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<document id="B5315B9966ED5DE78967AFAE046244FB" ID-DOI="10.5852/ejt.2024.923.2445" ID-GBIF-Dataset="01232941-b1fd-4c18-b269-e97169b18908" ID-ISSN="2118-9773" ID-Zenodo-Dep="10727741" ID-ZooBank="AD68CEDD-2F2D-4010-BE7A-1B1AE9E4A0F3" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatmentCitations_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1709215085450" checkinUser="plazi" docAuthor="Herbert, David G." docDate="2024" docId="E54587AB386B0B24FDD7FA83569FFE00" docLanguage="en" docName="EJT.2024.923.1-119.pdf.imf" docOrigin="European Journal of Taxonomy 923" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2445/10835" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Benthobrookula araneum Herbert 2024, sp. nov." docType="treatment" docUuid="51E19EE6-CE0C-4554-BE8A-10311EAE3267" docUuidSource="ZooBank" docVersion="9" lastPageNumber="57" masterDocId="197CFFD3385E0B1CFF9BFFC25776FFD3" masterDocTitle="The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species" masterLastPageNumber="119" masterPageNumber="1" pageNumber="54" updateTime="1732578736835" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D364287D222CD1AADE958D16A0355526">The Vetigastropoda (Mollusca) of Walters Shoal, with descriptions of two new genera and thirty new species</mods:title>
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<mods:name id="246B52C7CBA779AB0820300CAF12BBBA" type="personal">
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<mods:namePart id="74C39708127A75107AB481EFAA934056">Herbert, David G.</mods:namePart>
<mods:nameIdentifier id="DFFC066CE0CA5BFDB11A138BB8ECA25D" type="LSID">0C09EE45-6198-482E-857A-EF690C2A016F</mods:nameIdentifier>
<mods:affiliation id="5522205DA7EF55384CD323D42091FCD2">Department of Natural Sciences, National Museum Wales, Cathays Park, Cardiff, CF 10 3 NP, UK. Institute of Systematics, Evolution, Biodiversity (ISYEB), Muséum national dhistoire naturelle, CNRS, SU, EPHE, UA, CP 51, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
<mods:nameIdentifier id="B14B5071E315D427189C365560ADF354" type="email">phasianella@gmail.com</mods:nameIdentifier>
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<mods:title id="910241FF4D39C0EF6BAA82992F2EC09A">European Journal of Taxonomy</mods:title>
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<mods:date id="D1B4EC0E2DF93BCE1239E945C4F049D9">2024</mods:date>
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<mods:number id="66E2BF6FCABF5B0C971559C174724BCB">923</mods:number>
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<mods:url id="A0F582841772C9AFB159B3C57E54928B">https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2445/10835</mods:url>
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<mods:identifier id="B713FA2D1872B40C3E67C96DA761BEBE" type="DOI">10.5852/ejt.2024.923.2445</mods:identifier>
<mods:identifier id="388CC20052EE5B9854F5326140B1B77E" type="GBIF-Dataset">01232941-b1fd-4c18-b269-e97169b18908</mods:identifier>
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<treatment id="E54587AB386B0B24FDD7FA83569FFE00" ID-DOI="http://doi.org/10.5281/zenodo.10818372" ID-GBIF-Taxon="223179292" ID-Zenodo-Dep="10818372" LSID="urn:lsid:zoobank.org:act:51E19EE6-CE0C-4554-BE8A-10311EAE3267" httpUri="http://treatment.plazi.org/id/E54587AB386B0B24FDD7FA83569FFE00" lastPageId="56" lastPageNumber="57" pageId="53" pageNumber="54" scope_class="Gastropoda" scope_family="Trochaclididae" scope_order="Seguenziida" scope_phylum="Mollusca" scope_subClass="Vetigastropoda" scope_superFamily="Seguenzioidea">
<subSubSection id="25F66536386B0B29FDD7FA83549EFA8F" box="[588,1000,1345,1372]" pageId="53" pageNumber="54" type="nomenclature">
<paragraph id="6D5336BD386B0B29FDD7FA83549EFA8F" blockId="53.[368,1219,1345,1443]" box="[588,1000,1345,1372]" pageId="53" pageNumber="54">
<heading id="361B81D1386B0B29FDD7FA83549EFA8F" box="[588,1000,1345,1372]" centered="true" fontSize="11" level="2" pageId="53" pageNumber="54" reason="2">
<taxonomicName id="AAEC4D3E386B0B29FDD7FA8354F0FA8F" ID-CoL="C6HX6" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[588,902,1345,1372]" class="Gastropoda" family="Unassigned" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="araneum" status="sp. nov." superFamily="Seguenzioidea">
<emphasis id="5F98EAAF386B0B29FDD7FA8354F0FA8F" bold="true" box="[588,902,1345,1372]" italics="true" pageId="53" pageNumber="54">Benthobrookula araneum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D4386B0B29FC16FA80549EFA8F" box="[909,1000,1346,1372]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="25F66536386B0B29FEEBFAA754E7FA70" pageId="53" pageNumber="54" type="description">
<paragraph id="6D5336BD386B0B29FEEBFAA753B5FA53" blockId="53.[368,1219,1345,1443]" box="[368,1219,1381,1408]" pageId="53" pageNumber="54">
<uri id="197D3ABF386B0B29FEEBFAA753B5FA53" box="[368,1219,1381,1408]" pageId="53" pageNumber="54">
urn:lsid:zoobank.org:act:
<uuid id="194A0C68386B0B29FD15FAA753B5FA53" box="[654,1219,1381,1408]" pageId="53" pageNumber="54">51E19EE6-CE0C-4554-BE8A-10311EAE3267</uuid>
</uri>
</paragraph>
<paragraph id="6D5336BD386B0B29FD39FA4A54E7FA70" blockId="53.[368,1219,1345,1443]" box="[674,913,1416,1443]" pageId="53" pageNumber="54">
<figureCitation id="F5D72A38386B0B29FD39FA4A5445FA71" box="[674,819,1416,1442]" captionStart="Fig" captionStartId="55.[189,232,1751,1777]" captionTargetBox="[189,1399,265,1717]" captionTargetId="figure-114@55.[189,1399,265,1717]" captionTargetPageId="55" captionText="Fig. 24. AC. Benthobrookula araneum sp. nov. AB. Holotype, height 1.2 mm (MNHN-IM-2000- 38068). C. Paratype, diameter 1.1 mm (MNHN-IM-2000-38069). DF. Benthobrookula galeneae sp. nov. DE. Holotype, height 1.25 mm (MNHN-IM-2000-38075). F. Paratype, diameter 1.2 mm (MNHN-IM-2000-38076). GI. Benthobrookula laticostata sp. nov. GH. Holotype, diameter 1.0 mm (MNHN-IM-2000-38080).I.Paratype,diameter1.0mm(MNHN-IM-2000-38082).JL.Benthobrookula scalaroides sp. nov. JK. Holotype, height 1.14 mm (MNHN-IM-2000-38084). L. Paratype, diameter 1.0 mm (MNHN-IM-2000-38085). MO. Benthobrookula semisculpta sp. nov. MN. Holotype, height 1.4 mm (MNHN-IM-2000-38088). O. Paratype, diameter 1.4 mm (MNHN-IM-2000-38090)." figureDoi="http://doi.org/10.5281/zenodo.10727808" httpUri="https://zenodo.org/record/10727808/files/figure.png" pageId="53" pageNumber="54">Figs 24AC</figureCitation>
,
