treatments-xml/data/FE/18/87/FE18879D9B1D8313FF2BF9C2FAB9FC7A.xml
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<mods:title id="5BE4BF97BEF8DB7945447A5B00624880">New species and records in Phrynidius Lacordaire (Coleoptera, Cerambycidae, Lamiinae)</mods:title>
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<emphasis id="44C5EA999B1D8310FF2BF9C2FE52F98E" bold="true" box="[151,387,1590,1617]" italics="true" pageId="20" pageNumber="471">Phrynidius echinus</emphasis>
<bibRefCitation id="12204B7A9B1D8310FE36F9C3FDC9F98F" author="Bates, H. W." box="[394,536,1590,1617]" pageId="20" pageNumber="471" refId="ref16372" refString="Bates, H. W. (1880) Longicornia, Fam. Cerambycidae [continuation]. In: Godman, F. D., Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Taylor and Francis, London, pp. 17 - 152." type="book" year="1880">Bates, 1880</bibRefCitation>
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(
<figureCitation id="EE8A2A0E9B1D8310FF23F9AFFECEF9AA" box="[159,287,1626,1653]" captionStart-0="FIGURES 4651" captionStart-1="FIGURES 5256. 5254" captionStartId-0="21.[152,270,1943,1967]" captionStartId-1="22.[152,270,1506,1530]" captionTargetBox-0="[181,1406,181,1919]" captionTargetBox-1="[161,1436,186,1476]" captionTargetId-0="figure-17@21.[181,1406,181,1919]" captionTargetId-1="figure-26@22.[151,1436,181,1481]" captionTargetPageId-0="21" captionTargetPageId-1="22" captionText-0="FIGURES 4651. Phrynidius echinus Bates, 1880. 4649) Male: 46) Dorsal habitus; 47) Ventral habitus; 48) Lateral habitus; 49) Head, frontal view. 5051) Female: 50) Dorsal habitus; 51) Ventral habitus." captionText-1="FIGURES 5256. 5254) Phrynidius echinus Bates, 1880, holotype: 52) Dorsal habitus; 53) Lateral habitus; 54) Labels. 5556) Phrynidius armatus Linsley, 1933, holotype male: 55) Dorsal habitus; 56) Lateral habitus. Figures 5254 by Keita Matsumoto (BMNH); 5556 by Ranchel Diaz-Bastin (CASC)." figureDoi-0="http://doi.org/10.5281/zenodo.8212338" figureDoi-1="http://doi.org/10.5281/zenodo.8212340" httpUri-0="https://zenodo.org/record/8212338/files/figure.png" httpUri-1="https://zenodo.org/record/8212340/files/figure.png" pageId="20" pageNumber="471">Figs 4656</figureCitation>
)
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<emphasis id="44C5EA999B1D8310FF2BF956FE51F963" bold="true" box="[151,384,1698,1724]" pageId="20" pageNumber="471">Material examined.</emphasis>
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,
<emphasis id="44C5EA999B1D8310FDFFF957FD08F963" box="[579,729,1699,1724]" italics="true" pageId="20" pageNumber="471">
<collectingRegion id="B475F8699B1D8310FDFFF957FD08F963" box="[579,729,1699,1724]" country="Guatemala" name="Baja Verapaz" pageId="20" pageNumber="471">Baja Verapaz</collectingRegion>
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:
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S
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,
<collectorName id="DB44535D9B1D8310FB62F957FA4DF962" box="[1246,1436,1699,1725]" pageId="20" pageNumber="471">H. &amp; A. Howden</collectorName>
leg. (
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N
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leg. (
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.
