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<document id="78ABEF194431F65B898B9E9A37C359E6" ID-DOI="10.11646/zootaxa.2260.1.8" ID-ISSN="1175-5326" ID-Zenodo-Dep="5322208" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629645325754" checkinUser="felipe" docAuthor="Hughes, L. E. &amp; Lowry, J. K." docDate="2009" docId="038E87C89D30FFB2FF6ACEEA6EA7FBF0" docLanguage="en" docName="zt02260p219.pdf" docOrigin="Zootaxa 2260 (1)" docSource="https://www.biotaxa.org/Zootaxa/article/view/zootaxa.2260.1.8" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Cymadusa khbarnardi Hughes &amp; Lowry, sp. nov." docType="treatment" docVersion="4" lastPageNumber="191" masterDocId="FFB7FFB09D13FF94FFFDCF4C6D7CFFA2" masterDocTitle="Ampithoidae *" masterLastPageNumber="219" masterPageNumber="153" pageNumber="188" updateTime="1699088795311" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="DB6117AD65BFB0F4CEEB812E679141C2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1876CD5DA90510B7D8234242121E4E1B">
<mods:title id="32DD5E9929D44B65C783D5F0ABAEB26C">Ampithoidae *</mods:title>
</mods:titleInfo>
<mods:name id="08788C2589EBA38AFAB650231ED77AE2" type="personal">
<mods:role id="2FF595C0A38F1182C465396375C1688B">
<mods:roleTerm id="719FF399F2935BCA52AC398EAF7D9205">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1F97E76C346FBC1E857CED7371ADED6D">Hughes, L. E.</mods:namePart>
</mods:name>
<mods:name id="D957E63879348F4FF355B65353F20C71" type="personal">
<mods:role id="357839D48080B5C3FE0D54D0F7D7BFC0">
<mods:roleTerm id="640E1FECC6C93B0EA4832AC3FD440B4B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1B1B1A1E18999328B1691202289CD100">Lowry, J. K.</mods:namePart>
</mods:name>
<mods:typeOfResource id="05F1193EDA973C0F029AD016BD12DBFE">text</mods:typeOfResource>
<mods:relatedItem id="7F7FAC057672DAF3E796516022E78269" type="host">
<mods:titleInfo id="96D95FAA860FFC31011700356E8AE856">
<mods:title id="3C8A0F25678D75B69EEDBEEA42D70A7A">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FE6DF421DDFF0BA5548145E2D5F41B9B">
<mods:date id="E725EAE643F25F345FACE90995766BE8">2009</mods:date>
<mods:detail id="ED7780A69E1A603B88D7125203DD05B0" type="pubDate">
<mods:number id="DF0D12108DC071E7567A1131EE8B6727">2009-10-08</mods:number>
</mods:detail>
<mods:detail id="D973760FBBD0336DC973391808C84535" type="volume">
<mods:number id="33566F03D69465D45ADFAB6A4D770F6D">2260</mods:number>
</mods:detail>
<mods:detail id="2F4962F5A583B8279F569385468001B3" type="issue">
<mods:number id="F0A4A98B07BD5B93C6B70DBB7E2EEC3A">1</mods:number>
</mods:detail>
<mods:extent id="F24014B7EB02215F2AF76C64EBF27EC4" unit="page">
<mods:start id="AE1837656A4E8CAA438D10742342D068">153</mods:start>
<mods:end id="3C2553B04C66B26A9E78951968A41142">219</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="6DEFC85B0BFBFA20F2F3A06995FCCFBE">
<mods:url id="53A84B8B6C39D219FD594525E4BCFF7E">https://www.biotaxa.org/Zootaxa/article/view/zootaxa.2260.1.8</mods:url>
</mods:location>
<mods:classification id="26D73F105DD2325F96CB84034992FBFE">journal article</mods:classification>
<mods:identifier id="4DE2F0B7A1F65993669A25150A9F1573" type="DOI">10.11646/zootaxa.2260.1.8</mods:identifier>
<mods:identifier id="8AF7A76629BD87529E86D43E9E6CD28A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7023F408A9DF97F86E5325C8AF19F608" type="Zenodo-Dep">5322208</mods:identifier>
</mods:mods>
<treatment id="038E87C89D30FFB2FF6ACEEA6EA7FBF0" ID-DOI="http://doi.org/10.5281/zenodo.5334690" ID-Zenodo-Dep="5334690" LSID="urn:lsid:plazi:treatment:038E87C89D30FFB2FF6ACEEA6EA7FBF0" httpUri="http://treatment.plazi.org/id/038E87C89D30FFB2FF6ACEEA6EA7FBF0" lastPageId="38" lastPageNumber="191" pageId="35" pageNumber="188">
<subSubSection id="C33D65559D30FFB7FF6ACEEA6F96FE62" box="[151,746,422,448]" pageId="35" pageNumber="188" type="nomenclature">
<paragraph id="8B9836DE9D30FFB7FF6ACEEA6F96FE62" blockId="35.[151,746,422,483]" box="[151,746,422,448]" pageId="35" pageNumber="188">
<heading id="D0D081B29D30FFB7FF6ACEEA6F96FE62" bold="true" box="[151,746,422,448]" fontSize="11" level="1" pageId="35" pageNumber="188" reason="1">
<emphasis id="B953EACC9D30FFB7FF6ACEEA6F96FE62" bold="true" box="[151,746,422,448]" pageId="35" pageNumber="188">
<taxonomicName id="4C274D5D9D30FFB7FF6ACEEA6FFAFE62" authority="Hughes &amp; Lowry" authorityName="Hughes &amp; Lowry" box="[151,646,422,448]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="35" pageNumber="188" phylum="Arthropoda" rank="species" species="khbarnardi" status="sp. nov.">
