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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1153.99537" ID-Pensoft-Pub="1313-2970-1153-73" ID-Pensoft-UUID="9A8EFF777E2258F090F13B331AFDFAE2" ID-ZooBank="D9BFB5475F3F45F1A5563F05839B4413" ModsDocID="1313-2970-1153-73" checkinTime="1678945459464" checkinUser="pensoft" docAuthor="Hou, Yanmeng, Feng, Zegang &amp; Zhang, Feng" docDate="2023" docId="FE02CCC0BBCB5ECB98A8A3ACE836B8B6" docLanguage="en" docName="ZooKeys 1153: 73-95" docOrigin="ZooKeys 1153" docPubDate="2023-03-15" docSource="http://dx.doi.org/10.3897/zookeys.1153.99537" docTitle="Lagynochthonius daidaiensis Hou &amp; Feng &amp; Zhang 2023, sp. nov." docType="treatment" docUuid="D23220D7-2AF6-47AE-B848-941BED5096E1" docUuidSource="ZooBank" docVersion="1" id="9A8EFF777E2258F090F13B331AFDFAE2" lastPageNumber="73" masterDocId="9A8EFF777E2258F090F13B331AFDFAE2" masterDocTitle="Three new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from eastern Yunnan, China" masterLastPageNumber="95" masterPageNumber="73" pageNumber="73" updateTime="1678945459464" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Three new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from eastern Yunnan, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hou, Yanmeng</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-0059-3419</mods:nameIdentifier>
<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Feng, Zegang</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5809-5984</mods:nameIdentifier>
<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China &amp; Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; College of Life Sciences, University of Chinese Academy of Sciences, Beijing, 100049, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhang, Feng</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3347-1031</mods:nameIdentifier>
<mods:affiliation>The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Development, College of Life sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
<mods:nameIdentifier type="email">dudu06042001@163.com</mods:nameIdentifier>
</mods:name>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-03-15</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1153</mods:number>
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<mods:start>73</mods:start>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1153.99537</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1153.99537</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1153-73</mods:identifier>
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<subSection pageId="0" pageNumber="73" type="taxonomy">
<treatment LSID="urn:lsid:zoobank.org:act:D23220D7-2AF6-47AE-B848-941BED5096E1" httpUri="http://treatment.plazi.org/id/FE02CCC0BBCB5ECB98A8A3ACE836B8B6" lastPageNumber="73" pageId="0" pageNumber="73">
<subSubSection pageId="0" pageNumber="73" type="nomenclature">
<paragraph pageId="0" pageNumber="73">
<taxonomicName LSID="https://zoobank.org/D23220D7-2AF6-47AE-B848-941BED5096E1" authority="Hou &amp; Feng &amp; Zhang, 2023" authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius daidaiensis" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="daidaiensis" status="sp. nov.">Lagynochthonius daidaiensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="73">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="73" type="description">
<paragraph pageId="0" pageNumber="73">
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Daidai Cave (type locality), habitat of Lagynochthonius daidaiensis sp. nov. A entrance B inside the cave entrance C live female of L. daidaiensis sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1153.99537.figure10" httpUri="https://binary.pensoft.net/fig/825483" pageId="0" pageNumber="73">Figs 10</figureCitation>
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lagynochthonius daidaiensis sp. nov., holotype female, habitus (dorsal view). Scale bar: 0.50 mm." figureDoi="10.3897/zookeys.1153.99537.figure11" httpUri="https://binary.pensoft.net/fig/825484" pageId="0" pageNumber="73">, 11</figureCitation>
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lagynochthonius daidaiensis sp. nov., holotype female A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E female genital area (ventral view) F left pedipalp (minus chela, dorsal view). Scale bars: 0.20 mm (A, B, F); 0.10 mm (C-E)." figureDoi="10.3897/zookeys.1153.99537.figure12" httpUri="https://binary.pensoft.net/fig/825485" pageId="0" pageNumber="73">, 12</figureCitation>
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Lagynochthonius daidaiensis sp. nov., holotype female A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73">
, 13 Chinese name:
<normalizedToken originalValue="呆呆拉伪蝎">呆呆拉伪蝎</normalizedToken>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="73" type="type material">
<paragraph pageId="0" pageNumber="73">Type material.</paragraph>
<paragraph pageId="0" pageNumber="73">
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Holotype</emphasis>