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(cat.);
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(checklist);
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(cat.);
<treatmentCitation id="F710109A9B1D8310FCCCF8C7FB03F892" author="Chemsak, J. A. &amp; Linsley, E. G." box="[880,1234,1843,1869]" page="416" pageId="20" pageNumber="471" year="1970">
<bibRefCitation id="12204B7A9B1D8310FCCCF8C7FB03F892" author="Chemsak, J. A. &amp; Linsley, E. G." box="[880,1234,1843,1869]" pageId="20" pageNumber="471" pagination="404 - 417" refId="ref16666" refString="Chemsak, J. A. &amp; Linsley, E. G. (1970) Additional designations of lectotypes of neotropical Cerambycidae in the collections of the British Museum (Natural History) (Coleoptera). Journal of the Kansas Entomological Society, 43 (4), 404 - 417." type="journal article" year="1970">Chemsak &amp; Linsley, 1970: 416</bibRefCitation>
</treatmentCitation>
(lect.);
<treatmentCitation id="F710109A9B1D8310FA91F8C7FEDDF8AE" author="Breuning, S." page="325" pageId="20" pageNumber="471" year="1971">
<bibRefCitation id="12204B7A9B1D8310FA91F8C7FEDDF8AE" author="Breuning, S." pageId="20" pageNumber="471" pagination="209 - 335" refId="ref16624" refString="Breuning, S. (1971) Revision des especes americaines de la tribue des Apomecynini Lac. (Coleoptera, Cerambycidae). Entomologische Abhandlungen aus dem Staatliches Museum fur Tierkunde, 37 (3), 209 - 335." type="journal article" year="1971">Breuning, 1971: 325</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F710109A9B1D8310FEA6F8A3FDE9F8AE" author="Chemsak, J. &amp; Linsley, E. G. &amp; Mankins J. V." box="[282,568,1878,1905]" page="35" pageId="20" pageNumber="471" year="1980">
<bibRefCitation id="12204B7A9B1D8310FEA6F8A3FDE9F8AE" author="Chemsak, J. &amp; Linsley, E. G. &amp; Mankins J. V." box="[282,568,1878,1905]" pageId="20" pageNumber="471" pagination="26 - 37" refId="ref16720" refString="Chemsak, J. A, Linsley, E. G. &amp; Mankins J. V. (1980) Records of some Cerambycidae from Honduras (Coleoptera). The Pan-Pacific Entomologist, 56 (1), 26 - 37." type="journal article" year="1980">
Chemsak
<emphasis id="44C5EA999B1D8310FE36F8A3FE15F8AE" box="[394,452,1879,1905]" italics="true" pageId="20" pageNumber="471">et al.</emphasis>
, 1980: 35
</bibRefCitation>
</treatmentCitation>
(distr.);
<treatmentCitation id="F710109A9B1D8310FD26F8A3FC14F8AE" author="Chemsak, J. &amp; Linsley, E. G. &amp; Noguera, F. A." box="[666,965,1879,1905]" page="116" pageId="20" pageNumber="471" year="1992">
<bibRefCitation id="12204B7A9B1D8310FD26F8A3FC14F8AE" author="Chemsak, J. &amp; Linsley, E. G. &amp; Noguera, F. A." box="[666,965,1879,1905]" pageId="20" pageNumber="471" refId="ref16764" refString="Chemsak, J. A, Linsley, E. G. &amp; Noguera, F. A. (1992) Listados faunisticos de Mexico. II. Los Cerambycidae y Disteniidae de Norteamerica, Centroamerica y las Indias Occidentales (Coleoptera). Universidad Nacional Autonoma, Mexico City, 204 pp." type="book" year="1992">
Chemsak
<emphasis id="44C5EA999B1D8310FCB7F8A3FC94F8AE" box="[779,837,1879,1905]" italics="true" pageId="20" pageNumber="471">et al.</emphasis>
, 1992: 116
</bibRefCitation>
</treatmentCitation>
(cat.); Monné, 1994: 34 (cat.);
<treatmentCitation id="F710109A9B1D8310FA93F8A3FEA2F84A" author="Monne, M. A. &amp; Giesbert, E. F." page="189" pageId="20" pageNumber="471" year="1994">