<emphasis id="B953EACC9D30FFB7FF6ACEEA6CDAFE62" bold="true" box="[151,422,422,448]" italics="true" pageId="35" pageNumber="188">Cymadusa khbarnardi</emphasis>
Hughes &amp; Lowry
</taxonomicName>
<taxonomicNameLabel id="A26057B79D30FFB7FD70CEEA6F96FE62" box="[653,746,422,448]" pageId="35" pageNumber="188" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D65559D30FFB7FF6ACE856CFBFE41" box="[151,391,457,483]" pageId="35" pageNumber="188" type="description">
<paragraph id="8B9836DE9D30FFB7FF6ACE856CFBFE41" blockId="35.[151,746,422,483]" box="[151,391,457,483]" pageId="35" pageNumber="188">
(
<figureCitation id="131C2A5B9D30FFB7FF62CE856D86FE41" box="[159,250,457,483]" captionStart="FIGURE 21" captionStartId="36.[151,256,1979,2003]" captionTargetBox="[181,1411,158,1954]" captionTargetId="figure-40@36.[172,1416,143,1972]" captionTargetPageId="36" captionText="FIGURE 21. Cymadusa khbarnardi Hughes &amp; Lowry sp. nov., holotype, male, 9.0 mm, AM P76756, Lizard Island, Great Barrier Reef." figureDoi="http://doi.org/10.5281/zenodo.5322288" httpUri="https://zenodo.org/record/5322288/files/figure.png" pageId="35" pageNumber="188">Figs 21</figureCitation>
,
<figureCitation id="131C2A5B9D30FFB7FEFBCE856C5AFE41" box="[262,294,457,483]" captionStart="FIGURE 22" captionStartId="37.[151,255,1960,1984]" captionTargetBox="[197,1383,184,1928]" captionTargetId="figure-44@37.[178,1409,168,1950]" captionTargetPageId="37" captionText="FIGURE 22. Cymadusa khbarnardi Hughes &amp; Lowry sp. nov., holotype, male, 9.0 mm, AM P76756, paratype, female, 11.0 mm, AM P76757, Lizard Island, Great Barrier Reef." figureDoi="http://doi.org/10.5281/zenodo.5322292" httpUri="https://zenodo.org/record/5322292/files/figure.png" pageId="35" pageNumber="188">22</figureCitation>
, Pl. 1G)
</paragraph>
</subSubSection>
<subSubSection id="C33D65559D30FFB7FF6ACD5A6CA2FC9D" pageId="35" pageNumber="188" type="materials_examined">
<paragraph id="8B9836DE9D30FFB7FF6ACD5A6D9FFD06" blockId="35.[151,1436,534,2030]" pageId="35" pageNumber="188">
<emphasis id="B953EACC9D30FFB7FF6ACD5A6C2FFD92" bold="true" box="[151,339,534,560]" pageId="35" pageNumber="188">Type material.</emphasis>
<materialsCitation id="3B4F3C839D30FFB7FEA7CD5A6C51FDDC" accessionNumber="P76756" collectingDate="2005-02-28" collectionCode="AM, T" county="Krapp-Schickel" elevation="4" latitude="-14.6485" location="Mermaid Cove" longLatPrecision="13" longitude="145.45433" municipality="Lizard Island" pageId="35" pageNumber="188" specimenCode="QLD 1732" specimenCount="1" typeStatus="holotype">
<typeStatus id="549C887C9D30FFB7FEA7CD5A6CB0FD92" box="[346,460,534,560]" pageId="35" pageNumber="188" type="holotype">Holotype</typeStatus>
, male, 9.0 mm, 2 slides,
<collectionCode id="ED36AE1B9D30FFB7FD07CD5A6E55FD92" box="[762,809,534,560]" country="Australia" name="Australian Museum" pageId="35" pageNumber="188" type="Museum">AM</collectionCode>
P76756,
<location id="8EF860059D30FFB7FC63CD5A6929FD92" LSID="urn:lsid:plazi:treatment:038E87C89D30FFB2FF6ACEEA6EA7FBF0:8EF860059D30FFB7FC63CD5A6929FD92" box="[926,1109,534,560]" county="Krapp-Schickel" latitude="-14.6485" longLatPrecision="13" longitude="145.45433" municipality="Lizard Island" name="Mermaid Cove" pageId="35" pageNumber="188">Mermaid Cove</location>
,
<collectingMunicipality id="6BFCACA49D30FFB7FB9ECD5A687EFD92" box="[1123,1282,534,560]" pageId="35" pageNumber="188">Lizard Island</collectingMunicipality>
(
<geoCoordinate id="EE1350199D30FFB7FAE9CD5A68E0FD92" box="[1300,1436,534,560]" degrees="14" direction="south" minutes="38.91" orientation="latitude" pageId="35" pageNumber="188" precision="9" value="-14.6485">14°38.91’S</geoCoordinate>
<geoCoordinate id="EE1350199D30FFB7FF6ACD716C48FDF5" box="[151,308,573,599]" degrees="145" direction="east" minutes="27.26" orientation="longitude" pageId="35" pageNumber="188" precision="9" value="145.45433">145°27.26’E</geoCoordinate>
), mixed brown algae, sand with rubble bottom,
<quantity id="4CDF9B3B9D30FFB7FC82CD716EB4FDF5" box="[895,968,573,599]" metricMagnitude="0" metricUnit="m" metricValue="3.5" pageId="35" pageNumber="188" unit="m" value="3.5">
<elevation id="000AD1ED9D30FFB7FC82CD716EB4FDF5" box="[895,968,573,599]" metricMagnitude="0" metricUnit="m" metricValue="3.5" pageId="35" pageNumber="188" unit="m" value="3.5">3.5 m</elevation>
</quantity>
,
<collectionCode id="ED36AE1B9D30FFB7FC28CD716E95FDF5" box="[981,1001,573,599]" name="Tavera, Department of Geology and Geophysics" pageId="35" pageNumber="188">T</collectionCode>
.
