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<mods:title id="97D09447AC31A90B02B1F90271BC72A4">New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe</mods:title>
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E-H; 27G, H)
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sp.
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Merle
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2011: 424
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, pl. 90, fig.7a, b.
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. —
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(
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; Early Miocene (
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.F.J09945 (
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coll.), figured specimen in
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<emphasis id="B953EADEFFC7E427FF3CF91CFF32F952" box="[132,179,1737,1761]" italics="true" pageId="33" pageNumber="526">et al.</emphasis>
(2011: 424
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, pl. 90, fig. 7), H:
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<figureCitation id="131C2A49FFC7E427FD8EF91FFD29F953" box="[566,680,1737,1761]" captionStart="FIG" captionStartId="32.[132,143,1770,1787]" captionTargetBox="[138,1450,251,1691]" captionTargetId="figure-254@32.[699,1263,826,1403]" captionTargetPageId="32" captionText="FIG. 18. — Spiral cords and internal denticles of the outer lip in Pterynotus (Pterymarchia) gaasensis n. sp. (A-D), P. (P.) pelouatensis n. sp. (E-H) and P. (P) sandbergeri (HÖrnes,1856) (I-L): A, B, holotype,MNHN.F.A32492 (Merle coll.), Gaas (Espibos),Landes,Aquitaine Basin,France,early Oligocene (Rupelian); C, D, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos);E, F, holotype, MNHN.F.J09945, Saucats (Peloua), Gironde, Aquitaine Basin, Early Miocene (Burdigalian); G, H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, lectotype designated herein,NHMW1855/0045/0435,Gainfarn,Baden, Austria,Middle Miocene (Langhian); K, L, Landau coll., Bad VÖslau, Baden. Scale bars: A-H, 5 mm; I-L, 10 mm. Credits:A-H, P. Loubry (MNHN/CNRS); I, J, M. Harzhauser (NHMW); K, L, B. Landau." figureDoi="http://doi.org/10.5281/zenodo.13985802" httpUri="https://zenodo.org/record/13985802/files/figure.png" pageId="33" pageNumber="526">Figs 18E, F</figureCitation>
;
<figureCitation id="131C2A49FFC7E427FD0AF91FFD63F952" box="[690,738,1737,1761]" captionStart="FIG" captionStartId="46.[132,143,1606,1623]" captionTargetBox="[132,1455,215,1546]" captionTargetId="figure-366@46.[699,1260,794,1359]" captionTargetPageId="46" captionText="FIG. 27. — New species of Timbellus De Gregorio, 1885, Pterynotus Swainson, 1833 (Pterymarchia Houart, 1995), Textiliomurex Merle, 2011, Ponderia Houart, 1986 and Flexopteron Shuto, 1969: A-C, Timbellus submicropterus n. sp.: A, holotype, MNHN.F.A24111 (MNHN coll.), Fresville, France, Lutetian; B, paratype, MNHN.F.A24126 (Munier-Chalmas coll.), Gourbesville, France, Lutetian; C, paratype, UCBL-EM 33438 (Bourdot coll.), Fresville; D, T. cf. submicropterus n. sp., MNHN.F.A90557 (Brébion coll.), Saint-Aignan-Grand-Lieu (Pierre-Aiguë), France, Bartonian; E, F, Pterynotus (Pterymarchia) gaasensis n. sp.: E, holotype, MNHN.F.A32492 (Merle coll.), Gaas (Espibos), France, Rupelian; F, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos); G, H, P. (P.) pelouatensis n. sp.: G, holotype, MNHN.F.J09945, Saucats (Peloua), France, Burdigalian; H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, Textiliomurex chodmonensis n. sp.: I, holotype, MNHN.F.A81144 (Pacaud coll.), Chaumont-en-Vexin (Carrière Darcy), France, Lutetian; J, paratype, MNHN.F.A81145 (Pons coll.), Chaumont-en- Vexin (Carrière Darcy); K, L, Ponderia remyi n. sp.: K, holotype, MNHN.F.A90548 (Remy coll.), Fresville, Lutetian; L, paratype, MNHN.F.A90555, Fresville (Remy coll.); M, N, Flexopteron constantinense n. sp.: M, paratype, MNHN.F.A77761 (MNHN coll.), Carentan-les-Marais (Cauvin), France, Priabonian; N, holotype, MNHN.F.A77760 (MNHN coll.), Carentan-les-Marais (Cauvin). Scale bars: A-D, I-L, 10 mm; E-H, N, 5 mm; M, 3 mm. Credits: A, B, D-F, N, P. Loubry (MNHN/ CNRS); C, E. Robert (UCBL)." figureDoi="http://doi.org/10.5281/zenodo.13985831" httpUri="https://zenodo.org/record/13985831/files/figure.png" pageId="33" pageNumber="526">27G</figureCitation>
).
</materialsCitation>
<materialsCitation id="3B4F3C91FFC7E427FF3CF930FE69F8AB" ID-GBIF-Occurrence="4958671392" collectionCode="MNHN" collectorName="Neuville" country="France" location="France" pageId="33" pageNumber="526" specimenCount="1" typeStatus="paratype">
<emphasis id="B953EADEFFC7E427FF3CF930FEA0F94D" bold="true" box="[132,289,1766,1790]" pageId="33" pageNumber="526">
<typeStatus id="549C886EFFC7E427FF3CF930FF5BF94D" box="[132,218,1766,1790]" pageId="33" pageNumber="526" type="paratype">Paratype</typeStatus>
.
