133 lines
13 KiB
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133 lines
13 KiB
XML
<document id="40D4DE4E071602F86A6AA1E8E8D21748" ID-CLB-Dataset="46698" ID-DOI="10.11646/zootaxa.3753.6.3" ID-GBIF-Dataset="715c7010-c4b2-4910-b707-62a84ba9a6d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="224299" ID-ZooBank="8DF3EFAA-F8CC-4D4C-A830-D07D76C2D1C0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460366938684" checkinUser="plazi" docAuthor="Fragoso, Carlos & Rojas, Patricia" docDate="2014" docId="FD1F87899A51FF8A1DECAD1FFD26FA0D" docLanguage="en" docName="zt03753p572.pdf" docOrigin="Zootaxa 3753 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ramiellona lasiura Graff 1957" docType="treatment" docVersion="11" lastPageNumber="568" masterDocId="0126FFF19A42FF991D7BAE78FFDBFFB9" masterDocTitle="New species and records of the earthworm genus Ramiellona (Annelida, Oligochaeta, Acanthodrilidae) from southern Mexico and Guatemala" masterLastPageNumber="572" masterPageNumber="549" pageNumber="568" updateTime="1698295306276" updateUser="ExternalLinkService">
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<mods:title id="4DCAFA1E3910323C8FC554F7484AC5C5">New species and records of the earthworm genus Ramiellona (Annelida, Oligochaeta, Acanthodrilidae) from southern Mexico and Guatemala</mods:title>
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<mods:namePart id="D76B22E6C65B140E21F5987BBF22AC31">Fragoso, Carlos</mods:namePart>
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<mods:namePart id="FFC7A4831CAC14BAC993548F719BB188">Rojas, Patricia</mods:namePart>
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<mods:title id="D6FF98F1AB2D5AD88F8A6DC71542BE05">Zootaxa</mods:title>
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<mods:date id="41031167C3351A21E79BBB97C6B0C597">2014</mods:date>
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<mods:number id="D13D09F0A98296F60220814928979A20">3753</mods:number>
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<mods:number id="DDBA8103601445E4FDD9E8934778A16A">6</mods:number>
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<mods:identifier id="A9ABE969881B970A3CE70B45EA4446C5" type="DOI">10.11646/zootaxa.3753.6.3</mods:identifier>
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<treatment id="FD1F87899A51FF8A1DECAD1FFD26FA0D" ID-DOI="http://doi.org/10.5281/zenodo.5616751" ID-GBIF-Taxon="119411005" ID-Zenodo-Dep="5616751" LSID="urn:lsid:plazi:treatment:FD1F87899A51FF8A1DECAD1FFD26FA0D" httpUri="http://treatment.plazi.org/id/FD1F87899A51FF8A1DECAD1FFD26FA0D" lastPageNumber="568" pageId="19" pageNumber="568">
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<subSubSection id="3DAC65149A51FF8A1DECAD1FFDF3FC3B" box="[151,552,871,898]" pageId="19" pageNumber="568" type="nomenclature">
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<paragraph id="7509369F9A51FF8A1DECAD1FFDF3FC3B" blockId="19.[151,552,871,898]" box="[151,552,871,898]" pageId="19" pageNumber="568">
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<heading id="2E4181F39A51FF8A1DECAD1FFDF3FC3B" bold="true" box="[151,552,871,898]" fontSize="11" level="1" pageId="19" pageNumber="568" reason="1">
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<taxonomicName id="B2B64D1C9A51FF8A1DECAD1FFDF3FC3B" authority="Graff, 1957" authorityName="Graff" authorityYear="1957" box="[151,552,871,898]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="19" pageNumber="568" phylum="Annelida" rank="species" species="lasiura">
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<emphasis id="47C2EA8D9A51FF8A1DECAD1FFDF3FC3B" bold="true" box="[151,552,871,898]" pageId="19" pageNumber="568">
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<emphasis id="47C2EA8D9A51FF8A1DECAD1FFEA5FC38" bold="true" box="[151,382,871,897]" italics="true" pageId="19" pageNumber="568">Ramiellona lasiura</emphasis>
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(
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<bibRefCitation id="11274B6E9A51FF8A1CF5AD10FDFBFC3B" author="Graff" box="[398,544,872,898]" pageId="19" pageNumber="568" refString="Graff, O. (1957) Regenwurmer aus El Salvador (Oligochaeta). Senckenbergiana Biologica, 38, 115 - 143." type="journal article" year="1957">Graff, 1957</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="3DAC65149A51FF8A1DECADD6FD26FA0D" pageId="19" pageNumber="568" type="reference_group">
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<paragraph id="7509369F9A51FF8A1DECADD6FDCFFC5A" blockId="19.[151,837,942,995]" pageId="19" pageNumber="568">