<collectingCounty id="62F94E529D30FFB7FC0BCD7169CBFDF5" box="[1014,1207,573,599]" pageId="35" pageNumber="188">Krapp-Schickel</collectingCounty>
,
<date id="FF99101E9D30FFB7FB39CD7168E0FDF5" box="[1220,1436,573,599]" pageId="35" pageNumber="188" value="2005-02-28">
<collectingDate id="EFDDE9F69D30FFB7FB39CD7168E0FDF5" box="[1220,1436,573,599]" pageId="35" pageNumber="188" value="2005-02-28">28 February 2005</collectingDate>
</date>
(
<specimenCode id="DB819EA59D30FFB7FF62CD286C58FDDC" box="[159,292,612,638]" collectionCode="QLD" pageId="35" pageNumber="188">QLD 1732</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B4F3C839D30FFB7FEC4CD286956FDDC" accessionNumber="P76757" box="[313,1066,612,638]" collectionCode="AM" pageId="35" pageNumber="188" specimenCode="QLD 1732" specimenCount="1" typeStatus="paratype">
<typeStatus id="549C887C9D30FFB7FEC4CD286CD3FDDC" box="[313,431,612,638]" pageId="35" pageNumber="188" type="paratype">Paratypes</typeStatus>
: female, 11.0 mm, 3 slides,
<collectionCode id="ED36AE1B9D30FFB7FD06CD286E56FDDC" box="[763,810,612,638]" country="Australia" name="Australian Museum" pageId="35" pageNumber="188" type="Museum">AM</collectionCode>
P76757 (
<specimenCode id="DB819EA59D30FFB7FC61CD28695DFDDC" box="[924,1057,612,638]" collectionCode="QLD" pageId="35" pageNumber="188">QLD 1732</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B4F3C839D30FFB7FBCBCD286831FDDC" accessionNumber="P76758" box="[1078,1357,612,638]" collectionCode="AM" pageId="35" pageNumber="188" specimenCount="1" typeStatus="paratype">
3 unsexed,
<collectionCode id="ED36AE1B9D30FFB7FB40CD286990FDDC" box="[1213,1260,612,638]" country="Australia" name="Australian Museum" pageId="35" pageNumber="188" type="Museum">AM</collectionCode>
P76758
</materialsCitation>
<materialsCitation id="3B4F3C839D30FFB7FAABCD286DA3FD06" accessionNumber="P76758" collectionCode="QLD" isTailEnumeration="true" pageId="35" pageNumber="188" specimenCode="QLD 1732" specimenCount="1" typeStatus="paratype">
(
<specimenCode id="DB819EA59D30FFB7FAA3CD286DAAFD06" collectionCode="QLD" pageId="35" pageNumber="188">QLD 1732</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9836DE9D30FFB7FF3BCDFC6930FD50" blockId="35.[151,1436,534,2030]" pageId="35" pageNumber="188">
<emphasis id="B953EACC9D30FFB7FF3BCDFC6F3EFD68" bold="true" box="[198,578,688,714]" pageId="35" pageNumber="188">Additional material examined.</emphasis>
1 unsexed,
<collectionCode id="ED36AE1B9D30FFB7FD31CDFD6F87FD69" box="[716,763,689,715]" country="Australia" name="Australian Museum" pageId="35" pageNumber="188" type="Museum">AM</collectionCode>
P76760 (
<collectionCode id="ED36AE1B9D30FFB7FC96CDFD6EDDFD69" box="[875,929,689,715]" pageId="35" pageNumber="188">JDT</collectionCode>
/LIZ-14); male, photo 6,
<collectionCode id="ED36AE1B9D30FFB7FB3CCDFD698CFD69" box="[1217,1264,689,715]" country="Australia" name="Australian Museum" pageId="35" pageNumber="188" type="Museum">AM</collectionCode>
P71123 (
<collectionCode id="ED36AE1B9D30FFB7FAA3CDFD68E0FD69" box="[1374,1436,689,715]" pageId="35" pageNumber="188">QLD</collectionCode>
1730); 1 unsexed,
<collectionCode id="ED36AE1B9D30FFB7FE92CD946CE2FD50" box="[367,414,728,754]" country="Australia" name="Australian Museum" pageId="35" pageNumber="188" type="Museum">AM</collectionCode>
P71321 (
<collectionCode id="ED36AE1B9D30FFB7FDF2CD946F31FD50" box="[527,589,728,754]" pageId="35" pageNumber="188">QLD</collectionCode>
1787); 1 unsexed,
<collectionCode id="ED36AE1B9D30FFB7FCD1CD946E27FD50" box="[812,859,728,754]" country="Australia" name="Australian Museum" pageId="35" pageNumber="188" type="Museum">AM</collectionCode>
76759 (
<collectionCode id="ED36AE1B9D30FFB7FC46CD946E85FD50" box="[955,1017,728,754]" pageId="35" pageNumber="188">QLD</collectionCode>
1799).
</paragraph>
<paragraph id="8B9836DE9D30FFB7FF3BCDB26CA2FC9D" blockId="35.[151,1436,534,2030]" pageId="35" pageNumber="188">
<materialsCitation id="3B4F3C839D30FFB7FF3BCDB26CA6FC9D" elevation="4" latitude="-14.6485" location="Mermaid Cove" longLatPrecision="13" longitude="145.45433" municipality="Lizard Island" pageId="35" pageNumber="188" specimenCount="1" typeStatus="holotype">
<emphasis id="B953EACC9D30FFB7FF3BCDB26C13FCBA" bold="true" box="[198,367,766,792]" pageId="35" pageNumber="188">
<typeStatus id="549C887C9D30FFB7FF3BCDB26C78FCBA" box="[198,260,766,792]" pageId="35" pageNumber="188">Type</typeStatus>
locality.