: China • ♀; Yunnan Province, Qiubei County, Shuanglongying Town, Pingtan Village, Daidai Cave;
<geoCoordinate degrees="24" direction="north" minutes="19.772" orientation="latitude" precision="1" value="24.329533">24°19.772'N</geoCoordinate>
,
<geoCoordinate degrees="104" direction="east" minutes="14.355" orientation="longitude" precision="1" value="104.23925">104°14.355'E</geoCoordinate>
; 1228 m a.s.l.; 20 Jul. 2021; Zegang Feng, Hongru Xu, Liu Fu and Nana Zhan leg.; under a stone in the dark zone; Ps.-MHBU-HBUARA#2021-176-01 (Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Study area, general cave locations, and type locality for each species, Yunnan Province, China. Each color represents an administrative region (green: Yunnan Province; red: Fuyuan County) A Daidai Cave (Lagynochthonius daidaiensis sp. nov.) B Xianren Cave (Tyrannochthonius capito sp. nov.) C, D unnamed cave and Dongtianfu Cave (T. calvatus sp. nov.)." figureDoi="10.3897/zookeys.1153.99537.figure1" httpUri="https://binary.pensoft.net/fig/825474" pageId="0" pageNumber="73">1A</figureCitation>
,
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Daidai Cave (type locality), habitat of Lagynochthonius daidaiensis sp. nov. A entrance B inside the cave entrance C live female of L. daidaiensis sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1153.99537.figure10" httpUri="https://binary.pensoft.net/fig/825483" pageId="0" pageNumber="73">10</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Paratypes</emphasis>
: • 2 ♀; the same data as the holotype; Ps.-MSWU-HBUARA#2021-176-02-HBUARA#2021-176-03.
</paragraph>
<caption doi="10.3897/zookeys.1153.99537.figure10" httpUri="https://binary.pensoft.net/fig/825483" pageId="0" pageNumber="73" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="73">
<emphasis bold="true" pageId="0" pageNumber="73">Figure 10.</emphasis>
Daidai Cave (type locality), habitat of
<taxonomicName authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius daidaiensis" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="daidaiensis">
<emphasis italics="true" pageId="0" pageNumber="73">Lagynochthonius daidaiensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="73">A</emphasis>
entrance
<emphasis bold="true" pageId="0" pageNumber="73">B</emphasis>
inside the cave entrance
<emphasis bold="true" pageId="0" pageNumber="73">C</emphasis>
live female of
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. daidaiensis" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="daidaiensis">
<emphasis italics="true" pageId="0" pageNumber="73">L. daidaiensis</emphasis>
</taxonomicName>
sp. nov. in its natural environment.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="73" type="diagnosis">
<paragraph pageId="0" pageNumber="73">Diagnosis</paragraph>
<paragraph pageId="0" pageNumber="73">
<emphasis bold="true" pageId="0" pageNumber="73">(♀).</emphasis>
Moderately sized troglomorphic species with elongated appendages; carapace without eyes or eyespots; anterior margin of carapace thin, finely denticulated, epistome pointed and small, triangular; posterior margin of carapace with two setae; tergites I-IV each with two setae. Pedipalps slender, femur 7.79-8.07
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad; chela 7.67-8.39
<normalizedToken originalValue="×">x</normalizedToken>
longer than deep; both chelal fingers without intercalary teeth but fixed chelal finger with a modified accessory tooth (
<emphasis italics="true" pageId="0" pageNumber="73">td</emphasis>
) on dorso-antiaxial face.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="73" type="etymology">
<paragraph pageId="0" pageNumber="73">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="73">Named after the type locality, Daidai Cave.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="73" type="description">
<paragraph pageId="0" pageNumber="73">Description.</paragraph>
<paragraph pageId="0" pageNumber="73">
<emphasis bold="true" pageId="0" pageNumber="73">Adult females</emphasis>
(male unknown) (Figs
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Daidai Cave (type locality), habitat of Lagynochthonius daidaiensis sp. nov. A entrance B inside the cave entrance C live female of L. daidaiensis sp. nov. in its natural environment." figureDoi="10.3897/zookeys.1153.99537.figure10" httpUri="https://binary.pensoft.net/fig/825483" pageId="0" pageNumber="73">10C</figureCitation>
,
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Lagynochthonius daidaiensis sp. nov., holotype female, habitus (dorsal view). Scale bar: 0.50 mm." figureDoi="10.3897/zookeys.1153.99537.figure11" httpUri="https://binary.pensoft.net/fig/825484" pageId="0" pageNumber="73">11</figureCitation>
-
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Lagynochthonius daidaiensis sp. nov., holotype female A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73">13</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Color</emphasis>
: generally pale yellow, chelicerae, pedipalps and tergites slightly darker, soft parts pale.
