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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.715.20497" ID-GBIF-Dataset="138d6982-b157-4301-a3a0-72029f9044ba" ID-PMC="PMC5740420" ID-Pensoft-Pub="1313-2970-715-69" ID-PubMed="29302233" ID-ZBK="763EF053AFAA41119CA96060F9E09E70" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-715-69" ModsDocOrigin="ZooKeys 715" ModsDocTitle="Revision of the genus Ptomaphagus Hellwig from eastern Asia (Coleoptera, Leiodidae, Cholevinae)" checkinTime="1510736981334" checkinUser="pensoft" docAuthor="Wang, Cheng-Bin, Perreau, Michel, Ruzicka, Jan &amp; Nishikawa, Masaaki" docDate="2017" docId="FF913CBEC2FDA5D6E14249DA9BE5D0EF" docLanguage="en" docName="ZooKeys 715: 69-92" docOrigin="ZooKeys 715" docSource="http://dx.doi.org/10.3897/zookeys.715.20497" docTitle="Ptomaphagus funiu Wang, Perreau, Ruzicka &amp; Nishikawa, 2017, sp. n." docType="treatment" docUuid="8DA08857-FB58-4688-8E00-38E054A82C3D" docUuidSource="ZooBank" docVersion="5" lastPageNumber="83" masterDocId="7A4E6F14FFE5FFD08743FFBA6574A77E" masterDocTitle="Revision of the genus Ptomaphagus Hellwig from eastern Asia (Coleoptera, Leiodidae, Cholevinae)" masterLastPageNumber="92" masterPageNumber="69" pageNumber="79" updateTime="1668165125337" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the genus Ptomaphagus Hellwig from eastern Asia (Coleoptera, Leiodidae, Cholevinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Cheng-Bin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Perreau, Michel</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ruzicka, Jan</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Nishikawa, Masaaki</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>715</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>69</mods:start>
<mods:end>92</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.715.20497</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.715.20497</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-715-69</mods:identifier>
<mods:identifier type="ZBK">763EF053AFAA41119CA96060F9E09E70</mods:identifier>
<mods:identifier type="ZooBank">763EF053AFAA41119CA96060F9E09E70</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="137306765" LSID="urn:lsid:zoobank.org:act:8DA08857-FB58-4688-8E00-38E054A82C3D" httpUri="http://treatment.plazi.org/id/FF913CBEC2FDA5D6E14249DA9BE5D0EF" lastPageId="14" lastPageNumber="83" pageId="10" pageNumber="79">
<subSubSection pageId="10" pageNumber="79" type="nomenclature">
<paragraph pageId="10" pageNumber="79">
<taxonomicName LSID="http://zoobank.org/8DA08857-FB58-4688-8E00-38E054A82C3D" class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus funiu" order="Coleoptera" pageId="10" pageNumber="79" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="funiu">Ptomaphagus (s. str.) funiu</taxonomicName>
<taxonomicNameLabel pageId="10" pageNumber="79">sp. n.</taxonomicNameLabel>
Figs 1
<normalizedToken originalValue="FG">F-G</normalizedToken>
; 6
<normalizedToken originalValue="AJ">A-J</normalizedToken>
; 7
<normalizedToken originalValue="AF">A-F</normalizedToken>
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="80" type="type material">
<paragraph pageId="11" pageNumber="80">
<pageBreakToken pageId="11" pageNumber="80" start="start">Type</pageBreakToken>
material.
</paragraph>
<paragraph pageId="11" pageNumber="80">
Holotype. ♂, China, W Henan, 9.-10.VI. / Funiu Shan [伏牛山],
<geoCoordinate direction="north" orientation="latitude" precision="925" value="33.516666">33°31'N</geoCoordinate>
,
<geoCoordinate direction="east" orientation="longitude" precision="925" value="111.933334">111°56'E</geoCoordinate>
/ BAOTIANMAN, 1500-1750 m / Jaroslav Turna leg., 2008 (CPMG). Paratypes. 1♀, same data as holotype (CJRZ); 1♂, same data as holotype except: 15.V.-5.VI / 2009 (CMPR).