</emphasis>
<location id="8EF860059D30FFB7FE84CDB26F53FCBA" LSID="urn:lsid:plazi:treatment:038E87C89D30FFB2FF6ACEEA6EA7FBF0:8EF860059D30FFB7FE84CDB26F53FCBA" box="[377,559,766,792]" latitude="-14.6485" longLatPrecision="13" longitude="145.45433" municipality="Lizard Island" name="Mermaid Cove" pageId="35" pageNumber="188">Mermaid Cove</location>
,
<collectingMunicipality id="6BFCACA49D30FFB7FDC3CDB26FA7FCBA" box="[574,731,766,792]" pageId="35" pageNumber="188">Lizard Island</collectingMunicipality>
(
<geoCoordinate id="EE1350199D30FFB7FD12CDB26E09FCBA" box="[751,885,766,792]" degrees="14" direction="south" minutes="38.91" orientation="latitude" pageId="35" pageNumber="188" precision="9" value="-14.6485">14°38.91’S</geoCoordinate>
<geoCoordinate id="EE1350199D30FFB7FC82CDB26964FCBA" box="[895,1048,766,792]" degrees="145" direction="east" minutes="27.26" orientation="longitude" pageId="35" pageNumber="188" precision="9" value="145.45433">145°27.26’E</geoCoordinate>
), mixed brown algae, sand with rubble bottom,
<elevation id="000AD1ED9D30FFB7FEB6CC696CA6FC9D" box="[331,474,805,831]" metricMagnitude="0" metricUnit="m" metricValue="3.5" pageId="35" pageNumber="188" unit="m" value="3.5">
<quantity id="4CDF9B3B9D30FFB7FEB6CC696CF2FC9D" box="[331,398,805,831]" metricMagnitude="0" metricUnit="m" metricValue="3.5" pageId="35" pageNumber="188" unit="m" value="3.5">3.5 m</quantity>
depth
</elevation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D65559D30FFB7FF3BCC076823FCC4" box="[198,1375,843,870]" pageId="35" pageNumber="188" type="etymology">
<paragraph id="8B9836DE9D30FFB7FF3BCC076823FCC4" blockId="35.[151,1436,534,2030]" box="[198,1375,843,870]" pageId="35" pageNumber="188">
<emphasis id="B953EACC9D30FFB7FF3BCC076C2FFCC7" bold="true" box="[198,339,843,869]" pageId="35" pageNumber="188">Etymology.</emphasis>
Named for Keppel Harcourt Barnard, the first student of Great Barrier Reef amphipods.
</paragraph>
</subSubSection>
<subSubSection id="C33D65559D30FFB2FF3BCC3E6CAEFD6D" lastPageId="38" lastPageNumber="191" pageId="35" pageNumber="188" type="description">
<paragraph id="8B9836DE9D30FFB7FF3BCC3E6EEAFC2E" blockId="35.[151,1436,534,2030]" box="[198,918,882,908]" pageId="35" pageNumber="188">
<emphasis id="B953EACC9D30FFB7FF3BCC3E6C22FC2E" bold="true" box="[198,350,882,908]" pageId="35" pageNumber="188">Description.</emphasis>
Based on
<typeStatus id="549C887C9D30FFB7FE24CC3E6F3EFC2E" box="[473,578,882,908]" pageId="35" pageNumber="188" type="holotype">holotype</typeStatus>
, male, 9.0 mm, AM P76756.
</paragraph>
<paragraph id="8B9836DE9D30FFB7FF3BCCD46E04FB60" blockId="35.[151,1436,534,2030]" pageId="35" pageNumber="188">
<emphasis id="B953EACC9D30FFB7FF3BCCD46C6CFC10" bold="true" box="[198,272,920,946]" pageId="35" pageNumber="188">Head.</emphasis>
<emphasis id="B953EACC9D30FFB7FEE5CCD56C2BFC10" box="[280,343,921,946]" italics="true" pageId="35" pageNumber="188">Head</emphasis>
slightly longer than deep.
<emphasis id="B953EACC9D30FFB7FD69CCD56E70FC10" box="[660,780,921,946]" italics="true" pageId="35" pageNumber="188">Antenna 1</emphasis>
longer than antenna 2; peduncular article 1 subequal to article 2; flagellum 30 articles; accessory flagellum with 1 article.
<emphasis id="B953EACC9D30FFB7FC56CC8C6958FC7B" box="[939,1060,960,985]" italics="true" pageId="35" pageNumber="188">Antenna 2</emphasis>
peduncle not densely setose on ventral margin; article 4 subequal in length to article 5; flagellum 26 articles.
<emphasis id="B953EACC9D30FFB7FBDDCCAB69ECFBA2" box="[1056,1168,999,1024]" italics="true" pageId="35" pageNumber="188">Mandible</emphasis>
molar well developed, triturating; accessory setal row with 5 robust serrate setae; palp 3–articulate, article 3 apically setose; article 1 longer than article 2; article 2 shorter than article 3; article 3 longer than article 1.
<emphasis id="B953EACC9D30FFB7FB72CB786879FBEF" box="[1167,1285,1076,1101]" italics="true" pageId="35" pageNumber="188">Lower lip</emphasis>
outer plates notched, forming a slight medial excavation; mandibular lobe with curved margins, rounded apically.
<emphasis id="B953EACC9D30FFB7FABCCB176DDAFB38" italics="true" pageId="35" pageNumber="188">Maxilla 1</emphasis>
inner plate with 2 setae; palp article 2 broad.