<collectingCountry id="F330765CFFC7E427FF5AF930FEA0F94D" box="[226,289,1766,1790]" name="France" pageId="33" pageNumber="526">France</collectingCountry>
</emphasis>
• 1 spm; same as for the holotype;
<collectionCode id="ED36AE09FFC7E427FDE1F930FD2AF94E" box="[601,683,1766,1789]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="526">MNHN</collectionCode>
.F.J17632 (
<collectorName id="26D2531AFFC7E427FF33F8D6FF5EF8AB" box="[139,223,1792,1816]" pageId="33" pageNumber="526">Neuville</collectorName>
coll.) (
<figureCitation id="131C2A49FFC7E427FE9DF8D6FE20F8A4" box="[293,417,1792,1816]" captionStart="FIG" captionStartId="32.[132,143,1770,1787]" captionTargetBox="[138,1450,251,1691]" captionTargetId="figure-254@32.[699,1263,826,1403]" captionTargetPageId="32" captionText="FIG. 18. — Spiral cords and internal denticles of the outer lip in Pterynotus (Pterymarchia) gaasensis n. sp. (A-D), P. (P.) pelouatensis n. sp. (E-H) and P. (P) sandbergeri (HÖrnes,1856) (I-L): A, B, holotype,MNHN.F.A32492 (Merle coll.), Gaas (Espibos),Landes,Aquitaine Basin,France,early Oligocene (Rupelian); C, D, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos);E, F, holotype, MNHN.F.J09945, Saucats (Peloua), Gironde, Aquitaine Basin, Early Miocene (Burdigalian); G, H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, lectotype designated herein,NHMW1855/0045/0435,Gainfarn,Baden, Austria,Middle Miocene (Langhian); K, L, Landau coll., Bad VÖslau, Baden. Scale bars: A-H, 5 mm; I-L, 10 mm. Credits:A-H, P. Loubry (MNHN/CNRS); I, J, M. Harzhauser (NHMW); K, L, B. Landau." figureDoi="http://doi.org/10.5281/zenodo.13985802" httpUri="https://zenodo.org/record/13985802/files/figure.png" pageId="33" pageNumber="526">Figs 18G, H</figureCitation>
;
<figureCitation id="131C2A49FFC7E427FE14F8D6FE5DF8AB" box="[428,476,1792,1816]" captionStart="FIG" captionStartId="46.[132,143,1606,1623]" captionTargetBox="[132,1455,215,1546]" captionTargetId="figure-366@46.[699,1260,794,1359]" captionTargetPageId="46" captionText="FIG. 27. — New species of Timbellus De Gregorio, 1885, Pterynotus Swainson, 1833 (Pterymarchia Houart, 1995), Textiliomurex Merle, 2011, Ponderia Houart, 1986 and Flexopteron Shuto, 1969: A-C, Timbellus submicropterus n. sp.: A, holotype, MNHN.F.A24111 (MNHN coll.), Fresville, France, Lutetian; B, paratype, MNHN.F.A24126 (Munier-Chalmas coll.), Gourbesville, France, Lutetian; C, paratype, UCBL-EM 33438 (Bourdot coll.), Fresville; D, T. cf. submicropterus n. sp., MNHN.F.A90557 (Brébion coll.), Saint-Aignan-Grand-Lieu (Pierre-Aiguë), France, Bartonian; E, F, Pterynotus (Pterymarchia) gaasensis n. sp.: E, holotype, MNHN.F.A32492 (Merle coll.), Gaas (Espibos), France, Rupelian; F, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos); G, H, P. (P.) pelouatensis n. sp.: G, holotype, MNHN.F.J09945, Saucats (Peloua), France, Burdigalian; H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, Textiliomurex chodmonensis n. sp.: I, holotype, MNHN.F.A81144 (Pacaud coll.), Chaumont-en-Vexin (Carrière Darcy), France, Lutetian; J, paratype, MNHN.F.A81145 (Pons coll.), Chaumont-en- Vexin (Carrière Darcy); K, L, Ponderia remyi n. sp.: K, holotype, MNHN.F.A90548 (Remy coll.), Fresville, Lutetian; L, paratype, MNHN.F.A90555, Fresville (Remy coll.); M, N, Flexopteron constantinense n. sp.: M, paratype, MNHN.F.A77761 (MNHN coll.), Carentan-les-Marais (Cauvin), France, Priabonian; N, holotype, MNHN.F.A77760 (MNHN coll.), Carentan-les-Marais (Cauvin). Scale bars: A-D, I-L, 10 mm; E-H, N, 5 mm; M, 3 mm. Credits: A, B, D-F, N, P. Loubry (MNHN/ CNRS); C, E. Robert (UCBL)." figureDoi="http://doi.org/10.5281/zenodo.13985831" httpUri="https://zenodo.org/record/13985831/files/figure.png" pageId="33" pageNumber="526">27H</figureCitation>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFC7E427FF3CF8E3FC81F8FF" box="[132,768,1845,1869]" pageId="33" pageNumber="526" type="etymology">