<bibRefCitation id="12204B7A9B1D8310FA93F8A3FEA2F84A" author="Monne, M. A. &amp; Giesbert, E. F." pageId="20" pageNumber="471" refId="ref17341" refString="Monne, M. A. &amp; Giesbert, E. F. (1994) Checklist of the Cerambycidae and Disteniidae (Coleoptera) of the Western Hemisphere. Wolfsgarden Books, Burbank, 409 pp." type="book" year="1994">Monné &amp; Giesbert, 1994: 189</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="F710109A9B1D8310FE41F88FFCC1F84A" author="Turnbow, R. H. &amp; Cave, R. D. &amp; Thomas, M. C." box="[509,784,1915,1941]" page="21" pageId="20" pageNumber="471" year="2003">
<bibRefCitation id="12204B7A9B1D8310FE41F88FFCC1F84A" author="Turnbow, R. H. &amp; Cave, R. D. &amp; Thomas, M. C." box="[509,784,1915,1941]" pageId="20" pageNumber="471" pagination="1 - 43" refId="ref17732" refString="Turnbow, R. H., Cave, R. D. &amp; Thomas, M. C. (2003) A list of the Cerambycidae of Honduras, with additions of previously unrecorded species. Ceiba, 44 (1), 1 - 43." type="journal article" year="2003">
Turnbow
<emphasis id="44C5EA999B1D8310FDD5F88FFD70F84A" box="[617,673,1915,1941]" italics="true" pageId="20" pageNumber="471">et al.</emphasis>
, 2003: 21
</bibRefCitation>
</treatmentCitation>
(distr.);
<treatmentCitation id="F710109A9B1D8310FCD2F88FFBEAF84A" author="Monne, M. A." box="[878,1083,1914,1941]" page="305" pageId="20" pageNumber="471" year="2005">
<bibRefCitation id="12204B7A9B1D8310FCD2F88FFBEAF84A" author="Monne, M. A." box="[878,1083,1914,1941]" pageId="20" pageNumber="471" pagination="1 - 759" refId="ref17246" refString="Monne, M. A. (2005) Catalogue of the Cerambycidae (Coleoptera) of the Neotropical Region. Part II. Subfamily Lamiinae. Zootaxa, 1023 (1), 1 - 759. https: // doi. org / 10.11646 / zootaxa. 1023.1.1" type="journal article" year="2005">Monné, 2005: 305</bibRefCitation>
</treatmentCitation>
(cat.);
<treatmentCitation id="F710109A9B1D8310FB35F88FFF10F866" author="Monne, M. A. &amp; Hovore, F. T." page="228" pageId="20" pageNumber="471" year="2006">
<bibRefCitation id="12204B7A9B1D8310FB35F88FFF10F866" author="Monne, M. A. &amp; Hovore, F. T." pageId="20" pageNumber="471" refId="ref17379" refString="Monne, M. A. &amp; Hovore, F. T. (2006) A Checklist of the Cerambycidae, or longhorned wood-boring beetles, of the Western Hemisphere. BioQuip Publications, Rancho Dominguez, 393 pp." type="book" year="2006">Monné &amp; Hovore, 2006: 228</bibRefCitation>
</treatmentCitation>
(checklist);
<treatmentCitation id="F710109A9B1D8310FEF3F86BFDF2F866" author="Hovore, F. T." box="[335,547,1950,1977]" page="375" pageId="20" pageNumber="471" year="2006">
<bibRefCitation id="12204B7A9B1D8310FEF3F86BFDF2F866" author="Hovore, F. T." box="[335,547,1950,1977]" pageId="20" pageNumber="471" refId="ref16946" refString="Hovore, F. T. (2006) The Cerambycidae (Coleoptera) of Guatemala. In: Cano, E. B. (Ed) Biodiversidad de Guatemala. Vol. 1. Universidad del Valle de Guatemala, Guatemala, Guatemala, Centroamerica, pp. 363 - 378." type="book" year="2006">Hovore, 2006: 375</bibRefCitation>
</treatmentCitation>
(distr.);
<treatmentCitation id="F710109A9B1D8310FD39F86BFCA4F866" author="Swift, I. P. &amp; Bezark, L. G. &amp; Nearns, E. H. &amp; Solis, A. &amp; Hovore, F. T." box="[645,885,1950,1977]" page="51" pageId="20" pageNumber="471" year="2010">
<bibRefCitation id="12204B7A9B1D8310FD39F86BFCA4F866" author="Swift, I. P. &amp; Bezark, L. G. &amp; Nearns, E. H. &amp; Solis, A. &amp; Hovore, F. T." box="[645,885,1950,1977]" pageId="20" pageNumber="471" pagination="1 - 68" refId="ref17631" refString="Swift, I. P., Bezark, L. G., Nearns, E. H., Solis, A. &amp; Hovore, F. T. (2010) Checklist of the Cerambycidae (Coleoptera) of Costa Rica. Insecta Mundi, 131, 1 - 68." type="journal article" year="2010">
Swift
<emphasis id="44C5EA999B1D8310FD75F86BFD2AF866" box="[713,763,1951,1977]" italics="true" pageId="20" pageNumber="471">et al</emphasis>
., 2010: 51
</bibRefCitation>
</treatmentCitation>
(distr.);
