE64886A8FFE95BAAB" bold="true" box="[151,295,947,970]" pageId="12" pageNumber="115">Description:</emphasis>
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Length
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<quantity id="4CDEF789FFE7FFBE65806A8FFDA3BAAB" box="[415,529,947,970]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="12" pageNumber="115" unit="mm" value="0.9">0.90 mm</quantity>
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, width
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<quantity id="4CDEF789FFE7FFBE66616A8FFD5DBAAB" box="[638,751,947,970]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="12" pageNumber="115" unit="mm" value="0.52">0.52 mm</quantity>
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, dorsoventral diameter
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<quantity id="4CDEF789FFE7FFBE65976AEFFE58BA8B" box="[392,490,979,1002]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="12" pageNumber="115" unit="mm" value="0.47">0.47 mm</quantity>
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. Head, body, femora and tibiae nearly uniformly light brown, elytra somewhat lighter near apices, tarsi and antennae lighter than tibiae. Length/width ratios of antennomeres as: III 10/5: IV 13/5: V 14/5: VI 12/5: VII 23/10: VIII 13/8: IX 24/10: X 27/11: XI 33/12. Pronotum finely punctate, not microsculptured. Scutellum concealed. Elytra not iridescent, not microsculptured, lacking parasutural striae, sutural striae deep, curved near base, discal punctation coarser than pronotal punctation. Hypomera not microsculptured, smooth, lacking longitudinal stria. Mesoventrite with low mesal ridge. Metaventrite weakly convex between mesocoxae, lacking median stria; entire median area rather coarsely and densely punctate, with some punctures about as large as puncture intervals, pubescence short. Lateral parts of metaventrite very finely punctate, not microsculptured. Submesocoxal areas about
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<quantity id="4CDEF789FFE7FFBE65DF6CD6FD91BF60" box="[448,547,1514,1537]" metricMagnitude="-5" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="115" unit="mm" value="0.03">0.03 mm</quantity>
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, as half of shortest interval to metacoxae, submesocoxal lines appearing impunctate. Metanepisterna about
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<quantity id="4CDEF789FFE7FFBE66086F15FDC9BF21" box="[535,635,1577,1600]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="115" unit="mm" value="0.05">0.05 mm</quantity>
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wide, flat, not narrowed anteriad, impunctate, with straight suture reaching metepimera. Tibiae straight. Abdomen not microsculptured, very finely punctate, coarse basal punctures of ventrite I excepted.
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</paragraph>
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<paragraph id="8B995A6CFFE7FFBE64886FF9FD2CBED8" blockId="12.[151,757,947,1977]" pageId="12" pageNumber="115">
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<emphasis id="B952867EFFE7FFBE64886FF9FEF7BFBC" box="[151,325,1733,1757]" italics="true" pageId="12" pageNumber="115">Male characters</emphasis>
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: Protarsomeres I to III strongly widened, about as wide as apices of protibiae. Aedeagus as in
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<figureCitation id="131D46E9FFE7FFBE66D96FD9FF6ABE7D" captionStart="Figs 23-29" captionStartId="13.[151,192,1925,1946]" captionTargetBox="[177,1384,257,1874]" captionTargetId="figure-4@13.[157,1440,254,1894]" captionTargetPageId="13" captionText="Figs 23-29. Scaphisoma donati sp. nov. (23) Parameres in ventral view, scale bar = 0.2 mm. Figs 24-27. Scaphobaeocera plana sp. nov. (24, 25) Aedeagus in dorsal and lateral views, scale = 0.1 mm. (26) Paramere in ventral view, scale = 0.05 mm. (27) Apical part of median lobe, in lateral view, scale = 0.05 mm. Figs 28, 29. Scaphoxium lobatum sp. nov. (28) Aedeagus in ventral view, scale = 0.1 mm. (29) Apex of median lobe with extruded internal sac and right paramere, in dorsal view, scale = 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6461034" httpUri="https://zenodo.org/record/6461034/files/figure.png" pageId="12" pageNumber="115">Figs 24-27</figureCitation>
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,
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<quantity id="4CDEF789FFE7FFBE64F66E39FE36BE7D" box="[233,388,1797,1820]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.5999999999999996" pageId="12" pageNumber="115" unit="mm" value="0.37" valueMax="0.38" valueMin="0.36">0.36-0.38 mm</quantity>
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long. Apical process of median lobe short, curved and strongly inflexed, in apical section nearly perpendicular to basal bulb. Articular process not prominent. Dorsal side of basal bulb produced to form bilobed, strongly sclerotized plate. Internal sac with strongly sclerotized, proximally hook-like piece.