<emphasis id="B953EACC9D30FFB7FD29CBCD6E34FB38" box="[724,840,1153,1178]" italics="true" pageId="35" pageNumber="188">Maxilla 2</emphasis>
inner plate narrower than outer plate.
<emphasis id="B953EACC9D30FFB7FAE7CBCD68E7FB38" box="[1306,1435,1153,1178]" italics="true" pageId="35" pageNumber="188">Maxilliped</emphasis>
outer plate with row of large robust setae along medial margin.
</paragraph>
<paragraph id="8B9836DE9D30FFB7FF3BCB826C5DF8F1" blockId="35.[151,1436,534,2030]" pageId="35" pageNumber="188">
<emphasis id="B953EACC9D30FFB7FF3BCB826C59FB4A" bold="true" box="[198,293,1230,1256]" pageId="35" pageNumber="188">Pereon.</emphasis>
<emphasis id="B953EACC9D30FFB7FED3CB836CD3FB4A" box="[302,431,1231,1256]" italics="true" pageId="35" pageNumber="188">Coxae 1–4</emphasis>
deeper than broad.
<emphasis id="B953EACC9D30FFB7FD62CB836E47FB4A" box="[671,827,1231,1256]" italics="true" pageId="35" pageNumber="188">Gnathopod 1</emphasis>
subequal in size to gnathopod 2; coxa subequal to coxa 2, slightly produced, anterior margin convex, anteroventral corner rounded, with a tuft of slender setae in the posteroventral corner; basis longer than coxa, anterodistal lobe absent; merus posterodistal lobe subacute; carpus about 3 x as long as broad, longer than propodus, anterior margin with slender setae distally, posterior margin straight; propodus broad, subtriangular; palm acute, straight, with posterodistal corner rounded, with 1 robust seta defining palm; dactylus subequal in length to palm, apically acute, inner margin crenate.
<emphasis id="B953EACC9D30FFB7FF6ACAFB6C48FA72" box="[151,308,1463,1488]" italics="true" pageId="35" pageNumber="188">Gnathopod 2</emphasis>
, coxa with a tuft of slender setae in the posteroventral corner; basis with sparse slender setae, anterodistal corner with 1 slender seta; merus margin with long, rounded anterodistal lobe; carpus much shorter than propodus, subtriangular; propodus broad, less than 1.5 x as long as broad, subquadrate, not produced into an anterodistally setose lobe; palm acute, excavate, with posterodistal corner subquadrate, without palm defining robust setae; dactylus shorter than palm, tapering evenly, apically acute, inner margin crenate, with 3 groups of long slender setae on anterior margin.
<emphasis id="B953EACC9D30FFB7FC75C9346944F933" box="[904,1080,1656,1681]" italics="true" pageId="35" pageNumber="188">Pereopods 3–4</emphasis>
basis narrow; merus subequal in length to carpus.
<emphasis id="B953EACC9D30FFB7FE78C9D36F4BF91A" box="[389,567,1695,1720]" italics="true" pageId="35" pageNumber="188">Pereopods 5–7</emphasis>
simple.
<emphasis id="B953EACC9D30FFB7FD62C9D36E55F91A" box="[671,809,1695,1720]" italics="true" pageId="35" pageNumber="188">Pereopod 5</emphasis>
without medial slender setae; distal articles slender; propodus not expanded distally, without anterodistal striated robust setae; dactylus slightly curved.
<emphasis id="B953EACC9D30FFB7FAD6C9896DDAF8A7" italics="true" pageId="35" pageNumber="188">Pereopod 6</emphasis>
basis posterior margin straight, with several marginal robust setae; distal articles slender; propodus not expanded distally, without anterodistal striated robust setae; dactylus slightly curved.
<emphasis id="B953EACC9D30FFB7FB65C85F685EF88E" box="[1176,1314,1811,1836]" italics="true" pageId="35" pageNumber="188">Pereopod 7</emphasis>
similar to pereopod 6.
</paragraph>
<paragraph id="8B9836DE9D30FFB2FF3BC8136CCDFED1" blockId="35.[151,1436,534,2030]" lastBlockId="38.[151,1436,152,1106]" lastPageId="38" lastPageNumber="191" pageId="35" pageNumber="188">
<emphasis id="B953EACC9D30FFB7FF3BC8136C6FF8DB" bold="true" box="[198,275,1887,1913]" pageId="35" pageNumber="188">Pleon.</emphasis>
<emphasis id="B953EACC9D30FFB7FEE1C82C6CC8F8DB" box="[284,436,1888,1913]" italics="true" pageId="35" pageNumber="188">Epimera 2–3</emphasis>
posteroventral corner broadly rounded.
<emphasis id="B953EACC9D30FFB7FC72C82C697CF8DB" box="[911,1024,1888,1913]" italics="true" pageId="35" pageNumber="188">Uropod 1</emphasis>
peduncle with 7 robust setae, with short fringe of slender setae (less than 0.5 x length of peduncle), with large, acute distoventral spine; outer ramus length about 5–6 x as long as broad, with 9 lateral robust setae; inner ramus with 10 lateral robust setae.
<emphasis id="B953EACC9D30FFB7FF6AC8986C75F84F" box="[151,265,2004,2029]" italics="true" pageId="35" pageNumber="188">Uropod 2</emphasis>
peduncle with 3 robust setae; outer ramus with 7 lateral robust setae; inner ramus with 12 lateral robust setae.