<treatmentCitation id="F710109A9B1D8310FC6AF86BFB22F866" author="Gutierrez, N. &amp; Toledo-Hernandez, V. H. &amp; Noguera, F. A." box="[982,1267,1950,1977]" page="55" pageId="20" pageNumber="471" year="2020">
<bibRefCitation id="12204B7A9B1D8310FC6AF86BFB22F866" author="Gutierrez, N. &amp; Toledo-Hernandez, V. H. &amp; Noguera, F. A." box="[982,1267,1950,1977]" pageId="20" pageNumber="471" pagination="45 - 57" refId="ref16819" refString="Gutierrez, N., Toledo-Hernandez, V. H. &amp; Noguera, F. A. (2020) Four new species of Phrynidius Lacordaire (Coleoptera, Cerambycidae, Lamiinae) from Mexico with an identification key for the genus. ZooKeys, 1000 (2), 45 - 57. https: // doi. org / 10.3897 / zookeys. 1000.56757" type="journal article" year="2020">
Gutiérrez
<emphasis id="44C5EA999B1D8310FBF4F86BFB50F866" box="[1096,1153,1951,1977]" italics="true" pageId="20" pageNumber="471">et al.</emphasis>
, 2020: 55
</bibRefCitation>
</treatmentCitation>
(key);
<treatmentCitation id="F710109A9B1D8310FAFAF86BFED9F802" author="Monne, M. A." page="468" pageId="20" pageNumber="471" year="2023">
<bibRefCitation id="12204B7A9B1D8310FAFAF86BFED9F802" author="Monne, M. A." pageId="20" pageNumber="471" refId="ref17296" refString="Monne, M. A. (2023) Catalogue of the Cerambycidae (Coleoptera) of the Neotropical region. Part II. Subfamily Lamiinae. Available from: https: // cerambycids. com / catalog / (accessed 3 March 2023)" type="url" year="2023">Monné, 2023: 468</bibRefCitation>
</treatmentCitation>
(cat.).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<caption id="22CE66039B1C8311FF24F863FC6FF80B" ID-DOI="http://doi.org/10.5281/zenodo.8212338" ID-Zenodo-Dep="8212338" httpUri="https://zenodo.org/record/8212338/files/figure.png" pageId="21" pageNumber="472" startId="21.[152,270,1943,1967]" targetBox="[181,1406,181,1919]" targetPageId="21" targetType="figure">
<paragraph id="760E368B9B1C8311FF24F863FC6FF80B" blockId="21.[151,1436,1943,2004]" pageId="21" pageNumber="472">
<emphasis id="44C5EA999B1C8311FF24F863FE8AF870" bold="true" box="[152,347,1943,1967]" pageId="21" pageNumber="472">FIGURES 4651.</emphasis>
<taxonomicName id="B1B14D089B1C8311FEDDF86CFD74F86F" authority="Bates, 1880" authorityName="Bates" authorityYear="1880" box="[353,677,1944,1968]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="472" phylum="Arthropoda" rank="species" species="echinus">
<emphasis id="44C5EA999B1C8311FEDDF86CFDF2F86F" box="[353,547,1944,1968]" italics="true" pageId="21" pageNumber="472">Phrynidius echinus</emphasis>
<bibRefCitation id="12204B7A9B1C8311FD96F86CFD74F86F" author="Bates, H. W." box="[554,677,1944,1968]" pageId="21" pageNumber="472" refId="ref16372" refString="Bates, H. W. (1880) Longicornia, Fam. Cerambycidae [continuation]. In: Godman, F. D., Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Taylor and Francis, London, pp. 17 - 152." type="book" year="1880">Bates, 1880</bibRefCitation>
</taxonomicName>
.
<emphasis id="44C5EA999B1C8311FD13F863FD3FF870" bold="true" box="[687,750,1943,1967]" pageId="21" pageNumber="472">4649</emphasis>
) Male:
<emphasis id="44C5EA999B1C8311FC83F863FCB1F870" bold="true" box="[831,864,1943,1967]" pageId="21" pageNumber="472">46)</emphasis>
Dorsal habitus;
<emphasis id="44C5EA999B1C8311FBBBF863FBF8F870" bold="true" box="[1031,1065,1943,1967]" pageId="21" pageNumber="472">47)</emphasis>
Ventral habitus;
<emphasis id="44C5EA999B1C8311FB6AF863FB26F870" bold="true" box="[1238,1271,1943,1967]" pageId="21" pageNumber="472">48)</emphasis>
Lateral habitus;
<emphasis id="44C5EA999B1C8311FF2BF84FFF68F80C" bold="true" box="[151,185,1979,2003]" pageId="21" pageNumber="472">49)</emphasis>
Head, frontal view.