<paragraph id="8B9836CCFFC7E427FF3CF8E3FC81F8FF" blockId="33.[132,768,1845,1869]" box="[132,768,1845,1869]" pageId="33" pageNumber="526">
<smallCapsWord id="8D7EA010FFC7E427FF3CF8E3FF7FF8F8" baselines="1862,1863" box="[132,254,1845,1868]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Etymology" pageId="33" pageNumber="526">ETYMOLOGY</smallCapsWord>
. — From the
<typeStatus id="549C886EFFC7E427FE33F8E3FE34F8FE" box="[395,437,1845,1869]" pageId="33" pageNumber="526">type</typeStatus>
locality, Peloua, Gironde,
<collectingCountry id="F330765CFFC7E427FD00F8E3FD7DF8FF" box="[696,764,1845,1868]" name="France" pageId="33" pageNumber="526">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFC7E427FF3CF8BCFD81F858" pageId="33" pageNumber="526" type="materials_examined">
<paragraph id="8B9836CCFFC7E427FF3CF8BCFE7BF82F" blockId="33.[132,775,1898,1948]" pageId="33" pageNumber="526">
<smallCapsWord id="8D7EA010FFC7E427FF3CF8BCFF34F8CC" baselines="1915,1915" box="[132,181,1898,1921]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="33" pageNumber="526">
<typeStatus id="549C886EFFC7E427FF3CF8BCFF34F8CC" box="[132,181,1898,1921]" pageId="33" pageNumber="526">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D7EA010FFC7E427FF05F8B9FE99F833" baselines="1916" box="[189,280,1903,1920]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="horizon" pageId="33" pageNumber="526">HORIZON</smallCapsWord>
. — Faluns de Saucats, Early Miocene (lower Burdigalian), see
<bibRefCitation id="EFB64B3DFFC7E427FEB1F852FE74F82F" author="PARIZE O. &amp; MULDER T. &amp; CAHUZAC B. &amp; FIET N. &amp; LONDEIX L. &amp; RUBINO J. - L." box="[265,501,1924,1948]" pageId="33" pageNumber="526" pagination="390 - 399" refId="ref48184" refString="PARIZE O., MULDER T., CAHUZAC B., FIET N. LONDEIX L. &amp; RUBINO J. - L. 2008. - Sedimentology and sequence stratigraphy of Aquitanian and Burdigalian stratotypes in the Bordeaux area (southwestern France). Comptes Rendus Geoscience 340 (6): 390 - 399. https: // doi. org / 10.1016 / j. crte. 2008.03.004" type="journal article" year="2008">
Parize
<emphasis id="B953EADEFFC7E427FEF1F853FEF9F82F" box="[329,376,1924,1948]" italics="true" pageId="33" pageNumber="526">et al.</emphasis>
(2008: 396)
</bibRefCitation>
.
</paragraph>
<paragraph id="8B9836CCFFC7E427FF3CF86FFD81F858" blockId="33.[132,774,1977,2027]" pageId="33" pageNumber="526">
<smallCapsWord id="8D7EA010FFC7E427FF3CF86FFF39F87C" baselines="1994,1995" box="[132,184,1977,2000]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Type" pageId="33" pageNumber="526">
<typeStatus id="549C886EFFC7E427FF3CF86FFF39F87C" box="[132,184,1977,2000]" pageId="33" pageNumber="526">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="8D7EA010FFC7E427FF78F868FE9EF87C" baselines="1995" box="[192,287,1982,1999]" lowerCaseFontSize="7" mainFontSize="10" normCase="lower" normString="locality" pageId="33" pageNumber="526">LOCALITY</smallCapsWord>
. —
<collectingCountry id="F330765CFFC7E427FEE9F86FFE19F863" box="[337,408,1977,2000]" name="France" pageId="33" pageNumber="526">France</collectingCountry>
,
<collectingRegion id="49E3F82EFFC7E427FE1CF86FFD89F862" box="[420,520,1977,2001]" country="France" name="Aquitaine" pageId="33" pageNumber="526">Aquitaine</collectingRegion>
Basin, Gironde, Saucats (Peloua), seeParize
<emphasis id="B953EADEFFC7E427FEFCF802FEF2F858" box="[324,371,2003,2027]" italics="true" pageId="33" pageNumber="526">et al.</emphasis>
(2008: fig. 1).