<emphasis id="B953EACC9D35FFB2FEC1CFD46CCEFF13" box="[316,434,152,177]" italics="true" pageId="38" pageNumber="191">Uropod 3</emphasis>
peduncle longer than broad, less than 2 x length of rami, marginal slender setae present, with 5 distal and 2 lateral robust setae, with 2 distal slender setae; outer ramus with 2 large recurved distal robust setae, with 1 lateral robust seta and 4 lateral slender setae; inner ramus with 6 distal slender setae, with 2 lateral robust setae.
<taxonomicName id="4C274D5D9D35FFB2FE08CE406F3AFE87" authorityName="Pearse" authorityYear="1952" box="[501,582,268,293]" class="Hexanauplia" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="38" pageNumber="191" phylum="Arthropoda" rank="genus">
<emphasis id="B953EACC9D35FFB2FE08CE406F3AFE87" box="[501,582,268,293]" italics="true" pageId="38" pageNumber="191">Telson</emphasis>
</taxonomicName>
subtrapezoidal, apically truncated, with small apical cusp on each distolateral corner, with 4 pair of oblique medial slender setae, with 4 pair of lateral slender setae, and 1 pair of apical plumose setae.
</paragraph>
<caption id="DF5866569D37FFB0FF6AC8F76C19F857" ID-DOI="http://doi.org/10.5281/zenodo.5322288" ID-Zenodo-Dep="5322288" httpUri="https://zenodo.org/record/5322288/files/figure.png" pageId="36" pageNumber="189" startId="36.[151,256,1979,2003]" targetBox="[181,1411,158,1954]" targetPageId="36">
<paragraph id="8B9836DE9D37FFB0FF6AC8F76C19F857" blockId="36.[151,1435,1979,2037]" pageId="36" pageNumber="189">
<emphasis id="B953EACC9D37FFB0FF6AC8F76C55F871" bold="true" box="[151,297,1979,2003]" pageId="36" pageNumber="189">FIGURE 21.</emphasis>
<taxonomicName id="4C274D5D9D37FFB0FECCC8F76F90F870" authority="Hughes &amp; Lowry" authorityName="Hughes &amp; Lowry" box="[305,748,1979,2002]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="36" pageNumber="189" phylum="Arthropoda" rank="species" species="khbarnardi" status="sp. nov.">
<emphasis id="B953EACC9D37FFB0FECCC8F76F59F870" box="[305,549,1979,2002]" italics="true" pageId="36" pageNumber="189">Cymadusa khbarnardi</emphasis>
Hughes &amp; Lowry
</taxonomicName>
<emphasis id="B953EACC9D37FFB0FD09C8F76E30F871" bold="true" box="[756,844,1979,2003]" pageId="36" pageNumber="189">
<taxonomicNameLabel id="A26057B79D37FFB0FD09C8F76E30F871" box="[756,844,1979,2003]" pageId="36" pageNumber="189" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male, 9.0 mm, AM P76756, Lizard Island, Great Barrier Reef.
</paragraph>
</caption>
<caption id="DF5866569D36FFB1FF6AC8E46F87F847" ID-DOI="http://doi.org/10.5281/zenodo.5322292" ID-Zenodo-Dep="5322292" httpUri="https://zenodo.org/record/5322292/files/figure.png" pageId="37" pageNumber="190" startId="37.[151,255,1960,1984]" targetBox="[197,1383,184,1928]" targetPageId="37">
<paragraph id="8B9836DE9D36FFB1FF6AC8E46F87F847" blockId="37.[151,1436,1960,2021]" pageId="37" pageNumber="190">
<emphasis id="B953EACC9D36FFB1FF6AC8E46C5BF862" bold="true" box="[151,295,1960,1984]" pageId="37" pageNumber="190">FIGURE 22.</emphasis>
<taxonomicName id="4C274D5D9D36FFB1FED0C8E56FA1F862" authority="Hughes &amp; Lowry" authorityName="Hughes &amp; Lowry" box="[301,733,1961,1984]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="37" pageNumber="190" phylum="Arthropoda" rank="species" species="khbarnardi" status="sp. nov.">
<emphasis id="B953EACC9D36FFB1FED0C8E56F60F862" box="[301,540,1961,1984]" italics="true" pageId="37" pageNumber="190">Cymadusa khbarnardi</emphasis>
Hughes &amp; Lowry
</taxonomicName>
<emphasis id="B953EACC9D36FFB1FD19C8E46E46F862" bold="true" box="[740,826,1960,1984]" pageId="37" pageNumber="190">
<taxonomicNameLabel id="A26057B79D36FFB1FD19C8E46E46F862" box="[740,826,1960,1984]" pageId="37" pageNumber="190" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male, 9.0 mm, AM P76756, paratype, female, 11.0 mm, AM P76757, Lizard Island, Great Barrier Reef.
</paragraph>
</caption>
<paragraph id="8B9836DE9D35FFB2FF3BCE336D8FFDF9" blockId="38.[151,1436,152,1106]" pageId="38" pageNumber="191">
<emphasis id="B953EACC9D35FFB2FF3BCE336C5DFE3B" bold="true" box="[198,289,383,409]" pageId="38" pageNumber="191">Female</emphasis>
(sexually dimorphic characters).
<materialsCitation id="3B4F3C839D35FFB2FD54CECC6D93FDF9" accessionNumber="P76757" collectionCode="AM" pageId="38" pageNumber="191" specimenCount="1" typeStatus="paratype">
Based on
<typeStatus id="549C887C9D35FFB2FCE1CECC6EF8FE38" box="[796,900,384,410]" pageId="38" pageNumber="191" type="paratype">paratype</typeStatus>
, female, 11.0 mm,
<collectionCode id="ED36AE1B9D35FFB2FB99CECC69EFFE38" box="[1124,1171,384,410]" country="Australia" name="Australian Museum" pageId="38" pageNumber="191" type="Museum">AM</collectionCode>
P76757.