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Cephalothorax</emphasis>
(Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lagynochthonius daidaiensis sp. nov., holotype female A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E female genital area (ventral view) F left pedipalp (minus chela, dorsal view). Scale bars: 0.20 mm (A, B, F); 0.10 mm (C-E)." figureDoi="10.3897/zookeys.1153.99537.figure12" httpUri="https://binary.pensoft.net/fig/825485" pageId="0" pageNumber="73">12C</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Lagynochthonius daidaiensis sp. nov., holotype female A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73">13A, C</figureCitation>
): carapace 0.97
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad, gently narrowed posteriorly; surface smooth, without furrows; no traces of eyes; anterior margin slightly serrate; epistome pointed and small, triangular; with 18 setae arranged s4s:4:4:2:2, most setae heavy, long and gently curved, anterolateral setae much shorter than others; with three pairs of lyrifissures, the first two pairs situated middle and lateral to the setae of ocular row respectively, the third situated exterior to the sole pair of setae of posterior row. Chaetotaxy of coxae: P 3, I 3, II 3-4, III 5, IV 5; manducatory process with two acuminate distal setae, anterior seta less than 1/2 length of medial seta; apex of coxa I with small, rounded anteromedial process; coxae II with 11 or 12 terminally indented coxal spines on each side, set as an oblique and arc row, longer spines present in the middle of the row, becoming shorter distally and proximally and incised for ca. half their length (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Lagynochthonius daidaiensis sp. nov., holotype female A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73">13C</figureCitation>
); intercoxal tubercle absent; without sub-oral seta.
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Chelicera</emphasis>
(Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lagynochthonius daidaiensis sp. nov., holotype female A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E female genital area (ventral view) F left pedipalp (minus chela, dorsal view). Scale bars: 0.20 mm (A, B, F); 0.10 mm (C-E)." figureDoi="10.3897/zookeys.1153.99537.figure12" httpUri="https://binary.pensoft.net/fig/825485" pageId="0" pageNumber="73">12D</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Lagynochthonius daidaiensis sp. nov., holotype female A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73">13B, D</figureCitation>
): large, ca. as long as carapace, 2.41
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad; five setae and two lyrifissures (exterior condylar lyrifissure and exterior lyrifissure) present on hand, all setae acuminate, ventrobasal seta shorter than others; movable finger with one medial seta. Cheliceral palm with moderate hispid granulation on both ventral and dorsal sides. Both fingers well provided with teeth, fixed finger with 15-17 teeth, distal one largest; movable finger with 12 retrorse contiguous small teeth; galea absent (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Lagynochthonius daidaiensis sp. nov., holotype female A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73">13B</figureCitation>
). Serrula exterior with 21 blades and serrula interior with 13 or 14 blades. Rallum with seven blades, the distal one longest and recumbent basally, with fine barbules and slightly set apart from the other blades, latter tightly grouped and with long pinnae, some of which are subdivided (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Lagynochthonius daidaiensis sp. nov., holotype female A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73">13D</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Pedipalp</emphasis>
(Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lagynochthonius daidaiensis sp. nov., holotype female A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E female genital area (ventral view) F left pedipalp (minus chela, dorsal view). Scale bars: 0.20 mm (A, B, F); 0.10 mm (C-E)." figureDoi="10.3897/zookeys.1153.99537.figure12" httpUri="https://binary.pensoft.net/fig/825485" pageId="0" pageNumber="73">12A, B, F</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Lagynochthonius daidaiensis sp. nov., holotype female A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73">13E-G</figureCitation>
): long and slender, trochanter 1.25-1.41, femur 7.79-8.07, patella 2.35-2.53, chela 7.67-8.39, hand 3.29-3.61
<normalizedToken originalValue="×">x</normalizedToken>
longer than broad; femur 2.83-2.87
<normalizedToken originalValue="×">x</normalizedToken>
longer than patella; movable chelal finger 1.32-1.35
<normalizedToken originalValue="×">x</normalizedToken>
longer than hand and 0.57-0.58
<normalizedToken originalValue="×">x</normalizedToken>
longer than chela. Setae generally long and acuminate; one distal lyrifissure present on patella (Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lagynochthonius daidaiensis sp. nov., holotype female A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E female genital area (ventral view) F left pedipalp (minus chela, dorsal view). Scale bars: 0.20 mm (A, B, F); 0.10 mm (C-E)." figureDoi="10.3897/zookeys.1153.99537.figure12" httpUri="https://binary.pensoft.net/fig/825485" pageId="0" pageNumber="73">12F</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Lagynochthonius daidaiensis sp. nov., holotype female A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73">13E</figureCitation>
). Chelal palm gradually constricted towards fingers, apodeme complex of movable chelal finger strongly sclerotized, with weak granulation dorsally at base of fixed chelal finger. Fixed chelal finger and hand with eight trichobothria, movable chelal finger with four trichobothria,
<emphasis italics="true" pageId="0" pageNumber="73">ib</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="73">isb</emphasis>
situated close together, submedially on dorsum of chelal hand;
<emphasis italics="true" pageId="0" pageNumber="73">eb</emphasis>
,
<emphasis italics="true" pageId="0" pageNumber="73">esb</emphasis>
, and
<emphasis italics="true" pageId="0" pageNumber="73">ist</emphasis>
forming an oblique row at base of fixed chelal finger;
<emphasis italics="true" pageId="0" pageNumber="73">it</emphasis>
slightly distal to
<emphasis italics="true" pageId="0" pageNumber="73">est</emphasis>
, situated subdistally;
<emphasis italics="true" pageId="0" pageNumber="73">et</emphasis>
slightly near to tip of fixed chelal finger, very close to chelal teeth;
<emphasis italics="true" pageId="0" pageNumber="73">dx</emphasis>
situated distal to
<emphasis italics="true" pageId="0" pageNumber="73">et</emphasis>
;
<emphasis italics="true" pageId="0" pageNumber="73">sb</emphasis>
situated midway between
<emphasis italics="true" pageId="0" pageNumber="73">b</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="73">st</emphasis>
;
<emphasis italics="true" pageId="0" pageNumber="73">b</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="73">t</emphasis>
situated subdistally,
<emphasis italics="true" pageId="0" pageNumber="73">t</emphasis>
situated distal to
<emphasis italics="true" pageId="0" pageNumber="73">b</emphasis>
;
<emphasis italics="true" pageId="0" pageNumber="73">est</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="73">it</emphasis>
situated between
<emphasis italics="true" pageId="0" pageNumber="73">b</emphasis>
and
<emphasis italics="true" pageId="0" pageNumber="73">t</emphasis>
(Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Lagynochthonius daidaiensis sp. nov., holotype female A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73">13F</figureCitation>
). A tiny antiaxial lyrifissure present at base of fixed chelal finger (situated distal to
<emphasis italics="true" pageId="0" pageNumber="73">ist</emphasis>
). Both chelal fingers with a row of teeth, homodentate, spaced regularly along the margin, larger teeth present in the middle of the row, becoming smaller and closer distally and proximally: fixed chelal finger with 35 or 36 macrodenticles, slightly retrorse and pointed, plus a modified accessory tooth on dorso-antiaxial face (