<emphasis id="44C5EA999B1C8311FE34F84FFE1EF80C" bold="true" box="[392,463,1979,2003]" pageId="21" pageNumber="472">5051)</emphasis>
Female:
<emphasis id="44C5EA999B1C8311FD92F84FFD9EF80C" bold="true" box="[558,591,1979,2003]" pageId="21" pageNumber="472">50)</emphasis>
Dorsal habitus;
<emphasis id="44C5EA999B1C8311FD4AF84FFCC6F80C" bold="true" box="[758,791,1979,2003]" pageId="21" pageNumber="472">51)</emphasis>
Ventral habitus.
</paragraph>
</caption>
<caption id="22CE66039B1F8312FF24FA16FCC2F99D" ID-DOI="http://doi.org/10.5281/zenodo.8212340" ID-Zenodo-Dep="8212340" httpUri="https://zenodo.org/record/8212340/files/figure.png" pageId="22" pageNumber="473" startId="22.[152,270,1506,1530]" targetBox="[161,1436,186,1476]" targetPageId="22" targetType="figure">
<paragraph id="760E368B9B1F8312FF24FA16FCC2F99D" blockId="22.[151,1437,1505,1602]" pageId="22" pageNumber="473">
<emphasis id="44C5EA999B1F8312FF24FA16FE7EFA26" bold="true" box="[152,431,1505,1530]" pageId="22" pageNumber="473">FIGURES 5256. 5254)</emphasis>
<taxonomicName id="B1B14D089B1F8312FE05FA16FCD7FA25" authority="Bates, 1880" authorityName="Bates" authorityYear="1880" box="[441,774,1506,1530]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="echinus">
<emphasis id="44C5EA999B1F8312FE05FA16FDAEFA25" box="[441,639,1506,1530]" italics="true" pageId="22" pageNumber="473">Phrynidius echinus</emphasis>
<bibRefCitation id="12204B7A9B1F8312FD35FA16FCD7FA25" author="Bates, H. W." box="[649,774,1506,1530]" pageId="22" pageNumber="473" refId="ref16372" refString="Bates, H. W. (1880) Longicornia, Fam. Cerambycidae [continuation]. In: Godman, F. D., Salvin, O. (Eds.), Biologia Centrali-Americana, Insecta, Coleoptera. Vol. 5. Taylor and Francis, London, pp. 17 - 152." type="book" year="1880">Bates, 1880</bibRefCitation>
</taxonomicName>
, holotype:
<emphasis id="44C5EA999B1F8312FCC1FA15FC4EFA26" bold="true" box="[893,927,1505,1529]" pageId="22" pageNumber="473">52)</emphasis>
Dorsal habitus;
<emphasis id="44C5EA999B1F8312FBECFA15FBA0FA26" bold="true" box="[1104,1137,1505,1529]" pageId="22" pageNumber="473">53)</emphasis>
Lateral habitus;
<emphasis id="44C5EA999B1F8312FA9BFA15FA99FA26" bold="true" box="[1319,1352,1505,1529]" pageId="22" pageNumber="473">54)</emphasis>
Labels.
<emphasis id="44C5EA999B1F8312FF2BF9F1FF0EF9C2" bold="true" box="[151,223,1541,1565]" pageId="22" pageNumber="473">5556)</emphasis>
<taxonomicName id="B1B14D089B1F8312FF54F9F2FD9DF9C1" authority="Linsley, 1933" authorityName="Linsley" authorityYear="1933" box="[232,588,1542,1566]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="473" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="44C5EA999B1F8312FF54F9F2FE65F9C1" box="[232,436,1542,1566]" italics="true" pageId="22" pageNumber="473">Phrynidius armatus</emphasis>
<bibRefCitation id="12204B7A9B1F8312FE01F9F2FD9DF9C1" author="Linsley, E. G." box="[445,588,1542,1566]" pageId="22" pageNumber="473" pagination="131 - 132" refId="ref17088" refString="Linsley, E. G. (1933) A new longicorn beetle from Central America. The Pan-Pacific Entomologist, 9 (3), 131 - 132." type="journal article" year="1933">Linsley, 1933</bibRefCitation>
</taxonomicName>
, holotype male:
<emphasis id="44C5EA999B1F8312FD42F9F1FCCEF9C2" bold="true" box="[766,799,1541,1565]" pageId="22" pageNumber="473">55)</emphasis>
Dorsal habitus;
<emphasis id="44C5EA999B1F8312FC73F9F1FC20F9C2" bold="true" box="[975,1009,1541,1565]" pageId="22" pageNumber="473">56)</emphasis>
Lateral habitus. Figures 5254 by Keita Matsumoto (BMNH); 5556 by Ranchel Diaz-Bastin (CASC).