<figureCitation id="F5D72A38386B0B29FCDBFA4A54E7FA70" box="[832,913,1416,1443]" captionStart="Fig" captionStartId="56.[189,232,1467,1493]" captionTargetBox="[189,1398,516,1416]" captionTargetId="figure-220@56.[189,1398,516,1419]" captionTargetPageId="56" captionText="Fig. 25. A, F. Benthobrookula araneum sp. nov. A. Protoconch (paratype, MNHN-IM-2000-38069). F. Sculptural detail at start of last adult whorl, holotype. B, G. Benthobrookula galeneae sp. nov. B. Protoconch (paratype, MNHN-IM-2000-38076). G. Sculptural detail at start of last adult whorl, holotype. C, H. Benthobrookula laticostata sp. nov. C. Protoconch, DW4899 (paratype, MNHN- IM-2000-38082. H. Sculptural detail at start of last adult whorl, holotype. D, I. Benthobrookula scalaroides sp. nov. D. Protoconch (paratype, MNHN-IM-2000-38085). I. Sculptural detail at start of last adult whorl, holotype. E. Benthobrookula semisculpta sp. nov., protoconch CP4911 (paratype, MNHN-IM-2000-38090). Scale bars=100 μm." figureDoi="http://doi.org/10.5281/zenodo.10727814" httpUri="https://zenodo.org/record/10727814/files/figure.png" pageId="53" pageNumber="54">25A, F</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="25F66536386B0B29FF26FA0B5201F94C" pageId="53" pageNumber="54" type="diagnosis">
<paragraph id="6D5336BD386B0B29FF26FA0B5640FA30" blockId="53.[189,310,1481,1507]" box="[189,310,1481,1507]" pageId="53" pageNumber="54">
<heading id="361B81D1386B0B29FF26FA0B5640FA30" bold="true" box="[189,310,1481,1507]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="5F98EAAF386B0B29FF26FA0B5640FA30" bold="true" box="[189,310,1481,1507]" pageId="53" pageNumber="54">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD386B0B29FF26FA3A5201F94C" blockId="53.[189,1399,1528,1695]" pageId="53" pageNumber="54">Shell turbiniform, H/D=0.831.0, whorls evenly rounded, broadly umbilicate; teleoconch sculptured with axial ribs, 2324 on first whorl, 2023 ribs on last adult whorl, never crowded behind outer lip; ribs prosocline, their intervals substantially wider than ribs and sculptured by fine, close-set, spiral threads; ribs less elevated with more rounded crests on base, continuing into umbilicus in some specimens; umbilical margin evenly rounded, its interior with indistinct spiral lirae; protoconch globose and exsert.</paragraph>
</subSubSection>
<subSubSection id="25F66536386B0B29FF26F904554AF8E1" pageId="53" pageNumber="54" type="etymology">
<paragraph id="6D5336BD386B0B29FF26F9045632F933" blockId="53.[189,324,1734,1760]" box="[189,324,1734,1760]" pageId="53" pageNumber="54">
<heading id="361B81D1386B0B29FF26F9045632F933" bold="true" box="[189,324,1734,1760]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="5F98EAAF386B0B29FF26F9045632F933" bold="true" box="[189,324,1734,1760]" pageId="53" pageNumber="54">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD386B0B29FF26F937554AF8E1" blockId="53.[189,1398,1781,1842]" pageId="53" pageNumber="54">
From the Latin
<taxonomicName id="AAEC4D3E386B0B29FEECF937569AF8DC" authorityName="Herbert" authorityYear="2024" box="[375,492,1781,1807]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="53" pageNumber="54" phylum="Mollusca" rank="species" species="araneum">
<emphasis id="5F98EAAF386B0B29FE1FF934569AF8DC" box="[388,492,1782,1807]" italics="true" pageId="53" pageNumber="54">araneum</emphasis>
</taxonomicName>
“a cobweb” (noun in apposition); in reference to the web-like interaction of axial cords and spiral threads.
</paragraph>
</subSubSection>
<subSubSection id="25F66536386B0B2AFF26F89B5668FD6D" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="6D5336BD386B0B29FF26F89B56DCF8A0" blockId="53.[189,426,1881,1907]" box="[189,426,1881,1907]" pageId="53" pageNumber="54">
<heading id="361B81D1386B0B29FF26F89B56DCF8A0" bold="true" box="[189,426,1881,1907]" fontSize="11" level="3" pageId="53" pageNumber="54" reason="3">
<emphasis id="5F98EAAF386B0B29FF26F89B56DCF8A0" bold="true" box="[189,426,1881,1907]" pageId="53" pageNumber="54">Material examined</emphasis>
</heading>
</paragraph>
<materialsCitation id="DD843CE0386B0B29FF71F84A5558F83B" ID-GBIF-Occurrence="4538174380" collectingDate="2017-05-03" collectionCode="MNHN" elevation="620" latitude="-33.283333" location="WALTERS SHOAL" longLatPrecision="1205" longitude="43.95" pageId="53" pageNumber="54" specimenCode="MNHN-IM-2000-38068" specimenCount="1" typeStatus="holotype">
<paragraph id="6D5336BD386B0B29FF71F84A562AF871" blockId="53.[189,1399,1928,2024]" box="[234,348,1928,1954]" pageId="53" pageNumber="54">
<emphasis id="5F98EAAF386B0B29FF71F84A562AF871" bold="true" box="[234,348,1928,1954]" pageId="53" pageNumber="54">
<typeStatus id="B257881F386B0B29FF71F84A562AF871" box="[234,348,1928,1954]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6D5336BD386B0B29FF26F8695558F83B" blockId="53.[189,1399,1928,2024]" pageId="53" pageNumber="54">
<location id="68336066386B0B29FF26F86956DAF816" LSID="urn:lsid:plazi:treatment:E54587AB386B0B24FDD7FA83569FFE00:68336066386B0B29FF26F86956DAF816" box="[189,428,1963,1989]" latitude="-33.283333" longLatPrecision="1205" longitude="43.95" name="WALTERS SHOAL" pageId="53" pageNumber="54">WALTERS SHOAL</location>
• empty shell; slopes, stn DW4887;
<geoCoordinate id="08D8507A386B0B29FCC5F86854B3F816" box="[862,965,1962,1990]" degrees="33" direction="south" minutes="17" orientation="latitude" pageId="53" pageNumber="54" precision="925" value="-33.283333">33°17 S</geoCoordinate>
,
<geoCoordinate id="08D8507A386B0B29FC48F868534BF815" box="[979,1085,1962,1990]" degrees="43" direction="east" minutes="57" orientation="longitude" pageId="53" pageNumber="54" precision="925" value="43.95">43°57 E</geoCoordinate>
; depth
<quantity id="AA149B58386B0B29FB0FF869526BF815" box="[1172,1309,1963,1990]" metricMagnitude="2" metricUnit="m" metricValue="6.195" metricValueMax="6.4" metricValueMin="5.99" pageId="53" pageNumber="54" unit="m" value="619.5" valueMax="640.0" valueMin="599.0">
<elevation id="E6C1D18E386B0B29FB0FF869526BF815" box="[1172,1309,1963,1990]" metricMagnitude="2" metricUnit="m" metricValue="6.195" metricValueMax="6.4" metricValueMin="5.99" pageId="53" pageNumber="54" unit="m" value="619.5" valueMax="640.0" valueMin="599.0">599640 m</elevation>
</quantity>
;
<date id="1952107D386B0B29FAB1F869578DF83B" pageId="53" pageNumber="54" value="2017-05-03">
<collectingDate id="0916E995386B0B29FAB1F869578DF83B" pageId="53" pageNumber="54" value="2017-05-03">3 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC6386B0B29FE9CF80C555CF83B" box="[263,554,1998,2024]" pageId="53" pageNumber="54">MNHN-IM-2000-38068</specimenCode>
.
