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<document id="E2CA87843C5C605EB714736B38704BAF" ID-CLB-Dataset="53270" ID-DOI="10.11646/zootaxa.5249.4.3" ID-GBIF-Dataset="44ba3149-bde1-4b85-bc58-43c9bfb6f1fa" ID-ISSN="1175-5326" ID-Zenodo-Dep="7694534" ID-ZooBank="F29E442D-4D11-4004-A867-03C9A07DC6BF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1677825808234" checkinUser="plazi" docAuthor="Jaitrong, Weeyawat, Yamane, Seiki &amp; Noon-Anant, Nawee" docDate="2023" docId="FD68B35DFFB4FF834EB9C283FAA6F873" docLanguage="en" docName="zootaxa.5249.4.3.pdf" docOrigin="Zootaxa 5249 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5249.4.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Polyrhachis varicolor Viehmeyer 1916" docType="treatment" docVersion="7" lastPageNumber="463" masterDocId="0151CB25FFBAFF924E2EC214FFB3FC46" masterDocTitle="The Thai species of the Polyrhachis (Myrmatopa) flavicornis Smith, 1857 species group, with description of a new species (Hymenoptera, Formicidae Formicinae)" masterLastPageNumber="464" masterPageNumber="446" pageNumber="460" updateTime="1699476145853" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="C65B72D028CB3E1DF8736B7BD98787C4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="90CEEBCC6007609EB6906D7478D5789D">
<mods:title id="82C66D4F8A5921F7E230E7FA493BA17C">The Thai species of the Polyrhachis (Myrmatopa) flavicornis Smith, 1857 species group, with description of a new species (Hymenoptera, Formicidae Formicinae)</mods:title>
</mods:titleInfo>
<mods:name id="6493AFFBCFCE7E2797D6B685BD11B7C3" type="personal">
<mods:role id="A86EE4F449F6C00B33DE7FCC213400F8">
<mods:roleTerm id="ECC245E296EBF1AD6B4FF49CD602F851">Author</mods:roleTerm>
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<mods:namePart id="384F7DCAF2220E265D28D2590212B29E">Jaitrong, Weeyawat</mods:namePart>
<mods:nameIdentifier id="001B87B077DFCE6A6A70F9979E8CC689" type="ORCID">0000-0003-1362-0754</mods:nameIdentifier>
<mods:affiliation id="4588A1024059EDBA8545BE990950768B">Office of Natural Science Research, National Science Museum, Technopolis, Khlong 5, Khlong Luang, Pathum Thani, 12120 Thailand. polyrhachis @ yahoo. com; https: // orcid. org / 0000 - 0003 - 1362 - 0754</mods:affiliation>
<mods:nameIdentifier id="E0F8D73CCFE7ED9691AED69BDD687913" type="email">polyrhachis@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="5BD7894FF1C0B46A64FCEA2CAE7002A5" type="personal">
<mods:role id="92FDEEAC7328DD4CEAE0E21667E9A50E">
<mods:roleTerm id="B8B657125D18328AF99E79817DA72DE0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="33E75F6A3AAC97E268FC7D55CECEEDCE">Yamane, Seiki</mods:namePart>
<mods:nameIdentifier id="481DC95CB2FA9C9611CD66161910B9B6" type="ORCID">0000-0002-6865-0865</mods:nameIdentifier>
<mods:affiliation id="6880FC6D3D1608C0763CA6A81E17854F">Haruyama-chô 1054 - 1, Kagoshima-shi, 899 - 2704 Japan. mayiopa 0 @ gmail. com; https: // orcid. org / 0000 - 0002 - 6865 - 0865 Division of Biological Science, Faculty of Science, Prince of Songkla University, Hat Yai, Songkhla, 90110, Thailand Corresponding author. nawee. n @ psu. ac. th; https: // orcid. org / 0000 - 0002 - 9999 - 1646</mods:affiliation>
<mods:nameIdentifier id="95EB78C1FD53FC9AF53A1FC9295FAEDD" type="email">mayiopa0@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="2EB6ACDCEB25A39A95ACA52178D9F260" type="personal">
<mods:role id="24E9B10237201090A842663F271E8220">
<mods:roleTerm id="A5DA63FF88F69330B9125AE19A1CF17E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DC814689157BBAB28C65875B256A767C">Noon-Anant, Nawee</mods:namePart>
<mods:nameIdentifier id="9DFC3B82BD0944732C410AFE83BC8562" type="ORCID">0000-0002-9999-1646</mods:nameIdentifier>
<mods:nameIdentifier id="7E0904BB5AD968BE5EB06FCF072FF167" type="email">nawee.n@psu.ac.th</mods:nameIdentifier>
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<mods:title id="8354E7462AF7D56D5B362D67C12D3E51">Zootaxa</mods:title>
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<mods:date id="CA4AB5BCAECFEFEBB1354C0814EAA114">2023</mods:date>
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<mods:number id="71B25B820A4A253F6AE5F49FA74CC3EE">2023-03-03</mods:number>
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<mods:number id="CDC8409AE14F8B2893022CEE4F44C42F">5249</mods:number>
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<treatment id="FD68B35DFFB4FF834EB9C283FAA6F873" ID-DOI="http://doi.org/10.5281/zenodo.7696786" ID-GBIF-Taxon="209409309" ID-Zenodo-Dep="7696786" LSID="urn:lsid:plazi:treatment:FD68B35DFFB4FF834EB9C283FAA6F873" httpUri="http://treatment.plazi.org/id/FD68B35DFFB4FF834EB9C283FAA6F873" lastPageId="17" lastPageNumber="463" pageId="14" pageNumber="460">
<subSubSection id="3DDB51C0FFB4FF9C4EB9C283FDC1FCF7" box="[151,626,151,178]" pageId="14" pageNumber="460" type="nomenclature">
<paragraph id="757E024BFFB4FF9C4EB9C283FDC1FCF7" blockId="14.[151,626,151,213]" box="[151,626,151,178]" pageId="14" pageNumber="460">
<heading id="2E36B527FFB4FF9C4EB9C283FDC1FCF7" bold="true" box="[151,626,151,178]" fontSize="11" level="1" pageId="14" pageNumber="460" reason="1">
<taxonomicName id="B2C179C8FFB4FF9C4EB9C283FDC1FCF7" authority="Viehmeyer, 1916" authorityName="Viehmeyer" authorityYear="1916" box="[151,626,151,178]" class="Insecta" family="Formicidae" genus="Polyrhachis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="460" phylum="Arthropoda" rank="species" species="varicolor">
<emphasis id="47B5DE59FFB4FF9C4EB9C283FDC1FCF7" bold="true" box="[151,626,151,178]" pageId="14" pageNumber="460">
<emphasis id="47B5DE59FFB4FF9C4EB9C283FE2EFCF4" bold="true" box="[151,413,151,178]" italics="true" pageId="14" pageNumber="460">Polyrhachis varicolor</emphasis>
Viehmeyer, 1916
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3DDB51C0FFB4FF9C4EB9C2AFFEB8FC93" box="[151,267,187,213]" pageId="14" pageNumber="460" type="description">
<paragraph id="757E024BFFB4FF9C4EB9C2AFFEB8FC93" blockId="14.[151,626,151,213]" box="[151,267,187,213]" pageId="14" pageNumber="460">
(
<figureCitation id="EDFA1ECEFFB4FF9C4EB1C2AFFEB0FC93" box="[159,259,187,213]" captionStart-0="FIGURE 8" captionStart-1="FIGURE 9" captionStartId-0="15.[152,255,1545,1569]" captionStartId-1="16.[152,255,1802,1826]" captionTargetBox-0="[163,1423,181,1521]" captionTargetBox-1="[151,1436,181,1778]" captionTargetId-0="figure-31@15.[163,1423,181,1521]" captionTargetId-1="figure-20@16.[151,1436,181,1778]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionText-0="FIGURE 8. Polyrhachis varicolor (non-type worker, TH21-WJT-003/1). A, Body in profile view; B, head in full-face view; C, body in dorsal view." captionText-1="FIGURE 9. Polyrhachis varicolor.A–C, Non-type dealate queen (TH21-WJT-003/2); D–F, non-type male (TH21-WJT-003/3); A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi-0="http://doi.org/10.5281/zenodo.7694558" figureDoi-1="http://doi.org/10.5281/zenodo.7694560" httpUri-0="https://zenodo.org/record/7694558/files/figure.png" httpUri-1="https://zenodo.org/record/7694560/files/figure.png" pageId="14" pageNumber="460">Figs 8–9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3DDB51C0FFB4FF9C4EB9C314FE7BFD13" pageId="14" pageNumber="460" type="reference_group">
<paragraph id="757E024BFFB4FF9C4EB9C314FB42FD5E" blockId="14.[151,1437,255,341]" box="[151,1265,255,280]" pageId="14" pageNumber="460">
<taxonomicName id="B2C179C8FFB4FF9C4EB9C314FC74FD51" ID-CoL="5KL7Q" authority="Viehmeyer, 1916: 163" authorityName="Viehmeyer" authorityPageNumber="163" authorityYear="1916" box="[151,967,255,280]" class="Insecta" family="Formicidae" genus="Polyrhachis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="460" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subGenus="Campomyrma" subSpecies="varicolor">
<emphasis id="47B5DE59FFB4FF9C4EB9C314FEA2FD5E" box="[151,273,256,280]" italics="true" pageId="14" pageNumber="460">Polyrhachis</emphasis>
(
<emphasis id="47B5DE59FFB4FF9C4F0EC314FE03FD51" box="[288,432,256,279]" italics="true" pageId="14" pageNumber="460">Campomyrma</emphasis>
)
<emphasis id="47B5DE59FFB4FF9C4F90C314FD80FD5E" box="[446,563,256,280]" italics="true" pageId="14" pageNumber="460">fruhstorferi</emphasis>
subsp.
<emphasis id="47B5DE59FFB4FF9C4CAEC314FD6DFD5E" box="[640,734,256,280]" italics="true" pageId="14" pageNumber="460">varicolor</emphasis>
Viehmeyer, 1916: 163
</taxonomicName>
, worker &amp; queen,
<collectingCountry id="0DD642DBFFB4FF9C4AADC314FB5EFD5E" box="[1155,1261,256,280]" name="Singapore" pageId="14" pageNumber="460">Singapore</collectingCountry>
.
</paragraph>
<paragraph id="757E024BFFB4FF9C4EB9C30AFE7BFD13" blockId="14.[151,1437,255,341]" pageId="14" pageNumber="460">
<treatmentCitationGroup id="55D12565FFB4FF9C4EB9C30AFE7BFD13" pageId="14" pageNumber="460">
<taxonomicName id="B2C179C8FFB4FF9C4EB9C30AFD84FD70" authority=": Kohout 2008: 259" authorityName=": Kohout" authorityPageNumber="259" authorityYear="2008" box="[151,567,286,310]" class="Insecta" family="Formicidae" genus="Polyrhachis" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="460" phylum="Arthropoda" rank="species" species="varicolor">
<emphasis id="47B5DE59FFB4FF9C4EB9C30AFEC7FD70" box="[151,372,286,310]" italics="true" pageId="14" pageNumber="460">Polyrhachis varicolor</emphasis>
: Kohout 2008: 259
</taxonomicName>
(raised to species); Noon-anant
<emphasis id="47B5DE59FFB4FF9C4D57C30BFC16FD70" box="[889,933,286,310]" italics="true" pageId="14" pageNumber="460">et al</emphasis>
. 2009: 177, figs 3–8;
<treatmentCitation id="F460245AFFB4FF9C4A50C30AFF50FD13" author="Khachonpisitsak, S. &amp; Yamane, S. &amp; Siriwichai, P. &amp; Jaitrong, W." page="75" pageId="14" pageNumber="460" year="2020">
<bibRefCitation id="11507FBAFFB4FF9C4A50C30AFF50FD13" author="Khachonpisitsak, S. &amp; Yamane, S. &amp; Siriwichai, P. &amp; Jaitrong, W." pageId="14" pageNumber="460" pagination="1 - 182" refId="ref10340" refString="Khachonpisitsak, S., Yamane, S., Siriwichai, P. &amp; Jaitrong, W. (2020) An updated checklist of the ants of Thailand (Hymenoptera, Formicidae). ZooKeys, 998, 1 - 182. https: // doi. org / 10.3897 / zookeys. 998.54902" type="journal article" year="2020">
Khachonpisitsak
<emphasis id="47B5DE59FFB4FF9C4B02C30BFAE4FD70" box="[1324,1367,286,310]" italics="true" pageId="14" pageNumber="460">et al</emphasis>
, 2020: 75
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="F460245AFFB4FF9C4EC3C329FE70FD13" author="Wang, W. Y. &amp; Soh, E. J. Y. &amp; Yong, G. W. J. &amp; Wong, M. K. L. &amp; Guenard, B. &amp; Economo, E. P. &amp; Yamane, S." box="[237,451,317,341]" page="59" pageId="14" pageNumber="460" year="2022">
<bibRefCitation id="11507FBAFFB4FF9C4EC3C329FE70FD13" author="Wang, W. Y. &amp; Soh, E. J. Y. &amp; Yong, G. W. J. &amp; Wong, M. K. L. &amp; Guenard, B. &amp; Economo, E. P. &amp; Yamane, S." box="[237,451,317,341]" pageId="14" pageNumber="460" pagination="015006" refId="ref10889" refString="Wang, W. Y., Soh, E. J. Y., Yong, G. W. J., Wong, M. K. L., Guenard, B., Economo, E. P. &amp; Yamane, S. (2022) Remarkable diversity in a little red dot: a comprehensive checklist of known ant species in Singapore (Hymenoptera: Formicidae) with notes on ecology and taxonomy. Asian Myrmecology, 15, e 015006. https: // doi. org / 10.20362 / am. 015006" type="journal article" year="2022">
Wang
<emphasis id="47B5DE59FFB4FF9C4F00C32AFEE8FD13" box="[302,347,317,341]" italics="true" pageId="14" pageNumber="460">et al</emphasis>
. 2022: 59
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="3DDB51C0FFB4FF9C4EB9C397FBCBFE6B" pageId="14" pageNumber="460" type="materials_examined">
<paragraph id="757E024BFFB4FF9C4EB9C397FBCBFE6B" blockId="14.[151,1437,387,2034]" pageId="14" pageNumber="460">
<emphasis id="47B5DE59FFB4FF9C4EB9C397FE46FDDB" bold="true" box="[151,501,387,413]" pageId="14" pageNumber="460">Non-type material examined.</emphasis>
<materialsCitation id="C5A90816FFB4FF9C4FD2C397FA8DFD87" ID-GBIF-Occurrence="4041564315" collectingDate="2006-03-08" collectionCode="PSUSC, SKYC" collectorName="N. Noon-anant" country="Thailand" elevation="180" location="Nakon Si Thamarat Prov." municipality="Prachuab Khiri Khan Prov." pageId="14" pageNumber="460" specimenCount="1">
<emphasis id="47B5DE59FFB4FF9C4FD2C397FD2FFDDB" bold="true" box="[508,668,387,413]" pageId="14" pageNumber="460">
<collectingCountry id="0DD642DBFFB4FF9C4FD2C397FD24FDDB" box="[508,663,387,413]" name="Thailand" pageId="14" pageNumber="460">THAILAND</collectingCountry>
:
</emphasis>
W Thailand,
<collectingMunicipality id="951A9831FFB4FF9C4D19C397FBD8FDD8" box="[823,1131,387,414]" pageId="14" pageNumber="460">Prachuab Khiri Khan Prov.</collectingMunicipality>
,
<location id="701E5490FFB4FF9C4A59C397FA2AFDD8" LSID="urn:lsid:plazi:treatment:FD68B35DFFB4FF834EB9C283FAA6F873:701E5490FFB4FF9C4A59C397FA2AFDD8" box="[1143,1433,387,414]" country="Thailand" municipality="Prachuab Khiri Khan Prov." name="Nakon Si Thamarat Prov." pageId="14" pageNumber="460">Nakon Si Thamarat Prov.</location>
,
<quantity id="B239AFAEFFB4FF9C4EB9C3B3FF6CFD84" box="[151,223,423,450]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="460" unit="m" value="180.0">
<elevation id="FEECE578FFB4FF9C4EB9C3B3FF6CFD84" box="[151,223,423,450]" metricMagnitude="2" metricUnit="m" metricValue="1.8" pageId="14" pageNumber="460" unit="m" value="180.0">180 m</elevation>
</quantity>
alt., lower-vegetation,
<date id="017F248BFFB4FF9C4FCAC3B3FDE5FD87" box="[484,598,423,449]" pageId="14" pageNumber="460" value="2006-03-08">
<collectingDate id="113BDD63FFB4FF9C4FCAC3B3FDE5FD87" box="[484,598,423,449]" pageId="14" pageNumber="460" value="2006-03-08">8.III.2006</collectingDate>
</date>
,
<collectorName id="D834679DFFB4FF9C4C4DC3BCFCB9FD84" box="[611,778,424,450]" pageId="14" pageNumber="460">N. Noon-anant</collectorName>
leg., NAW-POL6-024 (workers,
<collectionCode id="13D09A8EFFB4FF9C4AACC3B3FB69FD87" box="[1154,1242,423,449]" collectionName="PSUSC" pageId="14" pageNumber="460">PSUSC</collectionCode>
,
<collectionCode id="13D09A8EFFB4FF9C4AC8C3B3FA86FD87" box="[1254,1333,423,449]" pageId="14" pageNumber="460">SKYC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C5A90816FFB4FF9C4B64C3B3FBB7FE4F" ID-GBIF-Occurrence="4041564341" collectingDate="2021-03-26" collectionCode="THNHM" collectorName="W. Jaitrong" country="Thailand" county="Central Thailand" location="Nang Rong Temple" municipality="Muaeng Dist." pageId="14" pageNumber="460" specimenCount="1" stateProvince="Nakhon Nayok Prov.">
Central
<collectingCountry id="0DD642DBFFB4FF9C4EB9C3DFFF4EFDA3" box="[151,253,459,485]" name="Thailand" pageId="14" pageNumber="460">Thailand</collectingCountry>
,
<collectingRegion id="B705CCA9FFB4FF9C4F24C3DFFE4FFDA0" box="[266,508,459,486]" country="Thailand" name="Nakhon Nayok" pageId="14" pageNumber="460">Nakhon Nayok Prov.</collectingRegion>
,
<collectingMunicipality id="951A9831FFB4FF9C4C26C3D8FD16FDA3" box="[520,677,459,486]" pageId="14" pageNumber="460">Muaeng Dist.</collectingMunicipality>
, near
<location id="701E5490FFB4FF9C4CC4C3D8FC78FDA3" LSID="urn:lsid:plazi:treatment:FD68B35DFFB4FF834EB9C283FAA6F873:701E5490FFB4FF9C4CC4C3D8FC78FDA3" box="[746,971,459,486]" country="Thailand" county="Central Thailand" municipality="Muaeng Dist." name="Nang Rong Temple" pageId="14" pageNumber="460" stateProvince="Nakhon Nayok Prov.">Nang Rong Temple</location>
, nest under leaf,
<date id="017F248BFFB4FF9C4AA0C3DFFABDFDA3" box="[1166,1294,459,485]" pageId="14" pageNumber="460" value="2021-03-26">
<collectingDate id="113BDD63FFB4FF9C4AA0C3DFFABDFDA3" box="[1166,1294,459,485]" pageId="14" pageNumber="460" value="2021-03-26">26.III.2021</collectingDate>
</date>
,
<collectorName id="D834679DFFB4FF9C4B35C3D8FA2FFDA3" box="[1307,1436,459,486]" pageId="14" pageNumber="460">W. Jaitrong</collectorName>
leg., TH21-WJT-003 (
<collectionCode id="13D09A8EFFB4FF9C4FBFC3FBFE4FFE4F" box="[401,508,495,521]" pageId="14" pageNumber="460">THNHM</collectionCode>
, workers, dealate queen, alate queens, males)
</materialsCitation>
;
<materialsCitation id="C5A90816FFB4FF9C4A3FC3FBFBC7FE6B" ID-GBIF-Occurrence="4041564334" collectingDate="2008-09-27" collectionCode="SKYC" collectorName="Sk. Yamane" country="Thailand" location="Khao Nan" municipality="Khao" pageId="14" pageNumber="460" specimenCount="1" stateProvince="Nakhon Si Thammarat Prov.">
S
<collectingCountry id="0DD642DBFFB4FF9C4A06C3FBFB3DFE4F" box="[1064,1166,495,521]" name="Thailand" pageId="14" pageNumber="460">Thailand</collectingCountry>
,
<collectingRegion id="B705CCA9FFB4FF9C4AB5C3FBFF66FE68" country="Thailand" name="Nakhon Si Thammarat" pageId="14" pageNumber="460">Nakhon Si Thammarat Prov.</collectingRegion>
,
<location id="701E5490FFB4FF9C4EF1C007FEE1FE68" LSID="urn:lsid:plazi:treatment:FD68B35DFFB4FF834EB9C283FAA6F873:701E5490FFB4FF9C4EF1C007FEE1FE68" box="[223,338,531,558]" country="Thailand" municipality="Khao" name="Khao Nan" pageId="14" pageNumber="460" stateProvince="Nakhon Si Thammarat Prov.">Khao Nan</location>
NP,
<location id="701E5490FFB4FF9C4FA8C007FDD0FE6B" LSID="urn:lsid:plazi:treatment:FD68B35DFFB4FF834EB9C283FAA6F873:701E5490FFB4FF9C4FA8C007FDD0FE6B" box="[390,611,531,558]" country="Thailand" municipality="Khao" name="Yod Nam Waterfall" pageId="14" pageNumber="460" stateProvince="Nakhon Si Thammarat Prov.">Yod Nam Waterfall</location>
,
<date id="017F248BFFB4FF9C4C43C007FD5CFE6B" box="[621,751,531,557]" pageId="14" pageNumber="460" value="2008-09-27">
<collectingDate id="113BDD63FFB4FF9C4C43C007FD5CFE6B" box="[621,751,531,557]" pageId="14" pageNumber="460" value="2008-09-27">27.IX.2008</collectingDate>
</date>
,
<collectorName id="D834679DFFB4FF9C4CD5C007FCCCFE68" box="[763,895,531,558]" pageId="14" pageNumber="460">Sk. Yamane</collectorName>
leg. (worker,
<collectionCode id="13D09A8EFFB4FF9C4A32C007FBDFFE6B" box="[1052,1132,531,557]" pageId="14" pageNumber="460">SKYC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3DDB51C0FFB4FF834EE9C023FEB3FFAB" lastPageId="17" lastPageNumber="463" pageId="14" pageNumber="460" type="description">
<paragraph id="757E024BFFB4FF9C4EE9C023FD42FEF8" blockId="14.[151,1437,387,2034]" pageId="14" pageNumber="460">
<emphasis id="47B5DE59FFB4FF9C4EE9C023FDB4FE17" bold="true" box="[199,519,567,593]" pageId="14" pageNumber="460">Measurements and indices</emphasis>
. Worker (10 non-types). TL 3.05–4.65, HL 1.30–1.45, HW 1.00–1.25, SL 1.75– 1.95, PW 0.85–1.05, MTL 1.50–1.75, CI 80–83, SI 150–180. Queen (3 non-types). TL 5.00–5.45, HL 1.60–1.70, HW 1.25, SL 1.90-2.05, PW1.45–1.50, MTL 1.85–2.00, CI 74–78, SI 152–164. Male (non-type). TL 4.8, HL 0.9, HW 0.65, SL 1.00, PW 1.25, MTL 1.7, CI 72, SI 154.
</paragraph>
<paragraph id="757E024BFFB4FF9C4EE9C0D3FDECFEA7" blockId="14.[151,1437,387,2034]" box="[199,607,711,737]" pageId="14" pageNumber="460">
<emphasis id="47B5DE59FFB4FF9C4EE9C0D3FE7DFEA7" bold="true" box="[199,462,711,737]" pageId="14" pageNumber="460">Description of worker</emphasis>
(
<figureCitation id="EDFA1ECEFFB4FF9C4FF3C0D3FDACFEA7" box="[477,543,711,737]" captionStart="FIGURE 8" captionStartId="15.[152,255,1545,1569]" captionTargetBox="[163,1423,181,1521]" captionTargetId="figure-31@15.[163,1423,181,1521]" captionTargetPageId="15" captionText="FIGURE 8. Polyrhachis varicolor (non-type worker, TH21-WJT-003/1). A, Body in profile view; B, head in full-face view; C, body in dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7694558" httpUri="https://zenodo.org/record/7694558/files/figure.png" pageId="14" pageNumber="460">Fig. 8</figureCitation>
A-C)
</paragraph>
<paragraph id="757E024BFFB4FF9C4EE9C0FFFAE5F863" blockId="14.[151,1437,387,2034]" pageId="14" pageNumber="460">Head in full-face view distinctly longer than broad, with posterior margin evenly and roundly convex, lateral margins distinctly converging anteriad. Frontal carinae in full-face view posteriorly reaching level of mid-length of eye, parallel anteriorly, divergent in middle, parallel posteriorly; frontal carina in profile distinctly elevated. Clypeus in dorsal view transverse, trapezoidal; its posteromedian margin straight to very shallowly emarginated; anterior margin slightly convex; anteromedian portion weakly produced with its anterior margin straight and usually with acute lateral angles. Mandible subtriangular, masticatory margin with five distinct teeth. Antennal scape long, ca. 1.30–1.37 times as long as head width including eyes; pedicel (antennal segment II) about 2 times as long as antennal segment III; all flagellomeres longer than broad with the first shortest. Eye in full-face view convex and prominent, distinctly breaking lateral margin of head, located posterior to mid-length on lateral face of head.</paragraph>
<paragraph id="757E024BFFB4FF9C4EE9C624FC01F9F7" blockId="14.[151,1437,387,2034]" pageId="14" pageNumber="460">Pronotum in dorsal view much broader than long, anteriorly and laterally weakly convex, with very shallowly concave posterior margins; humerus bluntly angled or produced as small process; in profile view lateral face of pronotum with shallow furrow along its anterior margins; posteroventral corner broadly rounded. Mesonotum in dorsal view demarcated anteriorly and posteriorly by distinct furrows, broader than long, gradually narrowed posteriad, posteriorly much narrower than pronotum; in profile view dorsal margin almost flat but lateral keel with angle in its posterior half; mesopleuron anteriorly demarcated from pronotum by sinuated deep furrow but completely fused with metapleuron. Metanotal groove short (‘narrow’) and shallow, continuing down to metathoracic spiracle that has an upward-directed small opening. Propodeum in dorsal view as broad as mesonotum posteriorly; dorsal face rectangular, slightly broader than long, gradually merging with posterior declivity; propodeal spine short, upward directed; lateral face not demarcated from metapleuron; propodeal lobe very small; propodeal spiracle located on lateral margin of declivity, prominent, with oblong opening.</paragraph>
<paragraph id="757E024BFFB4FF9C4EE9C7AFFB30FA07" blockId="14.[151,1437,387,2034]" pageId="14" pageNumber="460">Petiole in dorsal view broader than long; dorsal face not clearly differentiated from anterior and posterior slopes, with pair of long lateral spines that diverge in basal part and are parallel in apical two-thirds, without denticle between them; in profile view anterior slope of node straight, much shorter than posterior slope that is roundly convex; subpetiolar process weakly developed, subtrapezoid with straight ventral margin.</paragraph>
<paragraph id="757E024BFFB4FF9C4EE9C458FEBEFB24" blockId="14.[151,1437,387,2034]" pageId="14" pageNumber="460">Vertex and frons of head areolate-rugose, area between frontal carinae puncto-reticulate, areas between eyes and frontal carinae weaklier sculptured; gena behind eye and venter of head densely puncto-reticulate. Clypeus densely punctate. Mandible in basal half very densely and finely striate with scattered punctures from which setae arise; apical half with much sparser and weaker sculpture and shiny.Antenna densely micropunctate. Dorsum of pronotum puncto-striate, but with weak striation; mesonotum and dorsum of propodeum densely macropunctate; lateral face of mesosoma including lateral face of pronotum densely macropunctate. Petiole densely macropunctate. Gastral tergites finely micropunctate and matte; sternites with coarser punctures but interspaces shiny. Legs micropunctate and matte.</paragraph>
<paragraph id="757E024BFFB4FF9C4EE9C578FAA8FBB4" blockId="14.[151,1437,387,2034]" pageId="14" pageNumber="460">Dorsa of head, mesosoma, petiole, and gaster almost without standing hairs; few very short standing hairs present on outer margin of mandible, clypeus anteriorly, frontal lobe, petiolar sternite anteriorly and venter of gaster. Eye without standing hairs. Antennal scape without standing hairs. Body brown to dark reddish brown; gaster darker than elsewhere [in some specimens gaster with similar color or slightly paler than head and mesosoma]</paragraph>
<caption id="21BE52C3FFB5FF9D4EB6C41DFED6FA03" ID-DOI="http://doi.org/10.5281/zenodo.7694558" ID-Zenodo-Dep="7694558" httpUri="https://zenodo.org/record/7694558/files/figure.png" pageId="15" pageNumber="461" startId="15.[152,255,1545,1569]" targetBox="[163,1423,181,1521]" targetPageId="15">
<paragraph id="757E024BFFB5FF9D4EB6C41DFED6FA03" blockId="15.[151,1436,1545,1605]" pageId="15" pageNumber="461">
<emphasis id="47B5DE59FFB5FF9D4EB6C41DFEABFA67" bold="true" box="[152,280,1545,1569]" pageId="15" pageNumber="461">FIGURE 8.</emphasis>
<taxonomicName id="B2C179C8FFB5FF9D4F30C41DFE48FA67" authorityName=": Kohout" authorityYear="2008" box="[286,507,1545,1569]" class="Insecta" family="Formicidae" genus="Polyrhachis" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="461" phylum="Arthropoda" rank="species" species="varicolor">
<emphasis id="47B5DE59FFB5FF9D4F30C41DFE48FA67" box="[286,507,1545,1569]" italics="true" pageId="15" pageNumber="461">Polyrhachis varicolor</emphasis>
</taxonomicName>
(non-type worker, TH21-WJT-003/1). A, Body in profile view; B, head in full-face view; C, body in dorsal view.
</paragraph>
</caption>
<paragraph id="757E024BFFB5FF9D4EE9C467FDE7FACB" blockId="15.[151,1437,1651,2038]" box="[199,596,1651,1677]" pageId="15" pageNumber="461">
<emphasis id="47B5DE59FFB5FF9D4EE9C467FE0CFACB" bold="true" box="[199,447,1651,1677]" pageId="15" pageNumber="461">Description of queen</emphasis>
(
<figureCitation id="EDFA1ECEFFB5FF9D4FE0C467FDFFFACB" box="[462,588,1651,1677]" captionStart="FIGURE 9" captionStartId="16.[152,255,1802,1826]" captionTargetBox="[151,1436,181,1778]" captionTargetId="figure-20@16.[151,1436,181,1778]" captionTargetPageId="16" captionText="FIGURE 9. Polyrhachis varicolor.A–C, Non-type dealate queen (TH21-WJT-003/2); D–F, non-type male (TH21-WJT-003/3); A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7694560" httpUri="https://zenodo.org/record/7694560/files/figure.png" pageId="15" pageNumber="461">Fig. 9 A–C</figureCitation>
)
</paragraph>
<paragraph id="757E024BFFB5FF9D4EE9C483FE5FFBB3" blockId="15.[151,1437,1651,2038]" pageId="15" pageNumber="461">
General habitus similar to the worker except for caste-specific structures. Body slightly larger than the average worker with head width
<quantity id="B239AFAEFFB5FF9D4F88C4AFFDBAFA90" box="[422,521,1723,1750]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="15" pageNumber="461" unit="mm" value="1.25">1.25 mm</quantity>
(1.00–
<quantity id="B239AFAEFFB5FF9D4C79C4AFFD09FA90" box="[599,698,1723,1750]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="15" pageNumber="461" unit="mm" value="1.25">1.25 mm</quantity>
in worker). Eye larger and more prominent; ocelli arranged in low triangle. Anterolateral corner of pronotal dorsum round, without any projection. Mesoscutum without notauli, with weak parapsidal line in posterior half of scutum; prescutal carina absent; mesopleuron divided into upper and lower sections by groove; anterior margin of lower section with high acute keel (not conspicuous in the worker). Metanotum in dorsal view short (‘narrow’), indented, sharply demarcated from propodeum by deep furrow; metapleuron barely differentiated from mesopleuron, completely fused with propodeal side; propodeal spines short, blunt. Body slightly hairier; in addition to standing hairs mentioned for the worker, queen bears standing hairs as follows: one or two around ocelli, several on mesoscutum, some on mesoscutellum, some on second to fifth gastral tergites. Body color almost identical to the worker.
</paragraph>
<caption id="21BE52C3FFAAFF824EB6C51EFC69FB00" ID-DOI="http://doi.org/10.5281/zenodo.7694560" ID-Zenodo-Dep="7694560" httpUri="https://zenodo.org/record/7694560/files/figure.png" pageId="16" pageNumber="462" startId="16.[152,255,1802,1826]" targetBox="[151,1436,181,1778]" targetPageId="16">
<paragraph id="757E024BFFAAFF824EB6C51EFC69FB00" blockId="16.[151,1436,1802,1862]" pageId="16" pageNumber="462">
<emphasis id="47B5DE59FFAAFF824EB6C51EFEA4FB64" bold="true" box="[152,279,1802,1826]" pageId="16" pageNumber="462">FIGURE 9.</emphasis>
<emphasis id="47B5DE59FFAAFF824F33C51EFE4CFB64" box="[285,511,1802,1826]" italics="true" pageId="16" pageNumber="462">
<taxonomicName id="B2C179C8FFAAFF824F33C51EFE48FB64" authorityName=": Kohout" authorityYear="2008" box="[285,507,1802,1826]" class="Insecta" family="Formicidae" genus="Polyrhachis" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="462" phylum="Arthropoda" rank="species" species="varicolor">Polyrhachis varicolor</taxonomicName>
.
</emphasis>
A–C, Non-type dealate queen (TH21-WJT-003/2); D–F, non-type male (TH21-WJT-003/3); A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view.
</paragraph>
</caption>
<paragraph id="757E024BFFAAFF824EE9C560FDF6FBC8" blockId="16.[151,1436,1908,2043]" box="[199,581,1908,1934]" pageId="16" pageNumber="462">
<emphasis id="47B5DE59FFAAFF824EE9C560FE02FBC8" bold="true" box="[199,433,1908,1934]" pageId="16" pageNumber="462">Description of male</emphasis>
(
<figureCitation id="EDFA1ECEFFAAFF824FEEC560FD8FFBC8" box="[448,572,1908,1934]" captionStart="FIGURE 9" captionStartId="16.[152,255,1802,1826]" captionTargetBox="[151,1436,181,1778]" captionTargetId="figure-20@16.[151,1436,181,1778]" captionTargetPageId="16" captionText="FIGURE 9. Polyrhachis varicolor.A–C, Non-type dealate queen (TH21-WJT-003/2); D–F, non-type male (TH21-WJT-003/3); A, D, body in profile view; B, E, body in dorsal view; C, F, head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.7694560" httpUri="https://zenodo.org/record/7694560/files/figure.png" pageId="16" pageNumber="462">Fig. 9 D–F</figureCitation>
)
</paragraph>
<paragraph id="757E024BFFAAFF834EE9C58CFDA8FD5B" blockId="16.[151,1436,1908,2043]" lastBlockId="17.[151,1437,151,1077]" lastPageId="17" lastPageNumber="463" pageId="16" pageNumber="462">Head in full-face view longer than broad, but if including eyes, then distinctly broader than long. Clypeus in dorsal view broad, with transverse impression in its median portion, posteriorly clearly demarcated from frons, with posteromedian margin shallowly emarginated medially and anterior margin broadly convex. Mandible spatulate, weakly sclerotized, twice as long as broad with outer margin curved and apex bluntly pointed; masticatory margin short and edentate. Eye large, positioned at mid-length of lateral face of head; distance between mandibular base and anterior margin of eye (malar space) half as long as major diameter of eye. Antennal scape moderately long, as long as head width including eyes.</paragraph>
<paragraph id="757E024BFFABFF834EE9C333FB0EFDB3" blockId="17.[151,1437,151,1077]" pageId="17" pageNumber="463">Mesosoma in dorsal view much broader than head including eyes. Pronotum in dorsal view short (‘narrow’), almost entirely concealed by inflated mesoscutum. Mesoscutum with faint median line, without notauli; parapsidal line present in posterior half of scutum; scuto-scutellar sulcus prominent; mesoscutellum much narrower than mesoscutum, distinctly narrowed posteriad. Metanotum short (‘narrow’), indented, clearly defined from propodeum with deep furrow. Propodeum without separation of dorsum from declivity; spines absent. Petiole low and globular, in dorsal view as broad as long. Gaster rather weakly sclerotized, easily deformed after drying.</paragraph>
<paragraph id="757E024BFFABFF834EE9C3EBFCA8FFC7" blockId="17.[151,1437,151,1077]" pageId="17" pageNumber="463">
Entire body superficially and finely sculptured, weakly shiny. Short appressed hairs present over body surface; standing hairs confined to following parts: clypeus with two to three long hairs anteriorly; mandible with some long hairs of varying length on outer surface; standing hairs on maxillary and labial palpi numerous; mesoscutum without standing hairs; mesoscutellum without standing hairs but with sparse appressed hairs that are relatively short; a few standing hairs present in posterolateral area of propodeum and around propodeal spiracle; petiole with ca. 20 fine standing hairs dorsally, pair of standing hairs present on sternite posteriorly, and thick golden hairs anteriorly; gastral tergites generally lacking standing hairs except on apical tergites; gastral sternite with more standing hairs; coxae of all legs with sparse standing hairs. Eye without hairs that are recognized under high magnifications (&gt;100 x). Body brown to dark brown; mandible, legs and genitalia yellowish or light brown; maxillary and labial palpi yellowish. The male of
<taxonomicName id="B2C179C8FFABFF834FB3C157FD93FF1B" authorityName=": Kohout" authorityYear="2008" box="[413,544,835,861]" class="Insecta" family="Formicidae" genus="Polyrhachis" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="463" phylum="Arthropoda" rank="species" species="varicolor">
<emphasis id="47B5DE59FFABFF834FB3C157FD93FF1B" box="[413,544,835,861]" italics="true" pageId="17" pageNumber="463">P. varicolor</emphasis>
</taxonomicName>
is very similar to that of
<taxonomicName id="B2C179C8FFABFF834D1BC157FC7BFF1B" authorityName="Smith" authorityYear="1857" box="[821,968,835,861]" class="Insecta" family="Formicidae" genus="Polyrhachis" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="463" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="47B5DE59FFABFF834D1BC157FC7BFF1B" box="[821,968,835,861]" italics="true" pageId="17" pageNumber="463">P. flavicornis</emphasis>
</taxonomicName>
but is distinctly larger than the latter (HW
<quantity id="B239AFAEFFABFF834EB9C172FF48FFC7" box="[151,251,870,897]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="17" pageNumber="463" unit="mm" value="1.25">1.25 mm</quantity>
in this species; 0.95–1.00 mm in
<taxonomicName id="B2C179C8FFABFF834C5AC173FCB8FFC7" authorityName="Smith" authorityYear="1857" box="[628,779,871,897]" class="Insecta" family="Formicidae" genus="Polyrhachis" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="463" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="47B5DE59FFABFF834C5AC173FD36FFC6" box="[628,645,871,896]" italics="true" pageId="17" pageNumber="463">P</emphasis>
.
<emphasis id="47B5DE59FFABFF834CBDC173FCB8FFC7" box="[659,779,871,897]" italics="true" pageId="17" pageNumber="463">flavicornis</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="757E024BFFABFF834EE9C19EFEB3FFAB" blockId="17.[151,1437,151,1077]" pageId="17" pageNumber="463">
<emphasis id="47B5DE59FFABFF834EE9C19EFE99FFE2" bold="true" box="[199,298,906,932]" pageId="17" pageNumber="463">Habitat.</emphasis>
Nests of
<taxonomicName id="B2C179C8FFABFF834FB8C19FFDAAFFE3" authorityName=": Kohout" authorityYear="2008" box="[406,537,907,933]" class="Insecta" family="Formicidae" genus="Polyrhachis" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="463" phylum="Arthropoda" rank="species" species="varicolor">
<emphasis id="47B5DE59FFABFF834FB8C19FFDAAFFE3" box="[406,537,907,933]" italics="true" pageId="17" pageNumber="463">P. varicolor</emphasis>
</taxonomicName>
were found on the underside of plant leaves or between leaves in dry evergreen forests and evergreen forests (
<quantity id="B239AFAEFFABFF834FC7C1BAFD82FF8F" box="[489,561,942,969]" metricMagnitude="0" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="17" pageNumber="463" unit="m" value="3.5" valueMax="5.0" valueMin="2.0">2–5 m</quantity>
above the ground). Silk, carton and other materials were used in constructing the nests.
</paragraph>
</subSubSection>
<subSubSection id="3DDB51C0FFABFF834EE9C1E2FD4FF857" box="[199,764,1014,1041]" pageId="17" pageNumber="463" type="distribution">
<paragraph id="757E024BFFABFF834EE9C1E2FD4FF857" blockId="17.[151,1437,151,1077]" box="[199,764,1014,1041]" pageId="17" pageNumber="463">
<emphasis id="47B5DE59FFABFF834EE9C1E2FED2F856" bold="true" box="[199,353,1014,1040]" pageId="17" pageNumber="463">Distribution.</emphasis>
<collectingCountry id="0DD642DBFFABFF834F49C1E3FE78F857" box="[359,459,1015,1041]" name="Thailand" pageId="17" pageNumber="463">Thailand</collectingCountry>
and
<collectingCountry id="0DD642DBFFABFF834C2FC1E3FDC0F857" box="[513,627,1015,1041]" name="Singapore" pageId="17" pageNumber="463">Singapore</collectingCountry>
(
<figureCitation id="EDFA1ECEFFABFF834CACC1E3FD5DF857" box="[642,750,1015,1041]" captionStart="FIGURE 10" captionStartId="17.[152,255,1855,1879]" captionTargetBox="[155,1433,1101,1832]" captionTargetId="figure-427@17.[154,1433,1101,1832]" captionTargetPageId="17" captionText="FIGURE 10. Distribution map of Polyrhachis flavicornis species group. A) P. constructor and P. longipilosa sp. nov. B) P. flavicornis and P. varicolor." figureDoi="http://doi.org/10.5281/zenodo.7694564" httpUri="https://zenodo.org/record/7694564/files/figure.png" pageId="17" pageNumber="463">Fig. 10 B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3DDB51C0FFABFF834EE9C60FFAA6F873" box="[199,1301,1051,1077]" pageId="17" pageNumber="463" type="discussion">
<paragraph id="757E024BFFABFF834EE9C60FFAA6F873" blockId="17.[151,1437,151,1077]" box="[199,1301,1051,1077]" pageId="17" pageNumber="463">
<emphasis id="47B5DE59FFABFF834EE9C60FFE88F873" bold="true" box="[199,315,1051,1077]" pageId="17" pageNumber="463">Remarks.</emphasis>
<taxonomicName id="B2C179C8FFABFF834F6CC60FFD8BF873" authorityName=": Kohout" authorityYear="2008" box="[322,568,1051,1077]" class="Insecta" family="Formicidae" genus="Polyrhachis" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="463" phylum="Arthropoda" rank="species" species="varicolor">
<emphasis id="47B5DE59FFABFF834F6CC60FFD8BF873" box="[322,568,1051,1077]" italics="true" pageId="17" pageNumber="463">Polyrhachis varicolor</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="B2C179C8FFABFF834CE4C60FFCEDF873" authorityName="Smith" authorityYear="1857" box="[714,862,1051,1077]" class="Insecta" family="Formicidae" genus="Polyrhachis" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="463" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="47B5DE59FFABFF834CE4C60FFCEDF873" box="[714,862,1051,1077]" italics="true" pageId="17" pageNumber="463">P. flavicornis</emphasis>
</taxonomicName>
. See under ‘Remarks’ of
<taxonomicName id="B2C179C8FFABFF834A55C60FFABCF873" authorityName="Smith" authorityYear="1857" box="[1147,1295,1051,1077]" class="Insecta" family="Formicidae" genus="Polyrhachis" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="463" phylum="Arthropoda" rank="species" species="flavicornis">
<emphasis id="47B5DE59FFABFF834A55C60FFABCF873" box="[1147,1295,1051,1077]" italics="true" pageId="17" pageNumber="463">P. flavicornis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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