</paragraph>
</subSubSection>
<subSubSection pageId="11" pageNumber="80" type="diagnosis">
<paragraph pageId="11" pageNumber="80">Diagnosis.</paragraph>
<paragraph pageId="11" pageNumber="80">Aedeagus (Fig. 7A) very short and stout, with median lobe gradually narrowing towards a widely lanceolate apex and terminated to a rounded knob in dorsal view. In lateral view (Fig. 7B), median lobe very thick, gently bent ventrally, and gradually tapering towards a thin apex.</paragraph>
</subSubSection>
<subSubSection lastPageId="13" lastPageNumber="82" pageId="11" pageNumber="80" type="description">
<paragraph pageId="11" pageNumber="80">Description.</paragraph>
<paragraph pageId="11" pageNumber="80">
Male. EBL: 3.9 mm. Length of different body parts: HL: AL: PL: ELL = 0.6: 1.0: 1.1: 2.1 mm; width: HW: EW: PW: ELW = 1.0: 0.1: 1.5: 1.6 mm. Proportion of antennomeres from base to tip in
<normalizedToken originalValue="μm">μm</normalizedToken>
(length
<normalizedToken originalValue="×">x</normalizedToken>
width): 154
<normalizedToken originalValue="×">x</normalizedToken>
61, 117
<normalizedToken originalValue="×">x</normalizedToken>
61, 87
<normalizedToken originalValue="×">x</normalizedToken>
65, 55
<normalizedToken originalValue="×">x</normalizedToken>
72, 57
<normalizedToken originalValue="×">x</normalizedToken>
87, 46
<normalizedToken originalValue="×">x</normalizedToken>
100, 80
<normalizedToken originalValue="×">x</normalizedToken>
117, 34
<normalizedToken originalValue="×">x</normalizedToken>
124, 77
<normalizedToken originalValue="×">x</normalizedToken>
138, 89
<normalizedToken originalValue="×">x</normalizedToken>
146, 141
<normalizedToken originalValue="×">x</normalizedToken>
131.
</paragraph>
<paragraph pageId="11" pageNumber="80">Habitus (Fig. 1F) elongated oval, regularly convex and sublustrous. Well pigmented: mostly blackish brown; mouthparts, basal three or four antennomeres and apical half of ultimate antennomere, protarsi, and apex of meso- and metatarsi more or less paler. Dorsum continually clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous.</paragraph>
<paragraph pageId="11" pageNumber="80">Head quite transverse, HW/HL = 1.7. Clypeofrontal suture absent. Clypeus with anterior margin gently rounded. Compound eyes well developed, EW/HW = 0.1. Antennae (Fig. 6A) slender, AL/HW = 1.0; antennomere III shorter than II; VI with length/width = 0.5; XI pear-shape.</paragraph>
<paragraph pageId="11" pageNumber="80">Pronotum (Fig. 6B) transverse, widest directly before hind angles, PW/PL = 1.5. Sides gently arched, narrowing from posterior to anterior; hind angles slightly projected backwards and acute. Posterior margin widely protruded in middle part, emarginate near hind angles.</paragraph>
<paragraph pageId="11" pageNumber="80">Elytra oval, widest at about basal 1/5, ELL/EW = 1.3. Sides weakly arched, gradually narrowing from widest part to apices; apices (Fig. 6G) widely rounded. Sutural striae present. Metathoracic wings fully developed.</paragraph>
<paragraph pageId="12" pageNumber="81">
<pageBreakToken pageId="12" pageNumber="81" start="start">Prolegs</pageBreakToken>
robust, with basal three protarsomeres (Fig. 6C) less expanded: TW/BTW = 1.5. Protibiae (Fig. 6E) expanded towards apex. Profemora broad. Mesotibiae arcuate, mesotarsi simply linear. Metatibiae slender and straight.