</paragraph>
</materialsCitation>
<materialsCitation id="DD843CE038680B2AFF71FECB560EFEBB" ID-GBIF-Occurrence="4538174407" collectingDate="2017-05-03" collectionCode="MNHN" elevation="620" latitude="-33.283333" location="WALTERS SHOAL" longLatPrecision="1205" longitude="43.95" pageId="54" pageNumber="55" specimenCode="MNHN- IM-2000-38069" specimenCount="7" typeStatus="paratype">
<paragraph id="6D5336BD38680B2AFF71FECB5611FEF0" blockId="54.[189,1399,265,533]" box="[234,359,265,291]" pageId="54" pageNumber="55">
<emphasis id="5F98EAAF38680B2AFF71FECB5611FEF0" bold="true" box="[234,359,265,291]" pageId="54" pageNumber="55">
<typeStatus id="B257881F38680B2AFF71FECB5611FEF0" box="[234,359,265,291]" pageId="54" pageNumber="55" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="6D5336BD38680B2AFF26FEE95463FDC6" blockId="54.[189,1399,265,533]" pageId="54" pageNumber="55">
<location id="6833606638680B2AFF26FEE956C5FE96" LSID="urn:lsid:plazi:treatment:E54587AB386B0B24FDD7FA83569FFE00:6833606638680B2AFF26FEE956C5FE96" box="[189,435,299,325]" name="WALTERS SHOAL" pageId="54" pageNumber="55">WALTERS SHOAL</location>
<emphasis id="5F98EAAF38680B2AFE7AFEE9555AFE96" bold="true" box="[481,556,299,325]" pageId="54" pageNumber="55">slopes</emphasis>
<specimenCount id="7BEAFD3438680B2AFDCDFEEE5513FE95" box="[598,613,300,326]" pageId="54" pageNumber="55">7</specimenCount>
empty shells; same collection data as for
<typeStatus id="B257881F38680B2AFB0FFEE95388FE96" box="[1172,1278,299,325]" pageId="54" pageNumber="55">holotype</typeStatus>
;
<specimenCode id="3D4A9EC638680B2AFA88FEEE560EFEBB" pageId="54" pageNumber="55">MNHN- IM-2000-38069</specimenCode>
<materialsCitation id="DD843CE038680B2AFE03FE8C56AAFE59" ID-GBIF-Occurrence="4538174596" collectingDate="2017-05-06" collectionCode="MNHN" elevation="660" latitude="-33.15" longLatPrecision="1206" longitude="44.016666" pageId="54" pageNumber="55" specimenCode="MNHN-IM-2000-38070" specimenCount="1" typeStatus="paratype">
1 empty shell; stn DW4898;
<geoCoordinate id="08D8507A38680B2AFD67FE8E5413FEBB" box="[764,869,332,360]" degrees="33" direction="south" minutes="09" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-33.15">33°09 S</geoCoordinate>
,
<geoCoordinate id="08D8507A38680B2AFCEDFE8E5494FEBB" box="[886,994,332,360]" degrees="44" direction="east" minutes="01" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="44.016666">44°01 E</geoCoordinate>
; depth
<quantity id="AA149B5838680B2AFBA5FE8F53BCFEBB" box="[1086,1226,333,360]" metricMagnitude="2" metricUnit="m" metricValue="6.6" metricValueMax="6.68" metricValueMin="6.52" pageId="54" pageNumber="55" unit="m" value="660.0" valueMax="668.0" valueMin="652.0">
<elevation id="E6C1D18E38680B2AFBA5FE8F53BCFEBB" box="[1086,1226,333,360]" metricMagnitude="2" metricUnit="m" metricValue="6.6" metricValueMax="6.68" metricValueMin="6.52" pageId="54" pageNumber="55" unit="m" value="660.0" valueMax="668.0" valueMin="652.0">652668 m</elevation>
</quantity>
;
<date id="1952107D38680B2AFB41FE8C5204FEBB" box="[1242,1394,334,360]" pageId="54" pageNumber="55" value="2017-05-06">
<collectingDate id="0916E99538680B2AFB41FE8C5204FEBB" box="[1242,1394,334,360]" pageId="54" pageNumber="55" value="2017-05-06">6 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638680B2AFF26FEB256AAFE59" box="[189,476,368,394]" pageId="54" pageNumber="55">MNHN-IM-2000-38070</specimenCode>
</materialsCitation>
<materialsCitation id="DD843CE038680B2AFE6CFEB2555EFE7E" ID-GBIF-Occurrence="4538174603" collectingDate="2017-05-06" collectionCode="MNHN" elevation="714" latitude="-33.15" longLatPrecision="1206" longitude="44.033333" pageId="54" pageNumber="55" specimenCode="MNHN-IM-2000-38071" specimenCount="8" typeStatus="paratype">
<specimenCount id="7BEAFD3438680B2AFE6CFEB25570FE59" box="[503,518,368,394]" pageId="54" pageNumber="55">8</specimenCount>
empty shells; stn DW4899;
<geoCoordinate id="08D8507A38680B2AFCC0FEAD54B5FE59" box="[859,963,367,395]" degrees="33" direction="south" minutes="09" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-33.15">33°09 S</geoCoordinate>
,
<geoCoordinate id="08D8507A38680B2AFC4AFEAD534DFE58" box="[977,1083,367,395]" degrees="44" direction="east" minutes="02" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="44.033333">44°02 E</geoCoordinate>
; depth
<quantity id="AA149B5838680B2AFB08FEB2526AFE58" box="[1171,1308,368,395]" metricMagnitude="2" metricUnit="m" metricValue="7.135" metricValueMax="7.2" metricValueMin="7.07" pageId="54" pageNumber="55" unit="m" value="713.5" valueMax="720.0" valueMin="707.0">
<elevation id="E6C1D18E38680B2AFB08FEB2526AFE58" box="[1171,1308,368,395]" metricMagnitude="2" metricUnit="m" metricValue="7.135" metricValueMax="7.2" metricValueMin="7.07" pageId="54" pageNumber="55" unit="m" value="713.5" valueMax="720.0" valueMin="707.0">707720 m</elevation>
</quantity>
;
<date id="1952107D38680B2AFAB2FEB2578DFE7E" pageId="54" pageNumber="55" value="2017-05-06">
<collectingDate id="0916E99538680B2AFAB2FEB2578DFE7E" pageId="54" pageNumber="55" value="2017-05-06">6 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638680B2AFE92FE51555EFE7E" box="[265,552,403,429]" pageId="54" pageNumber="55">MNHN-IM-2000-38071</specimenCode>
</materialsCitation>
<materialsCitation id="DD843CE038680B2AFDDEFE5155F2FE03" ID-GBIF-Occurrence="4538174323" collectingDate="2017-05-06" collectionCode="MNHN" elevation="665" latitude="-33.166668" longLatPrecision="1206" longitude="44.016666" pageId="54" pageNumber="55" specimenCode="MNHN-IM-2000-38072" specimenCount="2" typeStatus="paratype">