<emphasis italics="true" pageId="0" pageNumber="73">td</emphasis>
, close to
<emphasis italics="true" pageId="0" pageNumber="73">dx</emphasis>
), 36 or 37 in total; movable chelal finger with 20-22 macrodenticles (slightly smaller than teeth on fixed chelal finger), slightly retrorse and pointed, plus nine or ten vestigial, rounded and contiguous basal teeth, 30 or 31 in total (Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Lagynochthonius daidaiensis sp. nov., holotype female A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73">13F</figureCitation>
). Chelal fingers slightly curved in dorsal view; microsetae (
<emphasis italics="true" pageId="0" pageNumber="73">sc</emphasis>
) present on dorsum of chelal hand (Figs
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lagynochthonius daidaiensis sp. nov., holotype female A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E female genital area (ventral view) F left pedipalp (minus chela, dorsal view). Scale bars: 0.20 mm (A, B, F); 0.10 mm (C-E)." figureDoi="10.3897/zookeys.1153.99537.figure12" httpUri="https://binary.pensoft.net/fig/825485" pageId="0" pageNumber="73">12B</figureCitation>
,
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Lagynochthonius daidaiensis sp. nov., holotype female A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73">13G</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Opisthosoma</emphasis>
: generally typical, pleural membrane finely granulated. Tergites and sternites undivided; setae uniseriate and acuminate. Tergal chaetotaxy I-XII: 2:2:2:2:4:4-5:4-5:5:5:4:T2T:0, tergites VIII and IX each with an unpaired median seta. Sternal chaetotaxy IV-XII: 11-12:7-8:7:7-8:7-8:7:7-8:0:2. Anterior genital operculum with 10 setae, posterior margin with 13-14 marginal setae, 23-24 in total (Fig.
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Lagynochthonius daidaiensis sp. nov., holotype female A left chela (lateral view) B left chela (dorsal view) C carapace (dorsal view) D left chelicera (dorsal view) E female genital area (ventral view) F left pedipalp (minus chela, dorsal view). Scale bars: 0.20 mm (A, B, F); 0.10 mm (C-E)." figureDoi="10.3897/zookeys.1153.99537.figure12" httpUri="https://binary.pensoft.net/fig/825485" pageId="0" pageNumber="73">12E</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Legs</emphasis>
(Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Lagynochthonius daidaiensis sp. nov., holotype female A carapace (dorsal view) B left chelicera (dorsal view) with details of dentation C coxal spines on coxae II (ventral view) D rallum E left pedipalp (minus chela, dorsal view) F left chela (lateral view) with details of dentation and trichobothrial pattern G left chela (dorsal view) H leg I (lateral view) I leg IV (lateral view). Scale bars: 0.20 mm (A, B, E-I); 0.10 mm (C, D)." figureDoi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73">13H, I</figureCitation>
): generally typical, long and slender. Fine granulation present on anterodorsal faces of femur IV and patella IV. Femur of leg I 1.88-1.97
<normalizedToken originalValue="×">x</normalizedToken>
longer than patella and with one lyrifissure at the base of femur; tarsus 2.72-2.73
<normalizedToken originalValue="×">x</normalizedToken>
longer than tibia. Femoropatella of leg IV 4.00
<normalizedToken originalValue="×">x</normalizedToken>
longer than deep; tibia 6.20-6.33
<normalizedToken originalValue="×">x</normalizedToken>
longer than deep; with basal tactile setae on both tarsal segments: basitarsus 3.50-3.71
<normalizedToken originalValue="×">x</normalizedToken>
longer than deep (TS = 0.29-0.35), telotarsus 14.60-15.20
<normalizedToken originalValue="×">x</normalizedToken>
longer than deep and 2.71-2.81
<normalizedToken originalValue="×">x</normalizedToken>
longer than basitarsus (TS = 0.36). Arolium slightly shorter than the claws, not divided; claws simple.