</paragraph>
<paragraph pageId="12" pageNumber="81">Abdominal ventrite VIII (Fig. 6I) simply subrounded at posterior edge. Spiculum gastrale of genital segment (Fig. 6J) with about 1/3 of length protruding beyond anterior edge of epipleurite IX.</paragraph>
<caption pageId="12" pageNumber="81">
<paragraph pageId="12" pageNumber="81">
Figure 6.
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus funiu" order="Coleoptera" pageId="12" pageNumber="81" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="funiu">Ptomaphagus (s. str.) funiu</taxonomicName>
sp. n. (♂: paratype; ♀: paratype). A antenna ♂ (dorsal view) B pronotum ♂ (dorsal view) C protarsus ♂ (dorsal view) D protarsus ♀ (dorsal view) E protibia and profemur ♂ (ventral view) F protibia and profemur ♀ (ventral view) G elytral apex ♂ (dorsoapical view) H elytral apex ♀ (dorsoapical view) I ventrite VIII ♂ (ventral view) J genital segment ♂ (ventral view). Scale bars: 0.1 mm.
</paragraph>
</caption>
<paragraph lastPageId="13" lastPageNumber="82" pageId="12" pageNumber="81">
Aedeagus (Fig. 7A) very short and stout, with median lobe gradually narrowing towards a widely lanceolate apex and terminated to a rounded knob in dorsal view;
<pageBreakToken pageId="13" pageNumber="82" start="start">opening</pageBreakToken>
of genital orifice situated on dorsal surface, deeply cut inwards on preapical left margin of median lobe. Ventral surface of the apex of the median lobe (Fig. 7C) inserted with 5 ventrally oriented setae on the left side and 6 ventrally oriented setae on the right side; parameres narrow, reaching about apical 1/5 of median lobe, each apex (Fig. 7D) with 2 lateral setae and 1 similar apical seta. In lateral view (Fig. 7B), median lobe very thick, gently bent ventrally, and gradually tapering towards a thin apex. Endophallus with stylus quite slender, a subelliptical nodule in middle region, a cheliform complex just below base of stylus, and a circular complex in the basal region.
</paragraph>
<paragraph pageId="13" pageNumber="82">Female. Similar to male in general appearance (Fig. 1G), including elytral apices (Fig. 6H), but distinguished by the following characteristics: protarsi (Fig. 6D) simply linear; protibiae (Fig. 6F) slightly narrower; abdominal ventrite VIII (Fig. 7E) round at posterior edge; genital segment and ovipositor as shown in Fig. 7F; spermatheca (Fig. 7F) curved in distal part, coiled in proximal part, and stem slightly arcuate.</paragraph>
<caption pageId="13" pageNumber="82">
<paragraph pageId="13" pageNumber="82">
Figure 7.
<taxonomicName class="Insecta" family="Leiodidae" genus="Ptomaphagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ptomaphagus funiu" order="Coleoptera" pageId="13" pageNumber="82" phylum="Arthropoda" rank="species" sensu="(s. str.)" species="funiu">Ptomaphagus (s. str.) funiu</taxonomicName>
sp. n. (♂: paratype; ♀: paratype). A aedeagus (dorsal view) B aedeagus (lateral view) C aedeagal apex (ventral view) D paramere apex (lateral view) E ventrite VIII ♀ (ventral view) F spermatheca, genital segment and ovipositor (ventral view). Scale bars: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="14" pageNumber="83" type="distribution">
<paragraph pageId="14" pageNumber="83">
<pageBreakToken pageId="14" pageNumber="83" start="start">Distribution</pageBreakToken>
.
</paragraph>
<paragraph pageId="14" pageNumber="83">China (Funiu Mts. in Henan).</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="83" type="etymology">
<paragraph pageId="14" pageNumber="83">Etymology.</paragraph>
<paragraph pageId="14" pageNumber="83">The specific epithet is from the Chinese name (in Pinyin) of the type locality &quot;Funiu Shan&quot;, and means &quot;prostrate cow&quot;.</paragraph>
</subSubSection>
</treatment>
</document>