<specimenCount id="7BEAFD3438680B2AFDDEFE515522FE7E" box="[581,596,403,429]" pageId="54" pageNumber="55">2</specimenCount>
empty shells; stn DW4900;
<geoCoordinate id="08D8507A38680B2AFC37FE535362FE7E" box="[940,1044,401,429]" degrees="33" direction="south" minutes="10" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-33.166668">33°10 S</geoCoordinate>
,
<geoCoordinate id="08D8507A38680B2AFBB8FE5353F9FE7E" box="[1059,1167,401,429]" degrees="44" direction="east" minutes="01" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="44.016666">44°01 E</geoCoordinate>
; depth
<quantity id="AA149B5838680B2AFB7CFE515207FE7E" box="[1255,1393,403,429]" metricMagnitude="2" metricUnit="m" metricValue="6.65" metricValueMax="6.7" metricValueMin="6.6" pageId="54" pageNumber="55" unit="m" value="665.0" valueMax="670.0" valueMin="660.0">
<elevation id="E6C1D18E38680B2AFB7CFE515207FE7E" box="[1255,1393,403,429]" metricMagnitude="2" metricUnit="m" metricValue="6.65" metricValueMax="6.7" metricValueMin="6.6" pageId="54" pageNumber="55" unit="m" value="665.0" valueMax="670.0" valueMin="660.0">660670 m</elevation>
</quantity>
;
<date id="1952107D38680B2AFF26FE775623FE03" box="[189,341,437,464]" pageId="54" pageNumber="55" value="2017-05-06">
<collectingDate id="0916E99538680B2AFF26FE775623FE03" box="[189,341,437,464]" pageId="54" pageNumber="55" value="2017-05-06">6 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638680B2AFEFEFE7455F2FE03" box="[357,644,438,464]" pageId="54" pageNumber="55">MNHN-IM-2000-38072</specimenCode>
</materialsCitation>
<materialsCitation id="DD843CE038680B2AFD38FE7455A1FE21" ID-GBIF-Occurrence="4538174593" collectingDate="2017-05-07" collectionCode="MNHN" elevation="706" latitude="-33.133335" longLatPrecision="1207" longitude="44.033333" pageId="54" pageNumber="55" specimenCode="MNHN-IM-2000-38073" specimenCount="1" typeStatus="paratype">
<specimenCount id="7BEAFD3438680B2AFD38FE7455C4FE03" box="[675,690,438,464]" pageId="54" pageNumber="55">1</specimenCount>
empty shell; stn CP4902;
<geoCoordinate id="08D8507A38680B2AFC62FE765315FE03" box="[1017,1123,436,464]" degrees="33" direction="south" minutes="08" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-33.133335">33°08 S</geoCoordinate>
,
<geoCoordinate id="08D8507A38680B2AFBE8FE765396FE03" box="[1139,1248,436,464]" degrees="44" direction="east" minutes="02" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="44.033333">44°02 E</geoCoordinate>
; depth
<quantity id="AA149B5838680B2AFAA7FE74567CFE20" metricMagnitude="2" metricUnit="m" metricValue="7.055" metricValueMax="7.109999999999999" metricValueMin="7.0" pageId="54" pageNumber="55" unit="m" value="705.5" valueMax="711.0" valueMin="700.0">
<elevation id="E6C1D18E38680B2AFAA7FE74567CFE20" metricMagnitude="2" metricUnit="m" metricValue="7.055" metricValueMax="7.109999999999999" metricValueMin="7.0" pageId="54" pageNumber="55" unit="m" value="705.5" valueMax="711.0" valueMin="700.0">700 711 m</elevation>
</quantity>
;
<date id="1952107D38680B2AFE83FE1B56DDFE21" box="[280,427,472,499]" pageId="54" pageNumber="55" value="2017-05-07">
<collectingDate id="0916E99538680B2AFE83FE1B56DDFE21" box="[280,427,472,499]" pageId="54" pageNumber="55" value="2017-05-07">7 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638680B2AFE23FE1A55A1FE21" box="[440,727,472,498]" pageId="54" pageNumber="55">MNHN-IM-2000-38073</specimenCode>
</materialsCitation>
<materialsCitation id="DD843CE038680B2AFD69FE1A5463FDC6" ID-GBIF-Occurrence="4538174518" collectingDate="2017-05-09" collectionCode="MNHN" elevation="818" latitude="-33.433334" longLatPrecision="1205" longitude="44.0" pageId="54" pageNumber="55" specimenCode="MNHN-IM-2000-38074" specimenCount="2" typeStatus="paratype">
<specimenCount id="7BEAFD3438680B2AFD69FE1A5477FE21" box="[754,769,472,498]" pageId="54" pageNumber="55">2</specimenCount>
empty shells; stn CP4906;
<geoCoordinate id="08D8507A38680B2AFBD3FE1453D9FE21" box="[1096,1199,470,498]" degrees="33" direction="south" minutes="26" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-33.433334">33°26 S</geoCoordinate>
,
<geoCoordinate id="08D8507A38680B2AFB26FE14525EFE20" box="[1213,1320,470,499]" degrees="44" direction="east" minutes="00" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="44.0">44°00 E</geoCoordinate>
; depth
<quantity id="AA149B5838680B2AFF26FE395633FDC6" box="[189,325,507,533]" metricMagnitude="2" metricUnit="m" metricValue="8.18" metricValueMax="8.37" metricValueMin="7.99" pageId="54" pageNumber="55" unit="m" value="818.0" valueMax="837.0" valueMin="799.0">
<elevation id="E6C1D18E38680B2AFF26FE395633FDC6" box="[189,325,507,533]" metricMagnitude="2" metricUnit="m" metricValue="8.18" metricValueMax="8.37" metricValueMin="7.99" pageId="54" pageNumber="55" unit="m" value="818.0" valueMax="837.0" valueMin="799.0">799837 m</elevation>
</quantity>
;
<date id="1952107D38680B2AFECAFE395694FDC6" box="[337,482,507,533]" pageId="54" pageNumber="55" value="2017-05-09">
<collectingDate id="0916E99538680B2AFECAFE395694FDC6" box="[337,482,507,533]" pageId="54" pageNumber="55" value="2017-05-09">9 May 2017</collectingDate>
</date>
;
<specimenCode id="3D4A9EC638680B2AFE74FE395467FDC6" box="[495,785,507,533]" pageId="54" pageNumber="55">MNHN-IM-2000-38074</specimenCode>
.