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFC7E427FC95FF19FAC0FF55" box="[813,1345,206,231]" pageId="33" pageNumber="526" type="distribution">
<paragraph id="8B9836CCFFC7E427FC95FF19FAC0FF55" blockId="33.[813,1345,206,231]" box="[813,1345,206,231]" pageId="33" pageNumber="526">
<smallCapsWord id="8D7EA010FFC7E427FC95FF19FC3BFF56" baselines="224,225" box="[813,954,207,230]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Distribution" pageId="33" pageNumber="526">DISTRIBUTION</smallCapsWord>
. — Only known from the
<typeStatus id="549C886EFFC7E427FB7CFF19FB6FFF54" box="[1220,1262,207,231]" pageId="33" pageNumber="526">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C33D6547FFC7E424FC95FECEFDECFE02" lastPageId="34" lastPageNumber="527" pageId="33" pageNumber="526" type="description">
<paragraph id="8B9836CCFFC7E427FC95FECEFC3CFE83" blockId="33.[811,1458,280,1230]" box="[813,957,280,306]" pageId="33" pageNumber="526">
<smallCapsWord id="8D7EA010FFC7E427FC95FECEFC3CFE83" baselines="299,299" box="[813,957,280,306]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Description" pageId="33" pageNumber="526">DESCRIPTION</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFC7E427FC95FEE1FB10FB7D" blockId="33.[811,1458,280,1230]" pageId="33" pageNumber="526">
Two smooth protoconch whorls preserved delimited by sinusigera scar (protoconch tip not preserved) (see
<bibRefCitation id="EFB64B3DFFC7E427FAC8FE81FC60FE22" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." pageId="33" pageNumber="526" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
<taxonomicName id="4C274D4FFFC7E427FAC8FE81FA30FEC2" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[1392,1457,343,369]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
<emphasis id="B953EADEFFC7E427FC95FEA1FCE2FE22" box="[813,867,375,401]" italics="true" pageId="33" pageNumber="526">et al.</emphasis>
2011: 122
</bibRefCitation>
, text-fig. 46E). Teleoconch up to
<quantity id="4CDF9B29FFC7E427FAE5FEA1FA2EFE22" box="[1373,1455,375,401]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="33" pageNumber="526" unit="mm" value="21.0">21 mm</quantity>
in height, up to
<quantity id="4CDF9B29FFC7E427FC66FE40FBC3FE02" box="[990,1090,406,433]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="33" pageNumber="526" unit="mm" value="12.5">12.5 mm</quantity>
in width, biconic in profile, composed of five whorls. Moderately high spire with subcarinate whorls. Last whorl up to 73% of total length. Apical angle 64°. Spiral sculpture with marked primary cords. First whorl: appearance of P1 and P2; second whorl: appearance of IP; third whorl: appearance of abis, adis and s2. Fourth whorl: no change or appearance of threads. Fifth whorl: IP, adis and abis, P1 to P6, s1 to s6; s3 and s4 more developed than other secondary cords; P6 slightly atrophied, on base of convex part of whorl; ADP, MP, ads and ms; ABP missing. Surface of shell with numerous threads appearing on fourth whorl. Axial sculpture with thick varices. On first whorl: 10-11 lamellose varices; on second whorl: 10-12 varices; on third whorl: 11-12 varices; on fourth whorl: 10-12 varices; on fifth whorl 8-10 varices. From second whorl to fifth whorl: thick varices. No evidence of intervarices on last whorl. From third to last whorl numerous, fine microlamellae forming scabrous microsculpture covering entire surface. Ovate aperture up to 31% of diameter and up to 72% of length of last whorl (including siphonal canal). Columellar lip adherent posteriorly, becoming progressively more erect anteriorly. One or two weak columellar tubercles. Parietal lip adherent. Outer lip with strong internal denticles: ID, D2 to D6 present, D1 missing; D4 split, D6 poorly developed. Pseudoumbilicus moderately large. Siphonal canal open, up to 43% of aperture length, slightly dorsally recurved.
</paragraph>
<paragraph id="8B9836CCFFC7E427FC95FB22FC46FABF" blockId="33.[811,1457,1268,2028]" box="[813,967,1268,1294]" pageId="33" pageNumber="526">
<smallCapsWord id="8D7EA010FFC7E427FC95FB22FC46FABF" baselines="1287,1287" box="[813,967,1268,1294]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Comparisons" pageId="33" pageNumber="526">COMPARISONS</smallCapsWord>
</paragraph>
<paragraph id="8B9836CCFFC7E424FC95FAC5FDECFE02" blockId="33.[811,1457,1268,2028]" lastBlockId="34.[132,775,215,433]" lastPageId="34" lastPageNumber="527" pageId="33" pageNumber="526">
The European Neogene contains two species of
<taxonomicName id="4C274D4FFFC7E427FAFCFAC2FC4BFAFE" authorityName="Houart" authorityYear="1995" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="subGenus" subGenus="Pterymarchia">
<emphasis id="B953EADEFFC7E427FAFCFAC2FA30FA9E" box="[1348,1457,1300,1325]" italics="true" pageId="33" pageNumber="526">Pterynotus</emphasis>
(
<emphasis id="B953EADEFFC7E427FC8EFAE5FC41FAFE" box="[822,960,1331,1357]" italics="true" pageId="33" pageNumber="526">Pterymarchia</emphasis>
)
</taxonomicName>
:
<taxonomicName id="4C274D4FFFC7E427FC6CFAE2FB0AFAFE" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[980,1163,1331,1357]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="pelouatensis" status="n. sp." subGenus="Pterymarchia">
<emphasis id="B953EADEFFC7E427FC6CFAE2FC66FAFE" box="[980,999,1332,1357]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
(
<emphasis id="B953EADEFFC7E427FC4CFAE2FB87FAFE" box="[1012,1030,1332,1357]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
)
<emphasis id="B953EADEFFC7E427FBABFAE5FB0AFAFE" box="[1043,1163,1331,1357]" italics="true" pageId="33" pageNumber="526">pelouatensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC7E427FB29FAE2FB4BFAFD" box="[1169,1226,1332,1358]" pageId="33" pageNumber="526" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C274D4FFFC7E427FB42FAE2FC56FADD" authority="(Hornes, 1856)" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Hornes" baseAuthorityYear="1856" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="sandbergeri" status="n. comb." subGenus="Pterymurex">
<emphasis id="B953EADEFFC7E427FB42FAE2FA90FAFE" box="[1274,1297,1332,1357]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
(
<emphasis id="B953EADEFFC7E427FAA3FAE2FAAFFAFE" box="[1307,1326,1332,1357]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
)
<emphasis id="B953EADEFFC7E427FA83FAE5FA31FAFE" box="[1339,1456,1331,1357]" italics="true" pageId="33" pageNumber="526">sandbergeri</emphasis>
(Hörnes, 1856)
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC7E427FC67FA82FBC3FADE" box="[991,1090,1363,1390]" pageId="33" pageNumber="526" rank="species">n. comb.</taxonomicNameLabel>
from the Langhian of Paratethys (
<collectingCountry id="F330765CFFC7E427FC8EFAA5FC08FA3E" box="[822,905,1395,1421]" name="Austria" pageId="33" pageNumber="526">Austria</collectingCountry>
,
<collectingCountry id="F330765CFFC7E427FC2DFAA2FC76FA3D" box="[917,1015,1396,1422]" name="Hungary" pageId="33" pageNumber="526">Hungary</collectingCountry>
,
<collectingCountry id="F330765CFFC7E427FBBDFAA5FBEAFA3E" box="[1029,1131,1395,1421]" name="Romania" pageId="33" pageNumber="526">Romania</collectingCountry>
,
<collectingCountry id="F330765CFFC7E427FBC0FAA5FB44FA3E" box="[1144,1221,1395,1421]" name="Poland" pageId="33" pageNumber="526">Poland</collectingCountry>
) and the Serravallian of
<collectingCountry id="F330765CFFC7E427FCF3FA45FC19FA1E" box="[843,920,1427,1453]" name="Turkey" pageId="33" pageNumber="526">Turkey</collectingCountry>
. The type material of
<taxonomicName id="4C274D4FFFC7E427FB30FA42FAC0FA1E" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" baseAuthorityName="Hornes" baseAuthorityYear="1856" box="[1160,1345,1427,1453]" class="Gastropoda" family="Muricidae" genus="Murex" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="sandbergeri" subGenus="Pterymurex">
<emphasis id="B953EADEFFC7E427FB30FA42FB1EFA1E" box="[1160,1183,1428,1453]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
(
<emphasis id="B953EADEFFC7E427FB12FA42FB3CFA1E" box="[1194,1213,1428,1453]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
)
<emphasis id="B953EADEFFC7E427FB74FA45FAC0FA1E" box="[1228,1345,1427,1453]" italics="true" pageId="33" pageNumber="526">sandbergeri</emphasis>
</taxonomicName>
is housed in Naturhistorisches Museum of
<collectingRegion id="49E3F82EFFC7E427FB29FA65FB4CFA7E" box="[1169,1229,1459,1485]" country="Austria" name="Wien" pageId="33" pageNumber="526">Wien</collectingRegion>
and contains the figured
<typeStatus id="549C886EFFC7E427FCDDFA05FC36FA5E" box="[869,951,1491,1517]" pageId="33" pageNumber="526" type="syntype">syntype</typeStatus>
of the species (Hörnes 1856: pl. 51, fig. 5a, b, NHMW 1855/0045/0435). Here, we designate this specimen as the
<typeStatus id="549C886EFFC7E427FC10F9C4FB8BF99F" box="[936,1034,1554,1580]" pageId="33" pageNumber="526" type="lectotype">lectotype</typeStatus>
(
<figureCitation id="131C2A49FFC7E427FBA1F9C4FB0BF99E" box="[1049,1162,1554,1581]" captionStart="FIG" captionStartId="32.[132,143,1770,1787]" captionTargetBox="[138,1450,251,1691]" captionTargetId="figure-254@32.[699,1263,826,1403]" captionTargetPageId="32" captionText="FIG. 18. — Spiral cords and internal denticles of the outer lip in Pterynotus (Pterymarchia) gaasensis n. sp. (A-D), P. (P.) pelouatensis n. sp. (E-H) and P. (P) sandbergeri (HÖrnes,1856) (I-L): A, B, holotype,MNHN.F.A32492 (Merle coll.), Gaas (Espibos),Landes,Aquitaine Basin,France,early Oligocene (Rupelian); C, D, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos);E, F, holotype, MNHN.F.J09945, Saucats (Peloua), Gironde, Aquitaine Basin, Early Miocene (Burdigalian); G, H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, lectotype designated herein,NHMW1855/0045/0435,Gainfarn,Baden, Austria,Middle Miocene (Langhian); K, L, Landau coll., Bad VÖslau, Baden. Scale bars: A-H, 5 mm; I-L, 10 mm. Credits:A-H, P. Loubry (MNHN/CNRS); I, J, M. Harzhauser (NHMW); K, L, B. Landau." figureDoi="http://doi.org/10.5281/zenodo.13985802" httpUri="https://zenodo.org/record/13985802/files/figure.png" pageId="33" pageNumber="526">Fig. 18I, J</figureCitation>
).