</paragraph>
</caption>
<subSubSection id="3EAB65009B1F8313FF7BF960FA4AFD5A" lastPageId="23" lastPageNumber="474" pageId="22" pageNumber="473" type="description">
<paragraph id="760E368B9B1F8312FF7BF960FB39F8C5" blockId="22.[151,1437,1684,2035]" pageId="22" pageNumber="473">
<emphasis id="44C5EA999B1F8312FF7BF960FED5F971" bold="true" box="[199,260,1684,1710]" pageId="22" pageNumber="473">Male</emphasis>
(
<figureCitation id="EE8A2A0E9B1F8312FEAFF960FE49F971" box="[275,408,1684,1710]" captionStart="FIGURES 4651" captionStartId="21.[152,270,1943,1967]" captionTargetBox="[181,1406,181,1919]" captionTargetId="figure-17@21.[181,1406,181,1919]" captionTargetPageId="21" captionText="FIGURES 4651. Phrynidius echinus Bates, 1880. 4649) Male: 46) Dorsal habitus; 47) Ventral habitus; 48) Lateral habitus; 49) Head, frontal view. 5051) Female: 50) Dorsal habitus; 51) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212338" httpUri="https://zenodo.org/record/8212338/files/figure.png" pageId="22" pageNumber="473">Figs 4649</figureCitation>
). Integument mostly blackish; scape and pedicel dark brown; antennomere III brown; basal 2/3 of antennomere IV brown, and apical third reddish brown; antennomeres VXI reddish brown. Pubescence mostly greenish brown, with short, squamiform whitish setae interspersed, especially on frons, scape, antennomeres III and IV, femora, and tibiae, pubescence sparser on elytra, metaventrite, and abdominal ventrites.
</paragraph>
<paragraph id="760E368B9B1F8312FF7BF8D0FDAFF811" blockId="22.[151,1437,1684,2035]" pageId="22" pageNumber="473">
<emphasis id="44C5EA999B1F8312FF7BF8D0FEDFF8E1" bold="true" box="[199,270,1828,1854]" pageId="22" pageNumber="473">Head.</emphasis>
Antennal tubercles not close to each other basally, with area between them U-shaped. Antennae 1.45 times elytral length, almost reaching posterior quarter of elytra. Flagellomeres slender; antennomere III slightly arched. Antennal formula (ratio) based on length of antennomere III (only
<specimenCount id="60B7FD029B1F8312FC41F899FBA3F859" box="[1021,1138,1900,1927]" pageId="22" pageNumber="473" type="male">two males</specimenCount>
were measured): scape = 0.84/094; pedicel = 0.15/0.16; IV = 0.62/0.61; V = 0.33/0.36; VI = 0.29/0.33; VII = 0.29/0.33; VIII = 0.29/0.30; IX = 0.29/0.30; X = 0.25/0.28; XI = 0.27/0.33.
</paragraph>
<paragraph id="760E368B9B1F8313FF7BF82CFF2FFE57" blockId="22.[151,1437,1684,2035]" lastBlockId="23.[151,1437,150,933]" lastPageId="23" lastPageNumber="474" pageId="22" pageNumber="473">
<emphasis id="44C5EA999B1F8312FF7BF82CFEF6F82D" bold="true" box="[199,295,2008,2034]" pageId="22" pageNumber="473">Thorax.</emphasis>
Prothorax slightly wider than long; sides rounded from anterolateral angles to posterior eighth, more distinctly after middle, then parallel-sided to posterolateral angles. Pronotum abundantly, coarsely punctate; with three moderately elevated tubercles, one centrally, from anterior quarter to about posterior third, dorsally sinuous, more strongly elevated before its middle, and one with blunt apex on each side of maximum elevation of central tubercle.
<emphasis id="44C5EA999B1E8313FEBEFEF6FE87FEC3" bold="true" box="[258,342,258,284]" pageId="23" pageNumber="474">Elytra.</emphasis>
Sub-ovoid; abundantly, coarsely punctate; with elevated tubercles with blunt apex throughout, and smaller tubercles, with more acute apex basally, forming four arched, transverse rows on anterior 2/3, and two longitudinal rows on posterior third; without thick and erect setae.
<emphasis id="44C5EA999B1E8313FC07FEBEFC2BFEBB" bold="true" box="[955,1018,330,356]" pageId="23" pageNumber="474">Legs.</emphasis>
Femoral club gradually, not strongly widened.