</materialsCitation>
</paragraph>
</materialsCitation>
<paragraph id="6D5336BD38680B2AFF71FDF956D0FD86" blockId="54.[189,1399,571,702]" box="[234,422,571,597]" pageId="54" pageNumber="55">
<emphasis id="5F98EAAF38680B2AFF71FDF956D0FD86" bold="true" box="[234,422,571,597]" pageId="54" pageNumber="55">Other material</emphasis>
</paragraph>
<paragraph id="6D5336BD38680B2AFF26FD9C5668FD6D" blockId="54.[189,1399,571,702]" pageId="54" pageNumber="55">
<materialsCitation id="DD843CE038680B2AFF26FD9C56CBFD48" ID-GBIF-Occurrence="4538174354" collectingDate="2017-05-03" collectionCode="MNHN" elevation="578" latitude="-33.283333" location="WALTERS SHOAL" longLatPrecision="1205" longitude="43.933334" pageId="54" pageNumber="55" specimenCount="4">
WALTERS SHOAL
<emphasis id="5F98EAAF38680B2AFE4BFD9C556DFDAB" bold="true" box="[464,539,606,632]" pageId="54" pageNumber="55">slopes</emphasis>
<specimenCount id="7BEAFD3438680B2AFDA1FD9C553FFDAB" box="[570,585,606,632]" pageId="54" pageNumber="55">4</specimenCount>
empty shells; stn DW4886;
<geoCoordinate id="08D8507A38680B2AFC3DFD9E5379FDAB" box="[934,1039,604,632]" degrees="33" direction="south" minutes="17" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-33.283333">33°17 S</geoCoordinate>
,
<geoCoordinate id="08D8507A38680B2AFB84FD9E53FAFDAB" box="[1055,1164,604,632]" degrees="43" direction="east" minutes="56" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="43.933334">43°56 E</geoCoordinate>
; depth
<quantity id="AA149B5838680B2AFB7DFD9C5207FDAB" box="[1254,1393,606,633]" metricMagnitude="2" metricUnit="m" metricValue="5.775" metricValueMax="5.82" metricValueMin="5.7299999999999995" pageId="54" pageNumber="55" unit="m" value="577.5" valueMax="582.0" valueMin="573.0">
<elevation id="E6C1D18E38680B2AFB7DFD9C5207FDAB" box="[1254,1393,606,633]" metricMagnitude="2" metricUnit="m" metricValue="5.775" metricValueMax="5.82" metricValueMin="5.7299999999999995" pageId="54" pageNumber="55" unit="m" value="577.5" valueMax="582.0" valueMin="573.0">573582 m</elevation>
</quantity>
;
<date id="1952107D38680B2AFF26FD435622FD48" box="[189,340,641,667]" pageId="54" pageNumber="55" value="2017-05-03">
<collectingDate id="0916E99538680B2AFF26FD435622FD48" box="[189,340,641,667]" pageId="54" pageNumber="55" value="2017-05-03">3 May 2017</collectingDate>
</date>
;
<collectionCode id="0BFDAE7838680B2AFEF8FD4356CBFD48" box="[355,445,641,667]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="54" pageNumber="55">MNHN</collectionCode>
</materialsCitation>
<materialsCitation id="DD843CE038680B2AFE47FD43566DFD6D" ID-GBIF-Occurrence="4538174667" collectingDate="2017-05-09" collectionCode="MNHN" elevation="880" latitude="-33.45" location="WALTERS SHOAL" longLatPrecision="1205" longitude="44.0" pageId="54" pageNumber="55" specimenCount="2">
<specimenCount id="7BEAFD3438680B2AFE47FD43569DFD48" box="[476,491,641,667]" pageId="54" pageNumber="55">2</specimenCount>
empty shells; stn CP4907;
<geoCoordinate id="08D8507A38680B2AFCA0FDBD54D2FD48" box="[827,932,639,667]" degrees="33" direction="south" minutes="27" orientation="latitude" pageId="54" pageNumber="55" precision="925" value="-33.45">33°27 S</geoCoordinate>
,
<geoCoordinate id="08D8507A38680B2AFC2FFDBD5356FD48" box="[948,1056,639,667]" degrees="44" direction="east" minutes="00" orientation="longitude" pageId="54" pageNumber="55" precision="925" value="44.0">44°00 E</geoCoordinate>
; depth
<quantity id="AA149B5838680B2AFBE0FD4353BDFD48" box="[1147,1227,641,667]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="54" pageNumber="55" unit="m" value="880.0">
<elevation id="E6C1D18E38680B2AFBE0FD4353BDFD48" box="[1147,1227,641,667]" metricMagnitude="2" metricUnit="m" metricValue="8.8" pageId="54" pageNumber="55" unit="m" value="880.0">880 m</elevation>
</quantity>
;
<date id="1952107D38680B2AFB40FD435204FD48" box="[1243,1394,641,667]" pageId="54" pageNumber="55" value="2017-05-09">
<collectingDate id="0916E99538680B2AFB40FD435204FD48" box="[1243,1394,641,667]" pageId="54" pageNumber="55" value="2017-05-09">9 May 2017</collectingDate>
</date>
;
<collectionCode id="0BFDAE7838680B2AFF26FD66566DFD6D" box="[189,283,676,702]" collectionName="France, Paris, Museum National d'Histoire Naturelle" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="54" pageNumber="55">MNHN</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="25F6653638680B2AFF26FD26531FFA26" pageId="54" pageNumber="55" type="description">
<paragraph id="6D5336BD38680B2AFF26FD265638FD2D" blockId="54.[189,334,740,766]" box="[189,334,740,766]" pageId="54" pageNumber="55">
<heading id="361B81D138680B2AFF26FD265638FD2D" bold="true" box="[189,334,740,766]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="5F98EAAF38680B2AFF26FD265638FD2D" bold="true" box="[189,334,740,766]" pageId="54" pageNumber="55">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38680B2AFF26FCD056CEFB3D" blockId="54.[188,1401,786,1262]" pageId="54" pageNumber="55">
<smallCapsWord id="6BB5A06138680B2AFF26FCD05677FCF8" baselines="805,806" box="[189,257,786,812]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="54" pageNumber="55">SHELL</smallCapsWord>
. Very small (height up to
<quantity id="AA149B5838680B2AFDD7FCD055CAFCFF" box="[588,700,786,812]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="54" pageNumber="55" unit="mm" value="1.23">1.23 mm</quantity>
), turbiniform; diameter ≥ height (H/D =0.831.0); whorls rounded and suture distinctly indented; periphery evenly rounded, close to mid-whorl; base rounded, broadly umbilicate; teleoconch of 2.753.0 whorls. First teleoconch whorl with 2324 axial ribs of similar strength, first one short, not extending to abapical suture; ribs more elevated and more prosocline on subsequent whorls, but not increasing in number, narrow and blade-like, their intervals progressively wider; 2023 ribs on last adult whorl, persisting to end of whorl in most specimens (though last few rather indistinct), but cease up to one quarter-whorl behind outer lip in others; never crowded behind outer lip; intervals between ribs substantially wider than ribs, particularly on final two whorls; intervals sculptured by fine, close-set, spiral threads (
<figureCitation id="F5D72A3838680B2AFD53FBE45447FB92" box="[712,817,1062,1089]" captionStart="Fig" captionStartId="56.[189,232,1467,1493]" captionTargetBox="[189,1398,516,1416]" captionTargetId="figure-220@56.[189,1398,516,1419]" captionTargetPageId="56" captionText="Fig. 25. A, F. Benthobrookula araneum sp. nov. A. Protoconch (paratype, MNHN-IM-2000-38069). F. Sculptural detail at start of last adult whorl, holotype. B, G. Benthobrookula galeneae sp. nov. B. Protoconch (paratype, MNHN-IM-2000-38076). G. Sculptural detail at start of last adult whorl, holotype. C, H. Benthobrookula laticostata sp. nov. C. Protoconch, DW4899 (paratype, MNHN- IM-2000-38082. H. Sculptural detail at start of last adult whorl, holotype. D, I. Benthobrookula scalaroides sp. nov. D. Protoconch (paratype, MNHN-IM-2000-38085). I. Sculptural detail at start of last adult whorl, holotype. E. Benthobrookula semisculpta sp. nov., protoconch CP4911 (paratype, MNHN-IM-2000-38090). Scale bars=100 μm." figureDoi="http://doi.org/10.5281/zenodo.10727814" httpUri="https://zenodo.org/record/10727814/files/figure.png" pageId="54" pageNumber="55">Fig. 25F</figureCitation>
), and even finer and more close-set axial threads (visible only under SEM); ribs continue on to base, but become less elevated, their crests rounded or even flattened, rather than sharp; ribs continue into umbilicus in some specimens, but evanesce at its margin in others; umbilicus relatively wide, margin evenly rounded; interior of umbilicus variably sculptured with fine, rather indistinct spiral lirae. Aperture circular; peristome complete, its edge simple; interior not nacreous.