<taxonomicName id="4C274D4FFFC7E427FB25F9C5FA2EF99F" box="[1181,1455,1554,1581]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="sandbergeri" subGenus="Pterymurex">
<emphasis id="B953EADEFFC7E427FB25F9C5FA88F99F" box="[1181,1289,1555,1580]" italics="true" pageId="33" pageNumber="526">Pterynotus</emphasis>
(
<emphasis id="B953EADEFFC7E427FAA1F9C5FAAAF99F" box="[1305,1323,1555,1580]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
)
<emphasis id="B953EADEFFC7E427FA82F9C4FA2EF99F" box="[1338,1455,1554,1580]" italics="true" pageId="33" pageNumber="526">sandbergeri</emphasis>
</taxonomicName>
was often attributed to
<taxonomicName id="4C274D4FFFC7E427FB9BF9E4FA96F9FE" authority="Iredale, 1915" authorityName="Iredale" authorityYear="1915" box="[1059,1303,1586,1613]" class="Gastropoda" family="Muricidae" genus="Murexsul" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC7E427FB9BF9E4FB02F9FF" box="[1059,1155,1586,1612]" italics="true" pageId="33" pageNumber="526">Murexsul</emphasis>
Iredale, 1915
</taxonomicName>
(
<bibRefCitation id="EFB64B3DFFC7E427FA9EF9E4FCE4F9DF" author="LANDAU B. M. &amp; HARZHAUSER M. &amp; ISLAMOGLU Y. &amp; MARQUES DA SILVA C." pageId="33" pageNumber="526" pagination="3 - 584" refId="ref45777" refString="LANDAU B. M., HARZHAUSER M., ISLAMOGLU Y. &amp; MARQUES DA SILVA C. 2013. - Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin, Turkey. Cainozoic Research 11 - 13: 3 - 584." type="book chapter" year="2013">
Landau
<emphasis id="B953EADEFFC7E427FAC5F9E5FA31F9FF" box="[1405,1456,1586,1612]" italics="true" pageId="33" pageNumber="526">et al.</emphasis>
2013
</bibRefCitation>
;
<bibRefCitation id="EFB64B3DFFC7E427FCC9F984FBB1F9DF" author="KOVACS Z. &amp; HIRMETZL T. &amp; VICIAN Z." box="[881,1072,1618,1644]" pageId="33" pageNumber="526" pagination="110 - 133" refId="ref45529" refString="KOVACS Z., HIRMETZL T. &amp; VICIAN Z. 2018. - Miocene Muricidae (Neogastropoda) assemblage from Letkes (Hungary). Bolletino malacologico 54 (2): 110 - 133." type="journal article" year="2018">
Kovács
<emphasis id="B953EADEFFC7E427FC79F985FC72F9DF" box="[961,1011,1618,1644]" italics="true" pageId="33" pageNumber="526">et al.</emphasis>
2018
</bibRefCitation>
; Kovács 2019). Although they share several similarities (biconic shells, internal denticles, columellar denticles), members of
<taxonomicName id="4C274D4FFFC7E427FB9FF944FB06F91F" authorityName="Iredale" authorityYear="1915" box="[1063,1159,1682,1708]" class="Gastropoda" family="Muricidae" genus="Murexsul" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC7E427FB9FF944FB06F91F" box="[1063,1159,1682,1708]" italics="true" pageId="33" pageNumber="526">Murexsul</emphasis>
</taxonomicName>
have a more rounded aperture, their internal denticles are close together, ID is weaker and their siphonal canal is narrower. The spires of
<taxonomicName id="4C274D4FFFC7E427FAFFF905FC08F8BF" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="pelouatensis" status="n. sp." subGenus="Pterymarchia">
<emphasis id="B953EADEFFC7E427FAFFF905FADAF95F" box="[1351,1371,1747,1772]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
(
<emphasis id="B953EADEFFC7E427FAD0F905FAFAF95F" box="[1384,1403,1747,1772]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
)
<emphasis id="B953EADEFFC7E427FA31F904FC08F8BF" italics="true" pageId="33" pageNumber="526">pelouatensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC7E427FC36F924FC46F8BF" box="[910,967,1778,1804]" pageId="33" pageNumber="526" rank="species">n. sp.</taxonomicNameLabel>
and the
<typeStatus id="549C886EFFC7E427FB99F924FB02F8BF" box="[1057,1155,1778,1804]" pageId="33" pageNumber="526" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C274D4FFFC7E427FB1AF925FAD9F8BF" box="[1186,1368,1778,1804]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="sandbergeri" subGenus="Pterymurex">
<emphasis id="B953EADEFFC7E427FB1AF925FB38F8BF" box="[1186,1209,1779,1804]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
(
<emphasis id="B953EADEFFC7E427FB7BF925FB57F8BF" box="[1219,1238,1779,1804]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
)
<emphasis id="B953EADEFFC7E427FB5BF924FAD9F8BF" box="[1251,1368,1778,1804]" italics="true" pageId="33" pageNumber="526">sandbergeri</emphasis>
</taxonomicName>
are very similar, but their last whorls are very different. The