<emphasis bold="true" italics="true" pageId="0" pageNumber="73">Dimensions of adult females</emphasis>
(length/breadth or, in the case of the legs, also for chela and hand, length/depth in mm): body length 1.85-2.04. Pedipalps: trochanter 0.20-0.24/0.16-0.17, femur 1.09-1.13/0.14, patella 0.38-0.40/0.15-0.17, chela 1.51-1.61/0.18-0.21, hand 0.65-0.69/0.18-0.21, movable chelal finger length 0.86-0.93. Chelicera 0.65-0.70/0.27-0.29, movable finger length 0.34-0.36. Carapace 0.56-0.58/0.58-0.60. Leg I: trochanter 0.16-0.18/0.15, femur 0.60-0.65/0.08, patella 0.32-0.33/0.07, tibia 0.25-0.26/0.05-0.06, tarsus 0.68-0.71/0.05. Leg IV: trochanter 0.25-0.26/0.15, femoropatella 0.84-0.92/0.21-0.23, tibia 0.57-0.62/0.09-0.10, basitarsus 0.26-0.28/0.07-0.08, telotarsus 0.73-0.76/0.05.
</paragraph>
<caption doi="10.3897/zookeys.1153.99537.figure11" httpUri="https://binary.pensoft.net/fig/825484" pageId="0" pageNumber="73" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="73">
<emphasis bold="true" pageId="0" pageNumber="73">Figure 11.</emphasis>
<taxonomicName authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius daidaiensis" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="daidaiensis">
<emphasis italics="true" pageId="0" pageNumber="73">Lagynochthonius daidaiensis</emphasis>
</taxonomicName>
sp. nov., holotype female, habitus (dorsal view). Scale bar: 0.50 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1153.99537.figure12" httpUri="https://binary.pensoft.net/fig/825485" pageId="0" pageNumber="73" start="Figure 12" startId="F12">
<paragraph pageId="0" pageNumber="73">
<emphasis bold="true" pageId="0" pageNumber="73">Figure 12.</emphasis>
<taxonomicName authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius daidaiensis" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="daidaiensis">
<emphasis italics="true" pageId="0" pageNumber="73">Lagynochthonius daidaiensis</emphasis>
</taxonomicName>
sp. nov., holotype female
<emphasis bold="true" pageId="0" pageNumber="73">A</emphasis>
left chela (lateral view)
<emphasis bold="true" pageId="0" pageNumber="73">B</emphasis>
left chela (dorsal view)
<emphasis bold="true" pageId="0" pageNumber="73">C</emphasis>
carapace (dorsal view)
<emphasis bold="true" pageId="0" pageNumber="73">D</emphasis>
left chelicera (dorsal view)
<emphasis bold="true" pageId="0" pageNumber="73">E</emphasis>
female genital area (ventral view)
<emphasis bold="true" pageId="0" pageNumber="73">F</emphasis>
left pedipalp (minus chela, dorsal view). Scale bars: 0.20 mm (
<emphasis bold="true" pageId="0" pageNumber="73">A, B, F</emphasis>
); 0.10 mm (
<emphasis bold="true" pageId="0" pageNumber="73">C-E</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1153.99537.figure13" httpUri="https://binary.pensoft.net/fig/825486" pageId="0" pageNumber="73" start="Figure 13" startId="F13">
<paragraph pageId="0" pageNumber="73">
<emphasis bold="true" pageId="0" pageNumber="73">Figure 13.</emphasis>
<taxonomicName authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius daidaiensis" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="daidaiensis">
<emphasis italics="true" pageId="0" pageNumber="73">Lagynochthonius daidaiensis</emphasis>
</taxonomicName>
sp. nov., holotype female
<emphasis bold="true" pageId="0" pageNumber="73">A</emphasis>
carapace (dorsal view)
<emphasis bold="true" pageId="0" pageNumber="73">B</emphasis>
left chelicera (dorsal view) with details of dentation
<emphasis bold="true" pageId="0" pageNumber="73">C</emphasis>
coxal spines on coxae II (ventral view)
<emphasis bold="true" pageId="0" pageNumber="73">D</emphasis>
rallum
<emphasis bold="true" pageId="0" pageNumber="73">E</emphasis>
left pedipalp (minus chela, dorsal view)
<emphasis bold="true" pageId="0" pageNumber="73">F</emphasis>