</paragraph>
<paragraph id="6D5336BD38680B2AFF26FAD756BFFAA0" blockId="54.[189,1399,1300,1396]" pageId="54" pageNumber="55">
<smallCapsWord id="6BB5A06138680B2AFF26FAD7563AFAFE" baselines="1320,1320" box="[189,332,1301,1327]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Protoconch" pageId="54" pageNumber="55">PROTOCONCH</smallCapsWord>
(
<figureCitation id="F5D72A3838680B2AFEC4FAD656BBFAFC" box="[351,461,1300,1327]" captionStart="Fig" captionStartId="56.[189,232,1467,1493]" captionTargetBox="[189,1398,516,1416]" captionTargetId="figure-220@56.[189,1398,516,1419]" captionTargetPageId="56" captionText="Fig. 25. A, F. Benthobrookula araneum sp. nov. A. Protoconch (paratype, MNHN-IM-2000-38069). F. Sculptural detail at start of last adult whorl, holotype. B, G. Benthobrookula galeneae sp. nov. B. Protoconch (paratype, MNHN-IM-2000-38076). G. Sculptural detail at start of last adult whorl, holotype. C, H. Benthobrookula laticostata sp. nov. C. Protoconch, DW4899 (paratype, MNHN- IM-2000-38082. H. Sculptural detail at start of last adult whorl, holotype. D, I. Benthobrookula scalaroides sp. nov. D. Protoconch (paratype, MNHN-IM-2000-38085). I. Sculptural detail at start of last adult whorl, holotype. E. Benthobrookula semisculpta sp. nov., protoconch CP4911 (paratype, MNHN-IM-2000-38090). Scale bars=100 μm." figureDoi="http://doi.org/10.5281/zenodo.10727814" httpUri="https://zenodo.org/record/10727814/files/figure.png" pageId="54" pageNumber="55">Fig. 25A</figureCitation>
). Globose and exsert; diameter ±220 μm, ca 1.25 whorls; sculptured with fine, anastomosing threads creating an irregular pattern of microscopic pits; terminal lip straight or weakly convex, not thickened.
</paragraph>
<paragraph id="6D5336BD38680B2AFF26FA585493FA67" blockId="54.[189,997,1434,1461]" box="[189,997,1434,1461]" pageId="54" pageNumber="55">
<smallCapsWord id="6BB5A06138680B2AFF26FA58566EFA60" baselines="1453,1454" box="[189,280,1434,1460]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Colour" pageId="54" pageNumber="55">COLOUR</smallCapsWord>
. Uniformly white; freshest specimens somewhat translucent.
</paragraph>
<paragraph id="6D5336BD38680B2AFF26FA1E531FFA26" blockId="54.[189,1129,1499,1526]" box="[189,1129,1499,1526]" pageId="54" pageNumber="55">
<smallCapsWord id="6BB5A06138680B2AFF26FA1E5634FA27" baselines="1519,1519" box="[189,322,1500,1526]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Dimensions" pageId="54" pageNumber="55">DIMENSIONS</smallCapsWord>
.
<typeStatus id="B257881F38680B2AFECAFA1956B7FA26" box="[337,449,1499,1525]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
, height
<quantity id="AA149B5838680B2AFD86FA1955FFFA25" box="[541,649,1499,1526]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="54" pageNumber="55" unit="mm" value="1.23">1.23 mm</quantity>
, diameter
<quantity id="AA149B5838680B2AFC99FA19541DFA25" box="[770,875,1499,1526]" metricMagnitude="-3" metricUnit="m" metricValue="1.38" pageId="54" pageNumber="55" unit="mm" value="1.38">1.38 mm</quantity>
(= largest specimen).
</paragraph>
</subSubSection>
<subSubSection id="25F6653638680B2AFF26F9DE53FAF955" pageId="54" pageNumber="55" type="distribution">
<paragraph id="6D5336BD38680B2AFF26F9DE5620F9E5" blockId="54.[189,342,1564,1590]" box="[189,342,1564,1590]" pageId="54" pageNumber="55">
<heading id="361B81D138680B2AFF26F9DE5620F9E5" bold="true" box="[189,342,1564,1590]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="5F98EAAF38680B2AFF26F9DE5620F9E5" bold="true" box="[189,342,1564,1590]" pageId="54" pageNumber="55">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38680B2AFF26F98853FAF955" blockId="54.[189,1398,1610,1671]" pageId="54" pageNumber="55">
Known only from the slopes of Walters Shoal, at depths of
<quantity id="AA149B5838680B2AFCE2F9885377F9B7" box="[889,1025,1610,1637]" metricMagnitude="2" metricUnit="m" metricValue="7.31" metricValueMax="8.8" metricValueMin="5.82" pageId="54" pageNumber="55" unit="m" value="731.0" valueMax="880.0" valueMin="582.0">582880 m</quantity>
; on coarse sandy substrata with abundant crustaceans, echinoderms and solitary corals; living specimens unknown.
</paragraph>
</subSubSection>
<subSubSection id="25F6653638680B24FF26F96F569FFE00" lastPageId="56" lastPageNumber="57" pageId="54" pageNumber="55" type="discussion">
<paragraph id="6D5336BD38680B2AFF26F96F5659F914" blockId="54.[189,303,1709,1735]" box="[189,303,1709,1735]" pageId="54" pageNumber="55">
<heading id="361B81D138680B2AFF26F96F5659F914" bold="true" box="[189,303,1709,1735]" fontSize="11" level="3" pageId="54" pageNumber="55" reason="3">
<emphasis id="5F98EAAF38680B2AFF26F96F5659F914" bold="true" box="[189,303,1709,1735]" pageId="54" pageNumber="55">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="6D5336BD38680B2AFF26F9195530F83A" blockId="54.[189,1399,1755,2025]" pageId="54" pageNumber="55">
<taxonomicName id="AAEC4D3E38680B2AFF26F91955DBF925" authority="(Thiele, 1925)" baseAuthorityName="Thiele" baseAuthorityYear="1925" box="[189,685,1755,1782]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="aethiopica">
<emphasis id="5F98EAAF38680B2AFF26F919568BF926" box="[189,509,1755,1781]" italics="true" pageId="54" pageNumber="55">Benthobrookula aethiopica</emphasis>
(
<bibRefCitation id="097D4B4C38680B2AFD96F91955D3F925" author="Thiele J." box="[525,677,1755,1782]" pageId="54" pageNumber="55" pagination="35 - 382" refId="ref63529" refString="Thiele J. 1925. Gastropoda der deutschen Tiefsee-Expedition, II. Wissenschaftliche Ergebnisse der deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot;, 1898 - 1899, 17 (2): 35 - 382 [dual pagination; also numbered 1 - 348]. Gustav Fischer, Jena." type="journal article" year="1925">Thiele, 1925</bibRefCitation>
)
</taxonomicName>
from
<quantity id="AA149B5838680B2AFD6CF9195426F925" box="[759,848,1755,1782]" metricMagnitude="3" metricUnit="m" metricValue="1.134" pageId="54" pageNumber="55" unit="m" value="1134.0">1134 m</quantity>
off
<collectingCountry id="15FB762D38680B2AFC1AF9195492F926" box="[897,996,1755,1781]" name="Somalia" pageId="54" pageNumber="55">Somalia</collectingCountry>
, is of similar size and proportions (H/D ±1.0), but has 28 axial ribs on the last adult whorl and reportedly lacks spiral sculpture.