</paragraph>
<paragraph id="760E368B9B1E8313FF7BFE66FDDAFE0F" blockId="23.[151,1437,150,933]" pageId="23" pageNumber="474">
<emphasis id="44C5EA999B1E8313FF7BFE66FECFFE73" bold="true" box="[199,286,402,428]" pageId="23" pageNumber="474">Female</emphasis>
(
<figureCitation id="EE8A2A0E9B1E8313FE8DFE66FE7CFE72" box="[305,429,402,429]" captionStart="FIGURES 4651" captionStartId="21.[152,270,1943,1967]" captionTargetBox="[181,1406,181,1919]" captionTargetId="figure-17@21.[181,1406,181,1919]" captionTargetPageId="21" captionText="FIGURES 4651. Phrynidius echinus Bates, 1880. 4649) Male: 46) Dorsal habitus; 47) Ventral habitus; 48) Lateral habitus; 49) Head, frontal view. 5051) Female: 50) Dorsal habitus; 51) Ventral habitus." figureDoi="http://doi.org/10.5281/zenodo.8212338" httpUri="https://zenodo.org/record/8212338/files/figure.png" pageId="23" pageNumber="474">Fig 5051</figureCitation>
). Similar to males; differ by the antennae slightly shorter, 1.1 times elytral length, not reaching posterior third of elytra.
</paragraph>
<paragraph id="760E368B9B1E8313FF7BFE2EFC41FDC7" blockId="23.[151,1437,150,933]" pageId="23" pageNumber="474">
<emphasis id="44C5EA999B1E8313FF7BFE2EFEEEFE2B" bold="true" box="[199,319,474,500]" pageId="23" pageNumber="474">Variation.</emphasis>
The pronotal tubercles are somewhat variable and may be slightly elevated; maximum width of the metafemoral club from 1.8 to 2.0 times basal width of the peduncle.
</paragraph>
<paragraph id="760E368B9B1E8313FF7BFDD6FA4AFD5A" blockId="23.[151,1437,150,933]" pageId="23" pageNumber="474">
<emphasis id="44C5EA999B1E8313FF7BFDD6FE76FDE3" bold="true" box="[199,423,546,572]" pageId="23" pageNumber="474">Dimensions in mm</emphasis>
(males/females). Total length, 8.809.80/9.1010.30; prothoracic length, 2.652.85/2.70 3.00; anterior prothoracic width, 1.851.95/2.002.40; posterior prothoracic width, 2.302.25/2.352.55; maximum prothoracic width, 2.802.75/2.803.05; humeral width, 2.502.30/2.502.80; elytral length, 5.405.90/5.806.50.
</paragraph>
</subSubSection>
<subSubSection id="3EAB65009B1E8313FF7BFD7AFAB9FC7A" pageId="23" pageNumber="474" type="discussion">
<paragraph id="760E368B9B1E8313FF7BFD7AFAB9FC7A" blockId="23.[151,1437,150,933]" pageId="23" pageNumber="474">
<emphasis id="44C5EA999B1E8313FF7BFD7AFEEAFD77" bold="true" box="[199,315,654,680]" pageId="23" pageNumber="474">Remarks.</emphasis>
<bibRefCitation id="12204B7A9B1E8313FEFEFD7AFDEAFD77" author="Gutierrez, N. &amp; Toledo-Hernandez, V. H. &amp; Noguera, F. A." box="[322,571,654,680]" pageId="23" pageNumber="474" pagination="45 - 57" refId="ref16819" refString="Gutierrez, N., Toledo-Hernandez, V. H. &amp; Noguera, F. A. (2020) Four new species of Phrynidius Lacordaire (Coleoptera, Cerambycidae, Lamiinae) from Mexico with an identification key for the genus. ZooKeys, 1000 (2), 45 - 57. https: // doi. org / 10.3897 / zookeys. 1000.56757" type="journal article" year="2020">
Gutiérrez
<emphasis id="44C5EA999B1E8313FE0EFD7BFE3BFD77" box="[434,490,654,680]" italics="true" pageId="23" pageNumber="474">et al.</emphasis>
(2020)
</bibRefCitation>
included
<taxonomicName id="B1B14D089B1E8313FD15FD7BFCCBFD77" authorityName="Bates" authorityYear="1880" box="[681,794,654,680]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="echinus">
<emphasis id="44C5EA999B1E8313FD15FD7BFCCBFD77" box="[681,794,654,680]" italics="true" pageId="23" pageNumber="474">P. echinus</emphasis>
</taxonomicName>
in the alternative of couplet “9” (“Scape distinctly longer than third antennomere”), coming from the alternative of couplet “8” (“Scape equal in length or shorter than third antennomere”). The inclusion in the alternative of couplet “9” is incorrect, since species with the scape longer than the antennomere III must be included in the alternative of couplet “10.” However, the scape in