<emphasis id="B953EACC9D35FFB2FAFFCECC68E0FE3B" box="[1282,1436,384,409]" italics="true" pageId="38" pageNumber="191">Gnathopod 1</emphasis>
subequal in size to gnathopod 2; coxa anteroventrally produced, anterior margin concave; basis shorter than coxa, anterodistal lobe rounded, without setae; merus anterodistal lobe acute; carpus about 2 x as long as broad, shorter than propodus.
<emphasis id="B953EACC9D35FFB2FE0BCEB86FECFDAF" box="[502,656,500,525]" italics="true" pageId="38" pageNumber="191">Gnathopod 2</emphasis>
basis without setae, anterodistal lobe without setae; carpus shorter than propodus; propodus subrectangular; palm straight; dactylus subequal in length to palm, inner margin crenate
</materialsCitation>
.
</paragraph>
<paragraph id="8B9836DE9D35FFB2FF3BCD2B6D97FD0A" blockId="38.[151,1436,152,1106]" pageId="38" pageNumber="191">
<emphasis id="B953EACC9D35FFB2FF3BCD2B6C5FFD23" bold="true" box="[198,291,615,641]" pageId="38" pageNumber="191">Colour.</emphasis>
Body clear to white with small dark brown and white spots, eyes with red ommatidia in a white matrix.
</paragraph>
<paragraph id="8B9836DE9D35FFB2FF3BCDF96CAEFD6D" blockId="38.[151,1436,152,1106]" box="[198,466,693,719]" pageId="38" pageNumber="191">
<emphasis id="B953EACC9D35FFB2FF3BCDF96C51FD6D" bold="true" box="[198,301,693,719]" pageId="38" pageNumber="191">Habitat.</emphasis>
Brown algae.
</paragraph>
</subSubSection>
<subSubSection id="C33D65559D35FFB2FF3BCD976870FB89" pageId="38" pageNumber="191" type="discussion">
<paragraph id="8B9836DE9D35FFB2FF3BCD976F88FCC8" blockId="38.[151,1436,152,1106]" pageId="38" pageNumber="191">
<emphasis id="B953EACC9D35FFB2FF3BCD976C43FD57" bold="true" box="[198,319,731,757]" pageId="38" pageNumber="191">Remarks.</emphasis>
<taxonomicName id="4C274D5D9D35FFB2FEADCD906F23FD57" box="[336,607,732,757]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="38" pageNumber="191" phylum="Arthropoda" rank="species" species="khbarnardi" status="sp. nov.">
<emphasis id="B953EACC9D35FFB2FEADCD906F23FD57" box="[336,607,732,757]" italics="true" pageId="38" pageNumber="191">Cymadusa khbarnardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26057B79D35FFB2FD92CD976FAAFD54" box="[623,726,731,758]" pageId="38" pageNumber="191" rank="species">
<emphasis id="B953EACC9D35FFB2FD92CD976FB3FD57" bold="true" box="[623,719,731,757]" pageId="38" pageNumber="191">sp. nov</emphasis>
.
</taxonomicNameLabel>
is similar to
<taxonomicName id="4C274D5D9D35FFB2FC69CD906876FD54" authority="Ledoyer, 1984" authorityName="Ledoyer" authorityYear="1984" box="[916,1290,732,758]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="38" pageNumber="191" phylum="Arthropoda" rank="species" species="grossimana">
<emphasis id="B953EACC9D35FFB2FC69CD906EDBFD57" box="[916,935,732,757]" italics="true" pageId="38" pageNumber="191">C</emphasis>
.
<emphasis id="B953EACC9D35FFB2FC42CD90693BFD57" box="[959,1095,732,757]" italics="true" pageId="38" pageNumber="191">grossimana</emphasis>
<bibRefCitation id="EFB64B2F9D35FFB2FBA5CD906876FD54" author="Ledoyer" box="[1112,1290,732,758]" firstAuthor="Ledoyer" pageId="38" pageNumber="191" pagination="1 - 113" refId="ref26173" refString="Ledoyer, M. (1984) Les gammariens (Crustacea, Amphipoda) des herbiers de phanerogames marines de Nouvelle Caledonie (region de Noumea). Memoires du Museum National d'Histoire Naturelle, Series A, Zoology, 129, 1 - 113." type="journal article" year="1984">Ledoyer, 1984</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F330764E9D35FFB2FA98CD906C6DFCBE" name="New Caledonia" pageId="38" pageNumber="191">New Caledonia</collectingCountry>
, in the propodus and palm shape of gnathopod 2. These species differ as follows: in
<taxonomicName id="4C274D5D9D35FFB2FB0BCC4E68E0FCB9" box="[1270,1436,770,795]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="38" pageNumber="191" phylum="Arthropoda" rank="species" species="khbarnardi">
<emphasis id="B953EACC9D35FFB2FB0BCC4E68E0FCB9" box="[1270,1436,770,795]" italics="true" pageId="38" pageNumber="191">C. khbarnardi</emphasis>
</taxonomicName>
the carpus of gnathopod 1 is extremely elongate (longer than the propodus) and rectolinear, while in
<taxonomicName id="4C274D5D9D35FFB2FA7DCC656C63FCCB" authorityName="Ledoyer" authorityYear="1984" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="38" pageNumber="191" phylum="Arthropoda" rank="species" species="grossimana">
<emphasis id="B953EACC9D35FFB2FA7DCC656C63FCCB" italics="true" pageId="38" pageNumber="191">C. grossimana</emphasis>
</taxonomicName>
it is subequal in length to the propodus.