<taxonomicName id="AAEC4D3E38680B2AFF26F8E255D0F8E8" authority="(Thiele, 1925)" baseAuthorityName="Thiele" baseAuthorityYear="1925" box="[189,678,1824,1851]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="valdiviae">
<emphasis id="5F98EAAF38680B2AFF26F8E25686F8E9" box="[189,496,1824,1850]" italics="true" pageId="54" pageNumber="55">Benthobrookula valdiviae</emphasis>
(
<bibRefCitation id="097D4B4C38680B2AFD98F8E255E8F8E8" author="Thiele J." box="[515,670,1824,1851]" pageId="54" pageNumber="55" pagination="35 - 382" refId="ref63529" refString="Thiele J. 1925. Gastropoda der deutschen Tiefsee-Expedition, II. Wissenschaftliche Ergebnisse der deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot;, 1898 - 1899, 17 (2): 35 - 382 [dual pagination; also numbered 1 - 348]. Gustav Fischer, Jena." type="journal article" year="1925">Thiele, 1925</bibRefCitation>
)
</taxonomicName>
from a depth of
<quantity id="AA149B5838680B2AFCE5F8E254BAF8E8" box="[894,972,1824,1851]" metricMagnitude="2" metricUnit="m" metricValue="6.93" pageId="54" pageNumber="55" unit="m" value="693.0">693 m</quantity>
off the Kenya-Somalia border and
<emphasis id="5F98EAAF38680B2AFF26F8865608F88E" box="[189,382,1859,1885]" italics="true" pageId="54" pageNumber="55">Be. kerguelensis</emphasis>
(
<bibRefCitation id="097D4B4C38680B2AFE14F8815551F88D" author="Thiele J." box="[399,551,1859,1886]" pageId="54" pageNumber="55" pagination="35 - 382" refId="ref63529" refString="Thiele J. 1925. Gastropoda der deutschen Tiefsee-Expedition, II. Wissenschaftliche Ergebnisse der deutschen Tiefsee-Expedition auf dem Dampfer &quot; Valdivia &quot;, 1898 - 1899, 17 (2): 35 - 382 [dual pagination; also numbered 1 - 348]. Gustav Fischer, Jena." type="journal article" year="1925">Thiele, 1925</bibRefCitation>
) from an unspecified depth in the Southern Ocean, both have a similar number of axial ribs to
<emphasis id="5F98EAAF38680B2AFE44F8A4550EF8AC" box="[479,632,1894,1919]" italics="true" pageId="54" pageNumber="55">
Be.
<taxonomicName id="AAEC4D3E38680B2AFD8BF8A4550EF8AC" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[528,632,1894,1919]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="araneum" status="sp. nov.">araneum</taxonomicName>
</emphasis>
<taxonomicNameLabel id="44AB57D438680B2AFD18F8A45595F853" box="[643,739,1894,1920]" pageId="54" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
, but both are proportionately taller (H/D ± 1.171.19), have a narrower umbilicus and reach a larger size (height
<quantity id="AA149B5838680B2AFC19F84A54A9F871" box="[898,991,1928,1954]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="54" pageNumber="55" unit="mm" value="2.1">2.1 mm</quantity>
and
<quantity id="AA149B5838680B2AFB86F84A530CF871" box="[1053,1146,1928,1954]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="54" pageNumber="55" unit="mm" value="1.6">1.6 mm</quantity>
respectively). In
<emphasis id="5F98EAAF38680B2AFACBF84B5639F817" italics="true" pageId="54" pageNumber="55">Be. kerguelensis</emphasis>
, the
<typeStatus id="B257881F38680B2AFE12F8685686F817" box="[393,496,1962,1988]" pageId="54" pageNumber="55" type="holotype">holotype</typeStatus>
of which was recently figured by
<bibRefCitation id="097D4B4C38680B2AFC1AF868537EF816" author="Engl" box="[897,1032,1962,1989]" pageId="54" pageNumber="55" refId="ref59972" refString="Engl W. 2012. Shells of Antarctica. Conchbooks, Hackenheim, Germany." type="book" year="2012">Engl (2012</bibRefCitation>
: pl. 25 fig. 1), the axial ribs are more robust than in
<emphasis id="5F98EAAF38680B2AFE32F80D5549F83B" box="[425,575,1999,2024]" italics="true" pageId="54" pageNumber="55">
Be.
<taxonomicName id="AAEC4D3E38680B2AFE4CF80D5549F83B" authorityName="Herbert" authorityYear="2024" box="[471,575,1999,2024]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="araneum">araneum</taxonomicName>
</emphasis>
.
</paragraph>
<caption id="3993663538690B2BFF26F9155275F834" ID-DOI="http://doi.org/10.5281/zenodo.10727808" ID-Zenodo-Dep="10727808" httpUri="https://zenodo.org/record/10727808/files/figure.png" pageId="55" pageNumber="56" startId="55.[189,232,1751,1777]" targetBox="[189,1399,265,1717]" targetPageId="55">
<paragraph id="6D5336BD38690B2BFF26F9155275F834" blockId="55.[189,1399,1751,2024]" pageId="55" pageNumber="56">
<emphasis id="5F98EAAF38690B2BFF26F9155629F922" bold="true" box="[189,351,1751,1777]" pageId="55" pageNumber="56">Fig. 24. AC</emphasis>
.
<taxonomicName id="AAEC4D3E38690B2BFEEAF91555E9F922" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[369,671,1751,1777]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="araneum" status="sp. nov.">
<emphasis id="5F98EAAF38690B2BFEEAF91555E9F922" box="[369,671,1751,1777]" italics="true" pageId="55" pageNumber="56">Benthobrookula araneum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D438690B2BFD31F91A547EF921" box="[682,776,1752,1778]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="5F98EAAF38690B2BFC88F915543CF922" bold="true" box="[787,842,1751,1777]" pageId="55" pageNumber="56">AB</emphasis>
. Holotype, height 1.2 mm (MNHN-IM-2000- 38068).
<emphasis id="5F98EAAF38690B2BFEBEF938564CF8C7" bold="true" box="[293,314,1786,1812]" pageId="55" pageNumber="56">C</emphasis>
. Paratype, diameter 1.1 mm (MNHN-IM-2000-38069).
<emphasis id="5F98EAAF38690B2BFC6CF938535AF8C7" bold="true" box="[1015,1068,1786,1812]" pageId="55" pageNumber="56">DF</emphasis>
.
<taxonomicName id="AAEC4D3E38690B2BFBD9F9385200F8C7" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[1090,1398,1786,1812]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="galeneae" status="sp. nov.">
<emphasis id="5F98EAAF38690B2BFBD9F9385200F8C7" box="[1090,1398,1786,1812]" italics="true" pageId="55" pageNumber="56">Benthobrookula galeneae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D438690B2BFF26F8DC566AF8EB" box="[189,284,1822,1848]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="5F98EAAF38690B2BFEB3F8DF5629F8E4" bold="true" box="[296,351,1821,1847]" pageId="55" pageNumber="56">DE</emphasis>
. Holotype, height 1.25 mm (MNHN-IM-2000-38075).
<emphasis id="5F98EAAF38690B2BFB9DF8DF536EF8E4" bold="true" box="[1030,1048,1821,1847]" pageId="55" pageNumber="56">F</emphasis>
. Paratype, diameter 1.2 mm (MNHN-IM-2000-38076).
<emphasis id="5F98EAAF38690B2BFE65F8825559F889" bold="true" box="[510,559,1856,1882]" pageId="55" pageNumber="56">GI</emphasis>
.