<taxonomicName id="B1B14D089B1E8313FB01FD0FFAFFFCCB" authorityName="Bates" authorityYear="1880" box="[1213,1326,762,788]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="echinus">
<emphasis id="44C5EA999B1E8313FB01FD0FFAFFFCCB" box="[1213,1326,762,788]" italics="true" pageId="23" pageNumber="474">P. echinus</emphasis>
</taxonomicName>
is shorter than the antennomere III. Therefore, the length of scape and antennomere III cannot be used to separate
<taxonomicName id="B1B14D089B1E8313FA97FCEBFA4CFCE7" authorityName="Bates" authorityYear="1880" box="[1323,1437,798,824]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="echinus">
<emphasis id="44C5EA999B1E8313FA97FCEBFA4CFCE7" box="[1323,1437,798,824]" italics="true" pageId="23" pageNumber="474">P. echinus</emphasis>
</taxonomicName>
from
<taxonomicName id="B1B14D089B1E8313FF6AFCB7FD83FC83" authority="Gutierrez et al., 2020" authorityName="Gutierrez" authorityYear="2020" box="[214,594,834,861]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="cristinae">
<emphasis id="44C5EA999B1E8313FF6AFCB7FE85FC83" box="[214,340,835,860]" italics="true" pageId="23" pageNumber="474">P. cristinae</emphasis>
<bibRefCitation id="12204B7A9B1E8313FEE0FCB6FD83FC83" author="Gutierrez, N. &amp; Toledo-Hernandez, V. H. &amp; Noguera, F. A." box="[348,594,834,861]" pageId="23" pageNumber="474" pagination="45 - 57" refId="ref16819" refString="Gutierrez, N., Toledo-Hernandez, V. H. &amp; Noguera, F. A. (2020) Four new species of Phrynidius Lacordaire (Coleoptera, Cerambycidae, Lamiinae) from Mexico with an identification key for the genus. ZooKeys, 1000 (2), 45 - 57. https: // doi. org / 10.3897 / zookeys. 1000.56757" type="journal article" year="2020">
Gutiérrez
<emphasis id="44C5EA999B1E8313FE72FCB7FDD9FC83" box="[462,520,834,860]" italics="true" pageId="23" pageNumber="474">et al.</emphasis>
, 2020
</bibRefCitation>
</taxonomicName>
. Although these two species are very similar, they can be separated by the length of the antennomere IV, distinctly longer than half of antennomere III in
<taxonomicName id="B1B14D089B1E8313FBB7FC93FB59FC5F" box="[1035,1160,871,896]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="cristinae">
<emphasis id="44C5EA999B1E8313FBB7FC93FB59FC5F" box="[1035,1160,871,896]" italics="true" pageId="23" pageNumber="474">P. cristinae</emphasis>
</taxonomicName>
, about half the length of antennomere III in
<taxonomicName id="B1B14D089B1E8313FED3FC7FFE31FC7B" authorityName="Bates" authorityYear="1880" box="[367,480,906,932]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="echinus">
<emphasis id="44C5EA999B1E8313FED3FC7FFE31FC7B" box="[367,480,906,932]" italics="true" pageId="23" pageNumber="474">P. echinus</emphasis>
</taxonomicName>
, and pedicel about as long as wide in
<taxonomicName id="B1B14D089B1E8313FC36FC7FFBD6FC7B" box="[906,1031,907,932]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="cristinae">
<emphasis id="44C5EA999B1E8313FC36FC7FFBD6FC7B" box="[906,1031,907,932]" italics="true" pageId="23" pageNumber="474">P. cristinae</emphasis>
</taxonomicName>
, distinctly longer in
<taxonomicName id="B1B14D089B1E8313FB4CFC7FFAB0FC7B" authorityName="Bates" authorityYear="1880" box="[1264,1377,906,932]" class="Insecta" family="Cerambycidae" genus="Phrynidius" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="474" phylum="Arthropoda" rank="species" species="echinus">
<emphasis id="44C5EA999B1E8313FB4CFC7FFAB0FC7B" box="[1264,1377,906,932]" italics="true" pageId="23" pageNumber="474">P. echinus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>