<typeStatus id="549C886EFFC7E427FAF4F8C4FA2FF89F" box="[1356,1454,1810,1836]" pageId="33" pageNumber="526" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C274D4FFFC7E427FCF2F8E4FB85F8F8" box="[842,1028,1841,1868]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="sandbergeri" subGenus="Pterymurex">
<emphasis id="B953EADEFFC7E427FCF2F8E4FCE0F8F8" box="[842,865,1842,1867]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
(
<emphasis id="B953EADEFFC7E427FCD5F8E4FC01F8F8" box="[877,896,1842,1867]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
)
<emphasis id="B953EADEFFC7E427FC37F8E7FB85F8F8" box="[911,1028,1841,1867]" italics="true" pageId="33" pageNumber="526">sandbergeri</emphasis>
</taxonomicName>
has a larger aperture, a surface covered by numerous threads and atrophy of P2, P5 and P6; P4 is more developed and D6 is missing. The differences between the primary cords are great, but in other specimens attributed from
<taxonomicName id="4C274D4FFFC7E427FCD1F864FBA0F878" box="[873,1057,1969,1995]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="526" phylum="Mollusca" rank="species" species="sandbergeri" subGenus="Pterymurex">
<emphasis id="B953EADEFFC7E427FCD1F864FC01F878" box="[873,896,1970,1995]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
(
<emphasis id="B953EADEFFC7E427FC32F864FC1CF878" box="[906,925,1970,1995]" italics="true" pageId="33" pageNumber="526">P.</emphasis>
)
<emphasis id="B953EADEFFC7E427FC14F867FBA0F878" box="[940,1057,1969,1995]" italics="true" pageId="33" pageNumber="526">sandbergeri</emphasis>
</taxonomicName>
(
<figureCitation id="131C2A49FFC7E427FB89F867FB30F878" box="[1073,1201,1969,1996]" captionStart="FIG" captionStartId="32.[132,143,1770,1787]" captionTargetBox="[138,1450,251,1691]" captionTargetId="figure-254@32.[699,1263,826,1403]" captionTargetPageId="32" captionText="FIG. 18. — Spiral cords and internal denticles of the outer lip in Pterynotus (Pterymarchia) gaasensis n. sp. (A-D), P. (P.) pelouatensis n. sp. (E-H) and P. (P) sandbergeri (HÖrnes,1856) (I-L): A, B, holotype,MNHN.F.A32492 (Merle coll.), Gaas (Espibos),Landes,Aquitaine Basin,France,early Oligocene (Rupelian); C, D, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos);E, F, holotype, MNHN.F.J09945, Saucats (Peloua), Gironde, Aquitaine Basin, Early Miocene (Burdigalian); G, H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, lectotype designated herein,NHMW1855/0045/0435,Gainfarn,Baden, Austria,Middle Miocene (Langhian); K, L, Landau coll., Bad VÖslau, Baden. Scale bars: A-H, 5 mm; I-L, 10 mm. Credits:A-H, P. Loubry (MNHN/CNRS); I, J, M. Harzhauser (NHMW); K, L, B. Landau." figureDoi="http://doi.org/10.5281/zenodo.13985802" httpUri="https://zenodo.org/record/13985802/files/figure.png" pageId="33" pageNumber="526">Fig. 18K, L</figureCitation>
), the atrophy of P2, P5 and P6 is less strong than in the
<typeStatus id="549C886EFFC7E427FB36F807FB71F858" box="[1166,1264,2001,2027]" pageId="33" pageNumber="526" type="lectotype">lectotype</typeStatus>
from
<collectingCountry id="F330765CFFC7E427FA8BF807FA01F858" box="[1331,1408,2001,2027]" name="Austria" pageId="33" pageNumber="526">Austria</collectingCountry>
(see
<bibRefCitation id="EFB64B3DFFC4E424FF3CFF01FECDFF42" author="LANDAU B. M. &amp; HARZHAUSER M. &amp; ISLAMOGLU Y. &amp; MARQUES DA SILVA C." box="[132,332,215,241]" pageId="34" pageNumber="527" pagination="3 - 584" refId="ref45777" refString="LANDAU B. M., HARZHAUSER M., ISLAMOGLU Y. &amp; MARQUES DA SILVA C. 2013. - Systematics and palaeobiogeography of the gastropods of the middle Miocene (Serravallian) Karaman Basin, Turkey. Cainozoic Research 11 - 13: 3 - 584." type="book chapter" year="2013">
Landau
<emphasis id="B953EADEFFC4E424FF64FF0EFE8FFF42" box="[220,270,215,241]" italics="true" pageId="34" pageNumber="527">et al.</emphasis>
2013
</bibRefCitation>
;
<bibRefCitation id="EFB64B3DFFC4E424FEE1FF01FD9BFF42" author="KOVACS Z. &amp; HIRMETZL T. &amp; VICIAN Z." box="[345,538,215,241]" pageId="34" pageNumber="527" pagination="110 - 133" refId="ref45529" refString="KOVACS Z., HIRMETZL T. &amp; VICIAN Z. 2018. - Miocene Muricidae (Neogastropoda) assemblage from Letkes (Hungary). Bolletino malacologico 54 (2): 110 - 133." type="journal article" year="2018">
Kovács
<emphasis id="B953EADEFFC4E424FE12FF0EFE5CFF42" box="[426,477,215,241]" italics="true" pageId="34" pageNumber="527">et al.</emphasis>
2018
</bibRefCitation>
; Kovács 2019). Compared to extant
<emphasis id="B953EADEFFC4E424FE8FFF2EFECEFEA2" box="[311,335,248,273]" italics="true" pageId="34" pageNumber="527">P.</emphasis>