<taxonomicName id="AAEC4D3E38690B2BFDA5F882540BF889" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[574,893,1856,1882]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="laticostata" status="sp. nov.">
<emphasis id="5F98EAAF38690B2BFDA5F882540BF889" box="[574,893,1856,1882]" italics="true" pageId="55" pageNumber="56">Benthobrookula laticostata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D438690B2BFC1EF8835496F888" box="[901,992,1857,1883]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="5F98EAAF38690B2BFC7CF8825352F889" bold="true" box="[999,1060,1856,1882]" pageId="55" pageNumber="56">GH</emphasis>
. Holotype, diameter 1.0 mm (MNHN-IM-2000-38080).
<emphasis id="5F98EAAF38690B2BFE61F8A65573F8AD" bold="true" box="[506,517,1892,1918]" pageId="55" pageNumber="56">I</emphasis>
.Paratype,diameter1.0mm(MNHN-IM-2000-38082).
<emphasis id="5F98EAAF38690B2BFBE0F8A653DAF8AD" bold="true" box="[1147,1196,1892,1918]" pageId="55" pageNumber="56">JL</emphasis>
.
<taxonomicName id="AAEC4D3E38690B2BFB21F8A65635F872" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="scalaroides" status="sp. nov.">
<emphasis id="5F98EAAF38690B2BFB21F8A65635F872" italics="true" pageId="55" pageNumber="56">Benthobrookula scalaroides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D438690B2BFED7F84A56D1F871" box="[332,423,1928,1954]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="5F98EAAF38690B2BFE2BF8455695F872" bold="true" box="[432,483,1927,1953]" pageId="55" pageNumber="56">JK</emphasis>
. Holotype, height 1.14 mm (MNHN-IM-2000-38084).
<emphasis id="5F98EAAF38690B2BFBECF84553FFF872" bold="true" box="[1143,1161,1927,1953]" pageId="55" pageNumber="56">L</emphasis>
. Paratype, diameter 1.0 mm (MNHN-IM-2000-38085).
<emphasis id="5F98EAAF38690B2BFDC5F86855E9F817" bold="true" box="[606,671,1962,1988]" pageId="55" pageNumber="56">MO</emphasis>
.
<taxonomicName id="AAEC4D3E38690B2BFD35F868548AF817" authority="Herbert, 2024" authorityName="Herbert" authorityYear="2024" box="[686,1020,1962,1988]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="55" pageNumber="56" phylum="Mollusca" rank="species" species="semisculpta" status="sp. nov.">
<emphasis id="5F98EAAF38690B2BFD35F868548AF817" box="[686,1020,1962,1988]" italics="true" pageId="55" pageNumber="56">Benthobrookula semisculpta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44AB57D438690B2BFB99F869532BF816" box="[1026,1117,1963,1989]" pageId="55" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="5F98EAAF38690B2BFBF8F86853D5F817" bold="true" box="[1123,1187,1962,1988]" pageId="55" pageNumber="56">MN</emphasis>
. Holotype, height 1.4 mm (MNHN-IM-2000-38088).
<emphasis id="5F98EAAF38690B2BFDC4F80F5500F834" bold="true" box="[607,630,1997,2023]" pageId="55" pageNumber="56">O</emphasis>
. Paratype, diameter 1.4 mm (MNHN-IM-2000-38090).
</paragraph>
</caption>
<paragraph id="6D5336BD38660B24FF26FECB569FFE00" blockId="56.[188,1399,265,467]" pageId="56" pageNumber="57">
<taxonomicName id="AAEC4D3E38660B24FF26FECB55FDFEF7" authority="(Powell, 1951)" baseAuthorityName="Powell" baseAuthorityYear="1951" box="[189,651,265,292]" class="Gastropoda" genus="Benthobrookula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Seguenziida" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="pfefferi">
<emphasis id="5F98EAAF38660B24FF26FECB56A3FEF0" box="[189,469,265,291]" italics="true" pageId="56" pageNumber="57">Benthobrookula pfefferi</emphasis>
(Powell, 1951)
</taxonomicName>
from
<collectingCountry id="15FB762D38660B24FD48FECB540AFEF0" box="[723,892,265,291]" name="South Georgia and the South Sandwich Islands" pageId="56" pageNumber="57">South Georgia</collectingCountry>
is also somewhat similar, but it has a more elevated shell with more numerous axial ribs (3638 on last adult whorl), and attains a larger size (height
<quantity id="AA149B5838660B24FF27FE925650FEB9" box="[188,294,336,362]" metricMagnitude="-3" metricUnit="m" metricValue="1.83" pageId="56" pageNumber="57" unit="mm" value="1.83">1.83 mm</quantity>
) (
<bibRefCitation id="097D4B4C38660B24FEA6FE8D556CFEBA" author="Zelaya D. G. &amp; Absalao R. S. &amp; Pimenta A. D." box="[317,538,335,362]" pageId="56" pageNumber="57" pagination="77 - 87" refId="ref64671" refString="Zelaya D. G., Absalao R. S. &amp; Pimenta A. D. 2006. A revision of Benthobrookula Clarke, 1961 (Gastropoda, Trochoidea) in the southwestern Atlantic Ocean. Journal of Molluscan Studies 72 (1): 77 - 87. https: // doi. org / 10.1093 / mollus / eyi 050" type="journal article" year="2006">
Zelaya
<emphasis id="5F98EAAF38660B24FE03FE9256BDFEBA" box="[408,459,335,361]" italics="true" pageId="56" pageNumber="57">et al</emphasis>
. 2006
</bibRefCitation>
). In terms of overall shell shape and proportions, and the number of axial ribs, perhaps the most similar species is
<emphasis id="5F98EAAF38660B24FD2AFEB155A1FE5F" box="[689,727,371,396]" italics="true" pageId="56" pageNumber="57">Be.</emphasis>
<taxonomicName id="AAEC4D3E38660B24FD79FEB15485FE5F" authority="(Watson, 1886)" baseAuthorityName="Watson" baseAuthorityYear="1886" box="[738,1011,370,397]" class="Gastropoda" family="Fissurellidae" genus="Emarginula" kingdom="Animalia" order="Lepetellida" pageId="56" pageNumber="57" phylum="Mollusca" rank="species" species="conica">
<emphasis id="5F98EAAF38660B24FD79FEB15446FE5F" box="[738,816,371,396]" italics="true" pageId="56" pageNumber="57">conica</emphasis>
(Watson, 1886)
</taxonomicName>
from off
<collectingCountry id="15FB762D38660B24FBEBFEB053CEFE5F" box="[1136,1208,370,396]" name="Brazil" pageId="56" pageNumber="57">Brazil</collectingCountry>
(
<bibRefCitation id="097D4B4C38660B24FB50FEB0578AFE63" author="Absalao R. S. &amp; Miyaji C. &amp; Pimenta A. D." pageId="56" pageNumber="57" pagination="675 - 687" refId="ref58799" refString="Absalao R. S., Miyaji C. &amp; Pimenta A. D. 2001. The genus Brookula Iredale, 1912 (Gastropoda, Trochidae) from Brazil: description of a new species, with notes on other South American species. Zoosystema 23 (4): 675 - 687." type="journal article" year="2001">
Absalão
<emphasis id="5F98EAAF38660B24FAA3FEB15219FE5F" box="[1336,1391,370,396]" italics="true" pageId="56" pageNumber="57">et al</emphasis>
. 2001
</bibRefCitation>
), but that species has more robust axial ribs which remain strong in the umbilicus and the umbilical margin lacks spiral cords.
</paragraph>
</subSubSection>
</treatment>
</document>