</paragraph>
<paragraph id="8B9836DE9D35FFB2FF3BCC3B6870FB89" blockId="38.[151,1436,152,1106]" pageId="38" pageNumber="191">
<taxonomicName id="4C274D5D9D35FFB2FF3BCC3B6CB3FC32" box="[198,463,887,912]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="38" pageNumber="191" phylum="Arthropoda" rank="species" species="khbarnardi" status="sp. nov.">
<emphasis id="B953EACC9D35FFB2FF3BCC3B6CB3FC32" box="[198,463,887,912]" italics="true" pageId="38" pageNumber="191">Cymadusa khbarnardi</emphasis>
</taxonomicName>
<emphasis id="B953EACC9D35FFB2FE24CC3A6F45FC32" bold="true" box="[473,569,886,912]" pageId="38" pageNumber="191">
<taxonomicNameLabel id="A26057B79D35FFB2FE24CC3A6F45FC32" box="[473,569,886,912]" pageId="38" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
forms a group with two other GBR species
<taxonomicName id="4C274D5D9D35FFB2FBA8CC3B6DAFFC15" authority="Rabindranath, 1972" authorityName="Rabindranath" authorityYear="1972" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="38" pageNumber="191" phylum="Arthropoda" rank="species" species="imbroglio">
<emphasis id="B953EACC9D35FFB2FBA8CC3B6991FC32" box="[1109,1261,887,912]" italics="true" pageId="38" pageNumber="191">C. imbroglio</emphasis>
<bibRefCitation id="EFB64B2F9D35FFB2FB0ACC3A6DAFFC15" author="Rabindranath" firstAuthor="Rabindranath" pageId="38" pageNumber="191" pagination="161 - 178" refId="ref26839" refString="Rabindranath, P. (1972) Marine Gammaridea (Crustacea: Amphipoda) from the Indian region. Family Ampithoidae. Marine Biology, 14 (2), 161 - 178." type="journal article" year="1972">Rabindranath, 1972</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C274D5D9D35FFB2FEECCCD16CC1FC14" authorityName="Hughes &amp; Lowry" authorityYear="2009" box="[273,445,925,950]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="38" pageNumber="191" phylum="Arthropoda" rank="species" species="smilodonta" status="sp. nov.">
<emphasis id="B953EACC9D35FFB2FEECCCD16CC1FC14" box="[273,445,925,950]" italics="true" pageId="38" pageNumber="191">C. smilodonta</emphasis>
</taxonomicName>
<emphasis id="B953EACC9D35FFB2FE3BCCD16F54FC15" bold="true" box="[454,552,925,951]" pageId="38" pageNumber="191">
<taxonomicNameLabel id="A26057B79D35FFB2FE3BCCD16F54FC15" box="[454,552,925,951]" pageId="38" pageNumber="191" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
based on the large sized individuals with submassive, broadly rounded propodus of male gnathopod 2. These species may be distinguished from one another by the sculpting on the palm of gnathopod 2. In
<taxonomicName id="4C274D5D9D35FFB2FE40CCA66FB8FBA1" box="[445,708,1002,1027]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="38" pageNumber="191" phylum="Arthropoda" rank="species" species="khbarnardi">
<emphasis id="B953EACC9D35FFB2FE40CCA66FB8FBA1" box="[445,708,1002,1027]" italics="true" pageId="38" pageNumber="191">Cymadusa khbarnardi</emphasis>
</taxonomicName>
the palm of gnathopod 2 is excavate and smooth, while in
<taxonomicName id="4C274D5D9D35FFB2FA7CCCA66C75FB88" authorityName="Rabindranath" authorityYear="1972" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="38" pageNumber="191" phylum="Arthropoda" rank="species" species="imbroglio">
<emphasis id="B953EACC9D35FFB2FA7CCCA66C75FB88" italics="true" pageId="38" pageNumber="191">C. imbroglio</emphasis>
</taxonomicName>
it has a midmedial projection and in
<taxonomicName id="4C274D5D9D35FFB2FD40CB5D6E1EFB88" authorityName="Hughes &amp; Lowry" authorityYear="2009" box="[701,866,1041,1066]" class="Malacostraca" family="Ampithoidae" genus="Cymadusa" kingdom="Animalia" order="Amphipoda" pageId="38" pageNumber="191" phylum="Arthropoda" rank="species" species="smilodonta">
<emphasis id="B953EACC9D35FFB2FD40CB5D6E1EFB88" box="[701,866,1041,1066]" italics="true" pageId="38" pageNumber="191">C. smilodonta</emphasis>
</taxonomicName>
it has a proximal palmar projection.
</paragraph>
</subSubSection>
<subSubSection id="C33D65559D35FFB2FF3BCB7B6EA7FBF0" box="[198,987,1079,1106]" pageId="38" pageNumber="191" type="distribution">
<paragraph id="8B9836DE9D35FFB2FF3BCB7B6EA7FBF0" blockId="38.[151,1436,152,1106]" box="[198,987,1079,1106]" pageId="38" pageNumber="191">
<emphasis id="B953EACC9D35FFB2FF3BCB7B6C1BFBF3" bold="true" box="[198,359,1079,1105]" pageId="38" pageNumber="191">Distribution.</emphasis>
<emphasis id="B953EACC9D35FFB2FE93CB746CA7FBF3" box="[366,475,1080,1105]" italics="true" pageId="38" pageNumber="191">
<collectingCountry id="F330764E9D35FFB2FE93CB746CA7FBF3" box="[366,475,1080,1105]" name="Australia" pageId="38" pageNumber="191">Australia</collectingCountry>
</emphasis>
.
<collectingRegion id="49E3F83C9D35FFB2FE17CB746F05FBF0" box="[490,633,1080,1106]" country="Australia" name="Queensland" pageId="38" pageNumber="191">Queensland</collectingRegion>
: Lizard Island (current study).
</paragraph>
</subSubSection>
</treatment>
</document>
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