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</paragraph>
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<paragraph id="8B995A6CFFE7FFBE64886EEDFDF8B169" blockId="12.[151,756,2001,2056]" pageId="12" pageNumber="115">
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<emphasis id="B952867EFFE7FFBE64886EEDFF4ABE89" bold="true" box="[151,248,2001,2024]" pageId="12" pageNumber="115">Habitat:</emphasis>
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Secondary forests on limestone,
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<quantity id="4CDEF789FFE7FFBE666B6EEDFD5DBE89" box="[628,751,2001,2024]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="12" pageNumber="115" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200-400 m</quantity>
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, dead log and leaf litter infested by fungi.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C09E7FFE7FFBE6720683EFACCB878" box="[831,1406,258,281]" pageId="12" pageNumber="115" type="distribution">
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<paragraph id="8B995A6CFFE7FFBE6720683EFACCB878" blockId="12.[831,1406,258,281]" box="[831,1406,258,281]" pageId="12" pageNumber="115">
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<emphasis id="B952867EFFE7FFBE6720683EFC66B878" bold="true" box="[831,980,258,281]" pageId="12" pageNumber="115">Distribution:</emphasis>
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<collectingCountry id="F3311AFCFFE7FFBE67C5683EFBF4B878" box="[986,1094,258,281]" name="Indonesia" pageId="12" pageNumber="115">Indonesia</collectingCountry>
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:
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<collectingCountry id="F3311AFCFFE7FFBE604F683EFB08B878" box="[1104,1210,258,281]" name="Indonesia" pageId="12" pageNumber="115">Moluccas</collectingCountry>
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, Kai Besar Island.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33C09E7FFE7FFBE6720680DFC04BBA9" pageId="12" pageNumber="115" type="discussion">
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<paragraph id="8B995A6CFFE7FFBE6720680DFC04BBA9" blockId="12.[831,1437,305,712]" pageId="12" pageNumber="115">
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<emphasis id="B952867EFFE7FFBE6720680DFC03B829" bold="true" box="[831,945,305,328]" pageId="12" pageNumber="115">Remarks:</emphasis>
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The species differs from most congeners by its body comparatively weakly convex, lacking microsculpture, and the elytra lacking parasutural striae, in combination. The aedeagal characters of
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<taxonomicName id="4C2621EFFFE7FFBE610568ADFAC6B8C9" authorityName="Löbl" authorityYear="2022" box="[1306,1396,400,424]" class="Insecta" family="Staphylinidae" genus="Scaphobaeocera" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="115" phylum="Arthropoda" rank="species" species="plana">
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<emphasis id="B952867EFFE7FFBE610568ADFAC6B8C9" box="[1306,1396,400,424]" italics="true" pageId="12" pageNumber="115">S. plana</emphasis>
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</taxonomicName>
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are unique. The presence of a narrow apicodorsal process of the median lobe, the shape of the parameres and that of the sclerite of the internal sac lacking a flagellum are diagnostic. The generic assigned is therefore tentative. Several character states used to define the
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<taxonomicName id="4C2621EFFFE7FFBE67206B6DFC54BB09" box="[831,998,593,616]" pageId="12" pageNumber="103" rank="tribe" tribe="Scaphisomatini">Scaphisomatini</taxonomicName>
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genera (see
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<bibRefCitation id="EFB7279DFFE7FFBE60986B6DFA26BB09" author="Leschen R. A. B. & Lobl I." box="[1159,1428,593,616]" pageId="12" pageNumber="115" pagination="1 - 63" refId="ref11061" refString="Leschen R. A. B. & Lobl I. 2005. Phylogeny and classification of Scaphisomatini Staphylinidae: Scaphidiinae with notes on mycophagy, termitophily, and functional morphology. Coleopterists Society Monographs 3: 1 - 63." type="journal article" year="2005">Leschen & Löbl, 2005</bibRefCitation>
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) have not been verified as they may be seen only on slides of the head and thorax, requiring more available specimens.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |