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<document ID-CLB-Dataset="55738" ID-DOI="http://dx.doi.org/10.3897/zookeys.1098.73223" ID-GBIF-Dataset="8b26ea49-e662-41f7-a213-47f4b598c91c" ID-PMC="PMC9848896" ID-Pensoft-Pub="1313-2970-1098-1" ID-Pensoft-UUID="B22184E75B1A59C9B985C0DD9F092196" ID-PubMed="36761078" ID-ZooBank="B4F4033F296E43CCBE54B9413BC19268" ModsDocID="1313-2970-1098-1" checkinTime="1651618226429" checkinUser="pensoft" docAuthor="Rakotonirina, Jean Claude &amp; Fisher, Brian L." docDate="2022" docId="E59160DA8FA65AB8B878429A200E7150" docLanguage="en" docName="ZooKeys 1098: 1-180" docOrigin="ZooKeys 1098" docPubDate="2022-05-03" docSource="http://dx.doi.org/10.3897/zookeys.1098.73223" docTitle="Camponotus kelimaso Rakotonirina &amp; Fisher 2022, sp. nov." docType="treatment" docUuid="BDE5DB12-AFBD-41F7-95F5-966AF8E790F3" docUuidSource="ZooBank" docVersion="5" id="B22184E75B1A59C9B985C0DD9F092196" lastPageNumber="1" masterDocId="B22184E75B1A59C9B985C0DD9F092196" masterDocTitle="Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1732660529870" updateUser="ExternalLinkService">
<mods:mods id="30497D88FCEC3F10579616CF9AF41305" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2083458AEC47D4EE557310E132E8D06F">
<mods:title id="E09E0B913410D0A0970F2F4FDF756BBC">Revision of the Malagasy Camponotus subgenus Myrmosaga (Hymenoptera, Formicidae) using qualitative and quantitative morphology</mods:title>
</mods:titleInfo>
<mods:name id="9F30C11C8D4E2811DF854D38B4328D44" type="personal">
<mods:role id="9D6EBD5C7BED21F1961D5B52255D73D2">
<mods:roleTerm id="42A7F162210D05FB1457E13AD1916FFF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8D72F59742C50FBE85CF62F6AF8588D1">Rakotonirina, Jean Claude</mods:namePart>
<mods:affiliation id="572A2474136D66175C15EBD28089AE12">Madagascar Biodiversity Center, BP 6257, Parc Botanique et Zoologique de Tsimbazaza, Antananarivo, Madagascar &amp; Departement d'Entomologie-Culture, Elevage, Sante; Faculte des Sciences, BP 906, Universite d'Antananarivo, Antananarivo, Madagascar</mods:affiliation>
<mods:nameIdentifier id="3361C2B7B9B01FE8C1D74FE113B146F9" type="email">jcrakoto25@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="72444971B9F22CCAC9F3CDB3B8C95E06" type="personal">
<mods:role id="F22D42BB28D54EC2504C9A133D91253B">
<mods:roleTerm id="F9E7CDE8FDEA3030A37D87E9F2CF100A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4A47CC7DF4CE078517C26E7B82424A9D">Fisher, Brian L.</mods:namePart>
<mods:nameIdentifier id="F328ED4E2A4D287D94B628B9D17DE864" type="ORCID">https://orcid.org/0000-0002-4653-3270</mods:nameIdentifier>
<mods:affiliation id="3CFE47830A4258DF08B70854C93ABDFA">Entomology, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource id="257BFB6B3251BD056E105FB89BA37CCE">text</mods:typeOfResource>
<mods:relatedItem id="82F55B54A4B4B6AFB74A56400FAC5131" type="host">
<mods:titleInfo id="DF9A72D805B11EE5892E922C6A7AF5AA">
<mods:title id="82A15F53A582D9E022596430CB62E8AD">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="EAEED073C7ACE250318708B332183425">
<mods:date id="33EB3418F0238F67A4DD4305F202005B">2022</mods:date>
<mods:detail id="7A9384E7FDB30783ADD60F2908D2FE3A" type="pubDate">
<mods:number id="5DD96F598088876A4EAC66FEA9D38EF8">2022-05-03</mods:number>
</mods:detail>
<mods:detail id="22F685260ACD645BC75B1FD852997198" type="volume">
<mods:number id="DB50834830C175852186B5E5E8F1A8D4">1098</mods:number>
</mods:detail>
<mods:extent id="26F742FC8CB4DF8CEBF872E3B5473E6D" unit="page">
<mods:start id="D3ECCA8462CE133AEF76BE0287EC82E4">1</mods:start>
<mods:end id="A7B2168F6F9A2E84481D8B00FADC6B16">180</mods:end>
</mods:extent>
</mods:part>
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<mods:location id="15E7A17E03A3D57DD78BCED38CF0B74C">
<mods:url id="83CE61F7CDAF06E87585987046B7A1B1">http://dx.doi.org/10.3897/zookeys.1098.73223</mods:url>
</mods:location>
<mods:classification id="FE1BF6749C63666C9DE1EC84B448EC3F">journal article</mods:classification>
<mods:identifier id="F365F7CDEF53F4BE9BD2CB3542C24895" type="DOI">http://dx.doi.org/10.3897/zookeys.1098.73223</mods:identifier>
<mods:identifier id="9B0EF8DE66D134245C01D72F7B693E4B" type="Pensoft-Pub">1313-2970-1098-1</mods:identifier>
<mods:identifier id="2D818DC500FF46351FEEED9D2986E6AE" type="ZooBank">B4F4033F296E43CCBE54B9413BC19268</mods:identifier>
<mods:identifier id="F81DD16C99EFA5BC46949EDE33680F03" type="Pensoft-UUID">B22184E75B1A59C9B985C0DD9F092196</mods:identifier>
</mods:mods>
<treatment id="E59160DA8FA65AB8B878429A200E7150" ID-GBIF-Taxon="195190280" LSID="urn:lsid:zoobank.org:act:BDE5DB12-AFBD-41F7-95F5-966AF8E790F3" httpUri="http://treatment.plazi.org/id/E59160DA8FA65AB8B878429A200E7150" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="3CA360EB34DA12B34243C6C2A3B8174A" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="50AC9665A852C2322F9BA7B8B05C14DE" pageId="0" pageNumber="1">
<taxonomicName id="1F19FA37F13FF323373259ADD1FD021C" LSID="http://zoobank.org/BDE5DB12-AFBD-41F7-95F5-966AF8E790F3" authority="Rakotonirina &amp; Fisher, 2022" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus kelimaso" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kelimaso" status="sp. nov.">Camponotus kelimaso</taxonomicName>
<taxonomicNameLabel id="0AED3FFAFE3BA91273679E796166C314" pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3CDDD224C8944152248C7B6C4A407A5" pageId="0" pageNumber="1" type="description">
<paragraph id="48A1CCAFC95268B3F908E7A8B938BE7A" pageId="0" pageNumber="1">
<figureCitation id="987678E495C8B5AB5281B71993F2A05C" captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Head in full-face view A C. liandia (CASENT 0491145) B C. kelimaso (CASENT 0487718)." figureDoi="10.3897/zookeys.1098.73223.figure34" httpUri="https://binary.pensoft.net/fig/678093" pageId="0" pageNumber="1">Figs 34B</figureCitation>
<figureCitation id="8E367AD7495538053BE3EBCF5D78176B" captionStart="Figure 35" captionStartId="F35" captionText="Figure 35. Second and third abdominal tergites in dorsal view A C. kelimaso (CASENT 0487718) B C. quadrimaculatus (CASENT 0096044) C C. gibber (CASENT 0491497)." figureDoi="10.3897/zookeys.1098.73223.figure35" httpUri="https://binary.pensoft.net/fig/678094" pageId="0" pageNumber="1">, 35A</figureCitation>
<figureCitation id="3FDE60060634F25AB822A1CE08ECC838" captionStart="Figure 37" captionStartId="F37" captionText="Figure 37. Head in full-face view A C. kelimaso (CASENT 0487718) B C. lubbocki (CASENT 0486998)." figureDoi="10.3897/zookeys.1098.73223.figure37" httpUri="https://binary.pensoft.net/fig/678096" pageId="0" pageNumber="1">, 37A</figureCitation>
<figureCitation id="81F5AA9075C413C5E9DDD1CC4BED5A83" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Camponotus kelimaso A lateral view B head in full-face view C dorsal view of holotype minor worker CASENT 0487718 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure63" httpUri="https://binary.pensoft.net/fig/678122" pageId="0" pageNumber="1">, 63</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="3939251384696F1B1208B7FFB2E00D5E" pageId="0" pageNumber="1" type="holotype worker">
<paragraph id="E7D27DDD123AA9FFA5C9F2A8AF198658" pageId="0" pageNumber="1">Holotype worker.</paragraph>
<paragraph id="CB100A0CACF157A9202599C7C6234375" pageId="0" pageNumber="1">
<emphasis id="2090951D03BD26315C15C98BD16040E5" bold="true" pageId="0" pageNumber="1">Madagascar</emphasis>
: Province
<emphasis id="4F167D878CA49B4F7CE1F942385DDD8C" bold="true" pageId="0" pageNumber="1">Antsiranana</emphasis>
: PN Marojejy, Manantenina River, 27.6 km 35° NE Andapa, 9.6 km 327° NNW Manantenina, -14.435, 49.76, 775 m, rainforest, ex rotten log, 16 Nov 2003 (B.L. Fisher et al.) collection code: BLF09010, specimen code: CASENT0487718 (CAS).
</paragraph>
</subSubSection>
<subSubSection id="0FEE9A807A8D5FD2FBB71FA770806603" pageId="0" pageNumber="1" type="paratypes">
<paragraph id="FAF944978CD4E1A533710DCEE3155128" pageId="0" pageNumber="1">Paratypes.</paragraph>
<paragraph id="6D0F76A67809508C1B1625212C3CF0C9" pageId="0" pageNumber="1">3 minor workers of same data as holotype but specimens coded as: CASENT0837637, CASENT0837636, CASENT0837635 (NHMUK, PBZT, CAS).</paragraph>
</subSubSection>
<subSubSection id="3D98CE04B11E48626702B288BFC753A6" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="F1B1E8360C101A9D06AB1470DFE0E90B" pageId="0" pageNumber="1">Additional material examined.</paragraph>
<paragraph id="3E2303B507C000BCADAE217D972101FB" pageId="0" pageNumber="1">
<materialsCitation id="8DB2AA28F8C39D4054CAABBEFB406E5D" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="B. L. Fisher, PN Marojejy" country="Madagascar" county="Makirovana forest" elevation="715" latitude="-14.10295" location="Manantenina River" longLatPrecision="1" longitude="50.01984" municipality="Makirovana forest" specimenCount="1" stateProvince="Antsiranana">
<emphasis id="4BD5AC51CEBCA74EE6E9734815B1DB3B" bold="true" pageId="0" pageNumber="1">
<collectingCountry id="322EEE2DFE9AA851CB6CD9EAD7AF4DC9" name="Madagascar">Madagascar</collectingCountry>
:
<collectingRegion id="7F94CE9930AC7568C92A08DEB92A5AB1" country="Madagascar" name="Antsiranana">Antsiranana</collectingRegion>
</emphasis>
:
<collectingCounty id="50B812D72720423718F5E1000482D9D7">Makirovana forest</collectingCounty>
, -14.16666, 49.95,
<quantity id="C6BC4B67A102BAC993CF13B1084E35F2" metricMagnitude="2" metricUnit="m" metricValue="7.15" unit="m" value="715.0">
<elevation id="3A2D1DD3B8F10F98CDBEE067F05600EE" metricMagnitude="2" metricUnit="m" metricValue="7.15" unit="m" value="715.0">715 m</elevation>
</quantity>
, rainforest (
<collectorName id="006911E30443A22B7AC7ED97163CE42D">B.L. Fisher</collectorName>
et al.) (CAS);
<collectingMunicipality id="61A38F4E11C6D557FE30D2DAA9306BA3">Makirovana forest</collectingMunicipality>
,
<geoCoordinate id="ACC484F102DA8043B01C09F54F6569A5" degrees="14.10295" direction="south" orientation="latitude" precision="1" value="-14.10295">-14.10295</geoCoordinate>
,
<geoCoordinate id="8239FBD9A92CA798041BC67B122C6419" degrees="50.01984" direction="east" orientation="longitude" precision="1" value="50.01984">50.01984</geoCoordinate>
,
<quantity id="EE714C835B85F613D06B5BDD2DAE9B1E" metricMagnitude="2" metricUnit="m" metricValue="3.9" unit="m" value="390.0">
<elevation id="B669B46672B9497C6D42C4F2DB9AF611" metricMagnitude="2" metricUnit="m" metricValue="3.9" unit="m" value="390.0">390 m</elevation>
</quantity>
, rainforest (
<collectorName id="796E715F52D6C09212EDFFBD76D5BFB3">B.L. Fisher</collectorName>
et al.) (CAS);
<collectorName id="A57FCAB6D74D8F358EE4AAE382E8BAA0">PN Marojejy</collectorName>
,
<location id="501C61627AC43920AADFF347C729287D" LSID="urn:lsid:plazi:treatment:E59160DA8FA65AB8B878429A200E7150:501C61627AC43920AADFF347C729287D" country="Madagascar" county="Makirovana forest" latitude="-14.10295" longLatPrecision="1" longitude="50.01984" municipality="Makirovana forest" name="Manantenina River" stateProvince="Antsiranana">Manantenina River</location>
,
<locationDeviation id="DDBE7BC333F6D564FD77FBB7CD4CC246" location="Andapa">
<quantity id="26F99C37DC08483E809ED0F2BF424883" metricMagnitude="4" metricUnit="m" metricValue="2.76" unit="km" value="27.6">27.6 km</quantity>
35° NE Andapa
</locationDeviation>
,
<locationDeviation id="79DC17DE36BFCAD4BFFD2E71215B8B48" location="Manantenina">
<quantity id="F7A2800D6974F2E29B06A4D78688D4E3" metricMagnitude="3" metricUnit="m" metricValue="9.6" unit="km" value="9.6">9.6 km</quantity>
327° NNW Manantenina
</locationDeviation>
, -14.435, 49.76,
<quantity id="E3EB47149829E8B279F4E272E2163EAE" metricMagnitude="2" metricUnit="m" metricValue="7.75" unit="m" value="775.0">
<elevation id="BBF28DEB09DE4AEF20B7E27FBAF5B103" metricMagnitude="2" metricUnit="m" metricValue="7.75" unit="m" value="775.0">775 m</elevation>
</quantity>
, rainforest (
<collectorName id="82AC34230A4771313471DE0DA88849E0">B.L. Fisher</collectorName>
) (CAS)
</materialsCitation>
.
<materialsCitation id="3C5D06F2AB2E5F324AF1AD3BCDB84FC0" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="R. Harin'Hala, M. E. Irwin" country="Madagascar" county="Foret d'Ambalagoavy Nord" elevation="625" latitude="-21.857067" location="Ambatombe" longLatPrecision="1" longitude="47.37849" municipality="Ikongo" specimenCount="1" stateProvince="Fianarantsoa">
<emphasis id="793FB4D7947D338A39EAF8E03EFA70AB" bold="true" pageId="0" pageNumber="1">
<collectingRegion id="2A35525EFC6A56EBAB9103A81123504E" country="Madagascar" name="Fianarantsoa">Fianarantsoa</collectingRegion>
</emphasis>
:
<collectingCounty id="4D1448A83E1E75DC6C80B5B73B99B166">
<normalizedToken id="D0E06FD383CD4109050CEA88E4063616" originalValue="Forêt">Foret</normalizedToken>
<normalizedToken id="C822C373DC4517D0F333322229F04E69" originalValue="d’Ambalagoavy">d'Ambalagoavy</normalizedToken>
Nord
</collectingCounty>
,
<collectingMunicipality id="3662BAADA9974C584572AA2A4A20ED5A">Ikongo</collectingMunicipality>
,
<location id="7A70946CC7D53B4A670D4B8D2504B5BD" LSID="urn:lsid:plazi:treatment:E59160DA8FA65AB8B878429A200E7150:7A70946CC7D53B4A670D4B8D2504B5BD" country="Madagascar" county="Foret d'Ambalagoavy Nord" latitude="-21.857067" longLatPrecision="1" longitude="47.37849" municipality="Ikongo" name="Ambatombe" stateProvince="Fianarantsoa">Ambatombe</location>
,
<geoCoordinate id="873A2E6DEAFBBAFA35A754CFBF3574DE" degrees="21.857068" direction="south" orientation="latitude" precision="1" value="-21.857067">-21.857068</geoCoordinate>
,
<geoCoordinate id="01E94EA605C462D6FA0C470EF544B471" degrees="47.37849" direction="east" orientation="longitude" precision="1" value="47.37849">47.37849</geoCoordinate>
,
<quantity id="92BBB7293EB2CA4C18F89ECFC1A850B9" metricMagnitude="2" metricUnit="m" metricValue="6.25" unit="m" value="625.0">
<elevation id="8E5B22B2F49869A041C5AAB49E690349" metricMagnitude="2" metricUnit="m" metricValue="6.25" unit="m" value="625.0">625 m</elevation>
</quantity>
(
<collectorName id="7EDEC79ACBCE5AFEE553E9CCB5CB317F">
R.
<normalizedToken id="7126448A5FE9DE583801CC977D5C09C1" originalValue="Harin’Hala">Harin'Hala</normalizedToken>
</collectorName>
&amp;
<collectorName id="8FD3814B94AC87B051E47D917BD6FB30">M.E. Irwin</collectorName>
) (CAS)
</materialsCitation>
.
<materialsCitation id="55DD323AA92BD534A2FF8F070D53F212" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="RS Ambatovaky, B. L. Fisher" country="Madagascar" elevation="430" latitude="-16.7702" location="Sandrangato river" longLatPrecision="5" longitude="49.26638" municipality="Sandrangato river" specimenCount="1" stateProvince="Toamasina">
<emphasis id="8ABE8A5A8BD2B994785114062B370F28" bold="true" pageId="0" pageNumber="1">
<collectingRegion id="C7B89FE29CBC2039764BC484F1180144" country="Madagascar" name="Toamasina">Toamasina</collectingRegion>
</emphasis>
:
<collectorName id="099D5C84BC065AC218695FD8ED37A207">RS Ambatovaky</collectorName>
,
<collectingMunicipality id="6893BEF2D6CC49F9D66520750A526C88">Sandrangato river</collectingMunicipality>
,
<geoCoordinate id="D2247A509D5130192AE6D0F69A29DF80" degrees="16.7755" direction="south" orientation="latitude" precision="5" value="-16.7755">-16.7755</geoCoordinate>
,
<geoCoordinate id="E6391910F7201D5F3EDCF18F6A2AB9AB" degrees="49.26427" direction="east" orientation="longitude" precision="1" value="49.26427">49.26427</geoCoordinate>
,
<quantity id="D3EDA3B37E0F12DA6906ADF922AC1ED7" metricMagnitude="2" metricUnit="m" metricValue="4.3" unit="m" value="430.0">
<elevation id="33E3572F45839A9820C10B901A1709B7" metricMagnitude="2" metricUnit="m" metricValue="4.3" unit="m" value="430.0">430 m</elevation>
</quantity>
, rainforest (
<collectorName id="4DD7BCE825E8393BD140E48F025935ED">B.L. Fisher</collectorName>
et al.) (CAS)
<collectorName id="CECE36FAF8B8090D96FD6CC9BB1C6159">RS Ambatovaky</collectorName>
,
<location id="EA68BE2298368E5E25948A460584FC24" LSID="urn:lsid:plazi:treatment:E59160DA8FA65AB8B878429A200E7150:EA68BE2298368E5E25948A460584FC24" country="Madagascar" latitude="-16.7702" longLatPrecision="5" longitude="49.26638" municipality="Sandrangato river" name="Sandrangato river" stateProvince="Toamasina">Sandrangato river</location>
,
<geoCoordinate id="1EDF5415AD993B8BBBE72D0EC0F0E728" degrees="16.7702" direction="south" orientation="latitude" precision="5" value="-16.7702">-16.7702</geoCoordinate>
,
<geoCoordinate id="0532CCA09D54BF373764DD48F8EC08DB" degrees="49.26638" direction="east" orientation="longitude" precision="1" value="49.26638">49.26638</geoCoordinate>
,
<quantity id="051B345F88290D3D11D91723966EBD36" metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">
<elevation id="F1768F04E99615BDDE61EA77FA05DDCC" metricMagnitude="2" metricUnit="m" metricValue="4.7" unit="m" value="470.0">470 m</elevation>
</quantity>
, rainforest (
<collectorName id="C808AE520BFF5E10F79B0E6B661209D2">B.L. Fisher</collectorName>
et al.) (CAS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="630388D21AB8F3EB4199EDDB7224E285" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="855B7B5F18A781F140C340C858784F30" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="974BF11C4880049A33252E4829B4AE1F" pageId="0" pageNumber="1">In full-face view, lateral margins of head anterior to eye level diverging posteriorly; anterior clypeal margin truncate; dorsum of second and third abdominal tergites lacking white spot; propodeal dorsum straight; body color dark brown; antennal scape without erect hairs.</paragraph>
</subSubSection>
<subSubSection id="A104947CEF877C2ED9F5C9E72A235D20" pageId="0" pageNumber="1" type="description">
<paragraph id="FFAA04C722C0E15A8A6B844C46C8393B" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="B674ACBBF4A2FBD5CA4EA48388A6C45F" pageId="0" pageNumber="1">
<emphasis id="15BB0B3FC889E0A91A6844A4D33936EE" bold="true" pageId="0" pageNumber="1">Minor worker.</emphasis>
In full-face view, head sides diverging towards anterior to level of eye, evenly rounding to approximately straight posterior margin; head widest at eye level; eyes almost flattened and small (EL/CS: 0.19
<normalizedToken id="638D2A305FEF776F06F2E91B98E6AFEF" originalValue="±">+/-</normalizedToken>
0.01; 0.16-0.21), not breaking lateral cephalic margin, level of posterior margin located approximately at posterior 1/3 of head (PoOc/CL: 0.28
<normalizedToken id="330BDF2DDEAC77BB02BDF297EDEAA367" originalValue="±">+/-</normalizedToken>
0.01; 0.27-0.29); frontal carinae more or less wide (FR/CS: 0.31
<normalizedToken id="E337CD4D75D54EA87C54E2A51E36DB0E" originalValue="±">+/-</normalizedToken>
0.02; 0.28-0.34), posteriorly diverging, distance between them smaller than their smallest distance to eye; clypeus with anterolateral angle and approximately straight anteromedian margin; mandible with two apical teeth distantly spaced; antennal scape relatively short (SL/CS: 1.19
<normalizedToken id="A36F72E8C1E925CD413ACE4E418DFEEB" originalValue="±">+/-</normalizedToken>
0.05; 1.11-1.27). Promesonotum weakly convex; mesonotum with posterior portion flat immediately anterior to weakly visible metanotal groove; propodeal dorsum almost straight, its angle to declivity widely rounded; propodeal dorsum ca. 2
<normalizedToken id="7F8E69D66EEEFE37777854F24DF0656B" originalValue="×">x</normalizedToken>
as long as declivity. Petiolar node flattened anteroposteriorly, without obvious dorsal margin; anterior face almost the same height as posterior face; femur of hind leg rounded axially, not twisted basally.
</paragraph>
<paragraph id="D220B4DCD1C20B2E32A291BD46A45E58" pageId="0" pageNumber="1">First and second gastral tergites without a pair of white spots; lateral cephalic margin with erect hairs anterior to level of eyes; no erect hairs on lateral margin of head posterior to eye level; two erect hairs present near posterior margin; antennal scape only covered with appressed hairs; pronotum with few erect hairs; mesonotum with a pair of erect hairs; two erect hairs present on anterior to posterodorsal corner of propodeum. Body color reddish brown to pale brown.</paragraph>
<paragraph id="F34AB8F3F322A1F8D4F5770181366C1A" pageId="0" pageNumber="1">
<emphasis id="B6FEC1DE0B8C9F34257C0C95072922CA" bold="true" pageId="0" pageNumber="1">Major worker.</emphasis>
Differing from minor worker in the following characters: enlarged head (CS: 2.83
<normalizedToken id="CF6CFB42822B4179B6DC91D62CCE810C" originalValue="±">+/-</normalizedToken>
0.25; 2.35-3.15; CWb/CL: 0.99
<normalizedToken id="B46E180FA387325F284FF1366285A89C" originalValue="±">+/-</normalizedToken>
0.04; 0.92-1.04) with broadly concave posterior margin; apical 1/4 of antennal scape surpassing posterior cephalic margin; robust mesosoma with separate convexity of promesonotum, lower position of metanotum and propodeum, propodeal dorsum approximately the same length as declivity surface and with rounded junction; petiolar node more flattened anteroposteriorly.
</paragraph>
</subSubSection>
<subSubSection id="A2793B6F3ADA423B27521A914ACF2111" pageId="0" pageNumber="1" type="distribution">
<paragraph id="8156077FD206665D96FBC0F82A9064B8" pageId="0" pageNumber="1">Distribution and biology.</paragraph>
<paragraph id="FE75CC81708C1921DC92DDCE3D68821C" pageId="0" pageNumber="1">
The distribution of
<taxonomicName id="71C59564A0D56FECA2D1A28C562B6FE5" lsidName="C. kelimaso" pageId="0" pageNumber="1" rank="species" species="kelimaso">
<emphasis id="D89BB6454859108338FE1A8BF1CF76C6" italics="true" pageId="0" pageNumber="1">C. kelimaso</emphasis>
</taxonomicName>
is limited to the eastern lowland rainforest of Madagascar between the Makirovana forest in the north and the
<normalizedToken id="E18911BCF8BB67ADF55DD1EDB0E48669" originalValue="Forêt">Foret</normalizedToken>
<normalizedToken id="3F893BFE9B7FC14CBB532CFBC895E501" originalValue="d’Ambalagoavy">d'Ambalagoavy</normalizedToken>
Nord in the south (Fig.
<figureCitation id="4091F5ED69B39B7C524431B28BF649C3" captionStart="Figure 63" captionStartId="F63" captionText="Figure 63. Camponotus kelimaso A lateral view B head in full-face view C dorsal view of holotype minor worker CASENT 0487718 D distribution map." figureDoi="10.3897/zookeys.1098.73223.figure63" httpUri="https://binary.pensoft.net/fig/678122" pageId="0" pageNumber="1">63D</figureCitation>
). It nests mainly in rotten logs and rarely in root mat layers in the ground. Foraging is carried out on the ground and through leaf litter.
</paragraph>
<caption id="30290B21093A1E0D104F0B30A02F70B4" doi="10.3897/zookeys.1098.73223.figure63" httpUri="https://binary.pensoft.net/fig/678122" pageId="0" pageNumber="1" start="Figure 63" startId="F63">
<paragraph id="420E2B9AEDBC5EB245046EDEA6E53405" pageId="0" pageNumber="1">
<emphasis id="D394C911173DE9B786F4BF2ED93297D5" bold="true" pageId="0" pageNumber="1">Figure 63.</emphasis>
<taxonomicName id="C12B4507CF7BABE37415507125DEE36A" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus kelimaso" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kelimaso">
<emphasis id="07CA96C5E38136E00B1A95AA6EB6E035" italics="true" pageId="0" pageNumber="1">Camponotus kelimaso</emphasis>
</taxonomicName>
<emphasis id="DFC933224D759135043F39F6160D106E" bold="true" pageId="0" pageNumber="1">A</emphasis>
lateral view
<emphasis id="169153C6A46FC035F80BC69F3BB36A93" bold="true" pageId="0" pageNumber="1">B</emphasis>
head in full-face view
<emphasis id="9BB0B63378001781FC198C4173424149" bold="true" pageId="0" pageNumber="1">C</emphasis>
dorsal view of holotype minor worker CASENT0487718
<emphasis id="C464EE2B5F8F3ECD8E5C572AB28075A3" bold="true" pageId="0" pageNumber="1">D</emphasis>
distribution map.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="BCA5E52DDE92A68DF4FFC2924B043287" pageId="0" pageNumber="1" type="discussion">
<paragraph id="12932CD11E369BC0A3AFC006961C76A2" pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph id="0E6805991C7811A6C60068BF3D4C0987" pageId="0" pageNumber="1">
<taxonomicName id="0407E35A75BA5F458FD03581EADFA93E" authorityName="Rakotonirina &amp; Fisher" authorityYear="2022" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus kelimaso" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="kelimaso">
<emphasis id="5C7D8B35D3D3E88C121266F589129571" italics="true" pageId="0" pageNumber="1">Camponotus kelimaso</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="C11D3DD4AF34B7C8BBEADD608B4383D8" lsidName="C. immaculatus" pageId="0" pageNumber="1" rank="species" species="immaculatus">
<emphasis id="8826A5A33DBF902A9A8D2823F666A5FF" italics="true" pageId="0" pageNumber="1">C. immaculatus</emphasis>
</taxonomicName>
by its straight propodeal dorsum. It can be differentiated from
<taxonomicName id="5D9B3CF68739F152A3DFDCFB7B61B908" lsidName="C. lubbocki" pageId="0" pageNumber="1" rank="species" species="lubbocki">
<emphasis id="FE4412C9558DC4CF96F389A05808436E" italics="true" pageId="0" pageNumber="1">C. lubbocki</emphasis>
</taxonomicName>
by the approximately straight posterior margin of the head and its small compound eyes.
</paragraph>
<paragraph id="CF42E5A74AD781ECB1844258B3F7A6DD" pageId="0" pageNumber="1">
Based on the information provided by the NC-clustering method, the grouping of
<taxonomicName id="59D7221117589AEB22026233E6DDA07D" lsidName="C. kelimaso" pageId="0" pageNumber="1" rank="species" species="kelimaso">
<emphasis id="5FB0E0B5097D4BAF193AF3D8A2227AC8" italics="true" pageId="0" pageNumber="1">C. kelimaso</emphasis>
</taxonomicName>
supports the distinction of the species by conventional qualitative taxonomy. The confirmatory LDA identified the samples successfully at 100%.
</paragraph>
</subSubSection>
<subSubSection id="AD02011A8DB9C800CD13F3D3B94C2344" pageId="0" pageNumber="1" type="etymology">
<paragraph id="019525A63C0604FC2711B47EBA9B13DD" pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph id="144D333F1D0B92E076348FBE7ADA3A12" pageId="0" pageNumber="1">
The species name
<emphasis id="0A848701FDE4C4C632B8BE29697C8DEA" italics="true" pageId="0" pageNumber="1">kelimaso</emphasis>
is a non-latin word derived from the Malagasy word for &quot;small eye&quot;. It refers to the fact that its compound eyes are small compared to those of other species in the subgenus
<taxonomicName id="D2A98FAE0134747CDDAF67A765B23AB7" class="Insecta" family="Formicidae" genus="Camponotus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Camponotus (Myrmosaga)" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Myrmosaga">Camponotus Myrmosaga</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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