(
<taxonomicName id="4C274D4FFFC4E424FEE1FF21FE64FEA2" authorityName="Houart" authorityYear="1995" box="[345,485,247,273]" class="Gastropoda" family="Muricidae" genus="Pterymarchia" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="genus">
<emphasis id="B953EADEFFC4E424FEE1FF21FE64FEA2" box="[345,485,247,273]" italics="true" pageId="34" pageNumber="527">Pterymarchia</emphasis>
</taxonomicName>
),
<taxonomicName id="4C274D4FFFC4E424FE47FF2EFD40FEA2" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[511,705,247,273]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="species" species="pelouatensis" status="n. sp." subGenus="Pterymarchia">
<emphasis id="B953EADEFFC4E424FE47FF2EFD97FEA2" box="[511,534,248,273]" italics="true" pageId="34" pageNumber="527">P.</emphasis>
(
<emphasis id="B953EADEFFC4E424FD9AFF2EFDB4FEA2" box="[546,565,248,273]" italics="true" pageId="34" pageNumber="527">P.</emphasis>
)
<emphasis id="B953EADEFFC4E424FDFEFF21FD40FEA2" box="[582,705,247,273]" italics="true" pageId="34" pageNumber="527">pelouatensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057A5FFC4E424FD72FF2EFC86FEA1" box="[714,775,248,274]" pageId="34" pageNumber="527" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C274D4FFFC4E424FF0CFECEFEEDFE82" box="[180,364,279,305]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="species" species="sandbergeri" subGenus="Pterymurex">
<emphasis id="B953EADEFFC4E424FF0CFECEFF4AFE82" box="[180,203,280,305]" italics="true" pageId="34" pageNumber="527">P.</emphasis>
(
<emphasis id="B953EADEFFC4E424FF6DFECEFF69FE82" box="[213,232,280,305]" italics="true" pageId="34" pageNumber="527">P.</emphasis>
)
<emphasis id="B953EADEFFC4E424FF4FFEC1FEEDFE82" box="[247,364,279,305]" italics="true" pageId="34" pageNumber="527">sandbergeri</emphasis>
</taxonomicName>
share greatest morphological similarities with
<taxonomicName id="4C274D4FFFC4E424FF5DFEEEFDD6FEE2" authority="(Roding, 1798)" baseAuthorityName="Roding" baseAuthorityYear="1798" box="[229,599,311,337]" class="Gastropoda" family="Muricidae" genus="Pterymarchia" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="species" species="martinetana" subGenus="Pterymurex">
<emphasis id="B953EADEFFC4E424FF5DFEEEFF7DFEE2" box="[229,252,312,337]" italics="true" pageId="34" pageNumber="527">P.</emphasis>
(
<emphasis id="B953EADEFFC4E424FEBFFEEEFE9BFEE2" box="[263,282,312,337]" italics="true" pageId="34" pageNumber="527">P.</emphasis>
)
<emphasis id="B953EADEFFC4E424FE90FEEEFE28FEE2" box="[296,425,312,337]" italics="true" pageId="34" pageNumber="527">martinetana</emphasis>
(Röding, 1798)
</taxonomicName>
and particularly with specimens from the Red Sea (
<bibRefCitation id="EFB64B3DFFC4E424FE43FE81FD32FEC2" author="MERLE D. &amp; GARRIGUES B. &amp; POINTIER J. - P." box="[507,691,343,369]" pageId="34" pageNumber="527" refId="ref47234" refString="MERLE D., GARRIGUES B. &amp; POINTIER J. - P. 2011. - Fossil &amp; Recent Muricidae of the World. Part Muricinae. ConchBooks, Hackenheim, 652 p." type="book" year="2011">
<taxonomicName id="4C274D4FFFC4E424FE43FE81FDBBFEC2" authorityName="Merle &amp; Pacaud &amp; Ledon &amp; Goret" authorityYear="2024" box="[507,570,343,369]" class="Gastropoda" family="Muricidae" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="543" phylum="Mollusca" rank="subFamily" status="subfam. nov." subFamily="Merle">Merle</taxonomicName>
<emphasis id="B953EADEFFC4E424FDF9FE8EFDF4FEC2" box="[577,629,343,369]" italics="true" pageId="34" pageNumber="527">et al.</emphasis>
2011
</bibRefCitation>
: pl. 92, figs 11, 12), but
<taxonomicName id="4C274D4FFFC4E424FE82FEA1FE73FE22" box="[314,498,375,401]" class="Gastropoda" family="Muricidae" genus="Pterynotus" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="527" phylum="Mollusca" rank="species" species="sandbergeri" subGenus="Pterymurex">
<emphasis id="B953EADEFFC4E424FE82FEA1FED0FE23" box="[314,337,375,400]" italics="true" pageId="34" pageNumber="527">P.</emphasis>
(
<emphasis id="B953EADEFFC4E424FEE3FEA1FEEFFE23" box="[347,366,375,400]" italics="true" pageId="34" pageNumber="527">P.</emphasis>
)
<emphasis id="B953EADEFFC4E424FEC5FEA1FE73FE22" box="[381,498,375,401]" italics="true" pageId="34" pageNumber="527">sandbergeri</emphasis>
</taxonomicName>
differs by having develop cord spines on P1, P2, P4, P5, ADP and MP.
</paragraph>
</subSubSection>
</treatment>
</document>