left chela (lateral view) with details of dentation and trichobothrial pattern
<emphasis bold="true" pageId="0" pageNumber="73">G</emphasis>
left chela (dorsal view)
<emphasis bold="true" pageId="0" pageNumber="73">H</emphasis>
leg I (lateral view)
<emphasis bold="true" pageId="0" pageNumber="73">I</emphasis>
leg IV (lateral view). Scale bars: 0.20 mm (
<emphasis bold="true" pageId="0" pageNumber="73">A, B, E-I</emphasis>
); 0.10 mm (
<emphasis bold="true" pageId="0" pageNumber="73">C, D</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="73" type="remarks">
<paragraph pageId="0" pageNumber="73">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="73">
<taxonomicName authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius daidaiensis" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="daidaiensis">
<emphasis italics="true" pageId="0" pageNumber="73">Lagynochthonius daidaiensis</emphasis>
</taxonomicName>
sp. nov. is similar to
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. laoxueyanensis" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="laoxueyanensis">
<emphasis italics="true" pageId="0" pageNumber="73">L. laoxueyanensis</emphasis>
</taxonomicName>
Hou, Gao &amp; Zhang, 2022 (from Yunnan, China), but differs by the number of setae on tergites III-IV (2 vs. 4), a shorter chela (chela 7.67-8.39 vs. 6.88-7.22 (♀)
<normalizedToken originalValue="×">x</normalizedToken>
longer than deep, length 1.51-1.61 vs. 1.65-1.66 mm) and the number of coxal spines blades (11 or 12 vs. 9) (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.1097.82527" author="Hou, YM" journalOrPublisher="ZooKeys" pageId="0" pageNumber="73" pagination="65 - 83" refId="B12" refString="Hou, YM, Gao, ZZ, Zhang, F, 2022a. Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from Yunnan, China. ZooKeys 1097: 65 - 83, DOI: https://doi.org/10.3897/zookeys.1097.82527" title="Two new species of cave-adapted pseudoscorpions (Pseudoscorpiones, Chthoniidae) from Yunnan, China." url="https://doi.org/10.3897/zookeys.1097.82527" volume="1097" year="2022 a">Hou et al. 2022a</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="73">
<taxonomicName authorityName="Hou &amp; Feng &amp; Zhang" authorityYear="2023" class="Arachnida" family="Chthoniidae" genus="Lagynochthonius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lagynochthonius daidaiensis" order="Pseudoscorpiones" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="daidaiensis">
<emphasis italics="true" pageId="0" pageNumber="73">Lagynochthonius daidaiensis</emphasis>
</taxonomicName>
sp. nov. can be easily distinguished from
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. fengi" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="fengi">
<emphasis italics="true" pageId="0" pageNumber="73">L. fengi</emphasis>
</taxonomicName>
Hou, Gao &amp; Zhang, 2022,
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. retrorsus" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="retrorsus">
<emphasis italics="true" pageId="0" pageNumber="73">L. retrorsus</emphasis>
</taxonomicName>
Hou, Gao &amp; Zhang, 2022,
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. serratus" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="serratus">
<emphasis italics="true" pageId="0" pageNumber="73">L. serratus</emphasis>
</taxonomicName>
Hou, Gao &amp; Zhang, 2022,
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. spinulentus" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="spinulentus">
<emphasis italics="true" pageId="0" pageNumber="73">L. spinulentus</emphasis>
</taxonomicName>
Hou, Gao &amp; Zhang, 2022,
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. xiaolinensis" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="xiaolinensis">
<emphasis italics="true" pageId="0" pageNumber="73">L. xiaolinensis</emphasis>
</taxonomicName>
Hou, Gao &amp; Zhang, 2022 and
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. yaowangguensis" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="yaowangguensis">
<emphasis italics="true" pageId="0" pageNumber="73">L. yaowangguensis</emphasis>
</taxonomicName>
Hou, Gao &amp; Zhang, 2022 by the absence of intercalary teeth on both chelal fingers; from
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. crassus" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="crassus">
<emphasis italics="true" pageId="0" pageNumber="73">L. crassus</emphasis>
</taxonomicName>
Hou, Gao &amp; Zhang, 2022 by lacking a pair of anterior eyespots; from
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. magnidentatus" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="magnidentatus">
<emphasis italics="true" pageId="0" pageNumber="73">L. magnidentatus</emphasis>
</taxonomicName>
Hou, Gao &amp; Zhang, 2022 and
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. xinjiaoensis" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="xinjiaoensis">
<emphasis italics="true" pageId="0" pageNumber="73">L. xinjiaoensis</emphasis>
</taxonomicName>
Hou, Gao &amp; Zhang, 2022 by the presence of two carapaceal antero-median setae; from
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. bailongtanensis" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="bailongtanensis">
<emphasis italics="true" pageId="0" pageNumber="73">L. bailongtanensis</emphasis>
</taxonomicName>
Li, Liu &amp; Shi, 2019,
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. minimus" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="minimus">
<emphasis italics="true" pageId="0" pageNumber="73">L. minimus</emphasis>
</taxonomicName>
Hou, Gao &amp; Zhang, 2022 and
<taxonomicName class="Arachnida" kingdom="Animalia" lsidName="L. xibaiensis" pageId="0" pageNumber="73" phylum="Arthropoda" rank="species" species="xibaiensis">
<emphasis italics="true" pageId="0" pageNumber="73">L. xibaiensis</emphasis>
</taxonomicName>
Hou, Gao &amp; Zhang, 2022 by the number of setae on tergites I-II (2 vs. 3-4) (
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4571.1.2" author="Li, YC" journalOrPublisher="Zootaxa" pageId="0" pageNumber="73" pagination="28 - 34" refId="B19" refString="Li, YC, Liu, H, Shi, AM, 2019. A new cave-dwelling species of Lagynochthonius (Arachnida: Pseudoscorpiones: Chthoniidae) from Yunnan Province, China. Zootaxa 4571 (1): 28 - 34, DOI: https://doi.org/10.11646/zootaxa.4571.1.2" title="A new cave-dwelling species of Lagynochthonius (Arachnida: Pseudoscorpiones: Chthoniidae) from Yunnan Province, China." url="https://doi.org/10.11646/zootaxa.4571.1.2" volume="4571" year="2019">Li et al. 2019</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.5198.1.1" author="Hou, YM" journalOrPublisher="Zootaxa" pageId="0" pageNumber="73" pagination="001 - 065" refId="B13" refString="Hou, YM, Gao, ZZ, Zhang, F, 2022b. Diversity of cave-dwelling pseudoscorpions from eastern Yunnan in China, with the description of eleven new species of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae). Zootaxa 5198 (1): 001 - 065, DOI: https://doi.org/10.11646/zootaxa.5198.1.1" title="Diversity of cave-dwelling pseudoscorpions from eastern Yunnan in China, with the description of eleven new species of the genus Lagynochthonius (Pseudoscorpiones, Chthoniidae)." url="https://doi.org/10.11646/zootaxa.5198.1.1" volume="5198" year="2022 b">Hou et al. 2022b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="73" type="distribution">
<paragraph pageId="0" pageNumber="73">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="73">Known only from the type locality.</paragraph>
</subSubSection>
</treatment>
</subSection>
</document>