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<treatmentCitationGroup id="55A611B19A51FF8A1DECADD6FDCFFC5A" pageId="19" pageNumber="568">
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<taxonomicName id="B2B64D1C9A51FF8A1DECADD6FD1CFC7D" authority="Graff, 1957: 129" authorityName="Graff" authorityPageNumber="129" authorityYear="1957" box="[151,711,942,964]" class="Clitellata" family="Megascolecidae" genus="Howascolex" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opisthopora" pageId="19" pageNumber="568" phylum="Annelida" rank="species" species="lasiurus" subGenus="Graceevelynia">
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<emphasis id="47C2EA8D9A51FF8A1DECADD6FDCDFC7D" box="[151,534,942,964]" italics="true" pageId="19" pageNumber="568">Howascolex (Graceevelynia) lasiurus</emphasis>
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<treatmentCitation id="F417108E9A51FF8A1F67ADD6FD1CFC7D" author="Graff" box="[540,711,942,964]" page="129" pageId="19" pageNumber="568" year="1957">
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<bibRefCitation id="11274B6E9A51FF8A1F67ADD6FD4FFC7D" author="Graff" box="[540,660,942,964]" pageId="19" pageNumber="568" refString="Graff, O. (1957) Regenwurmer aus El Salvador (Oligochaeta). Senckenbergiana Biologica, 38, 115 - 143." type="journal article" year="1957">Graff, 1957</bibRefCitation>
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: 129
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</treatmentCitation>
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</taxonomicName>
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. Figs. 16-19
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<taxonomicName id="B2B64D1C9A51FF8A1DECADB5FDCBFC5A" authority="Gates 1962: 215" authorityName="Gates" authorityPageNumber="215" authorityYear="1962" box="[151,528,973,995]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="19" pageNumber="568" phylum="Annelida" rank="species" species="lasiura">
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<emphasis id="47C2EA8D9A51FF8A1DECADB5FE81FC5A" box="[151,346,973,995]" italics="true" pageId="19" pageNumber="568">Ramiellona lasiura</emphasis>
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:
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<treatmentCitation id="F417108E9A51FF8A1C1CADB5FDCBFC5A" author="Gates" box="[359,528,973,995]" page="215" pageId="19" pageNumber="568" year="1962">
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<bibRefCitation id="11274B6E9A51FF8A1C1CADB5FE05FC5A" author="Gates" box="[359,478,973,995]" pageId="19" pageNumber="568" refString="Gates, G. (1962) On some earthworms of Eisen's collections. Proceedings of the California Academy of Sciences, 4 (31), 185 - 225." type="journal article" year="1962">Gates 1962</bibRefCitation>
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: 215
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="7509369F9A51FF8A1DECAA68FBBBFB01" blockId="19.[151,1436,1039,1461]" pageId="19" pageNumber="568">
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<bibRefCitation id="11274B6E9A51FF8A1DECAA68FEEBFB91" author="Gates" box="[151,304,1039,1064]" pageId="19" pageNumber="568" refString="Gates, G. (1962) On some earthworms of Eisen's collections. Proceedings of the California Academy of Sciences, 4 (31), 185 - 225." type="journal article" year="1962">Gates (1962)</bibRefCitation>
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transferred this species to
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<taxonomicName id="B2B64D1C9A51FF8A1FF8AA77FCDFFB91" box="[643,772,1039,1064]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="19" pageNumber="568" phylum="Annelida" rank="genus">
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<emphasis id="47C2EA8D9A51FF8A1FF8AA77FCDFFB91" box="[643,772,1039,1064]" italics="true" pageId="19" pageNumber="568">Ramiellona</emphasis>
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</taxonomicName>
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, considering that it has only meronephridia and lacks holonephridia. This author pointed out that in the original description several characters were not indicated or wrongly placed. Thanks to the kindness of Dr. J. Römbke, in 1987 the first author of this paper had the opportunity to briefly see the
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<typeStatus id="AA0D883D9A51FF8A1C2CAA04FE62FB2C" box="[343,441,1148,1173]" pageId="19" pageNumber="568" type="holotype">holotype</typeStatus>
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of this species (Naturmuseum Senckenberg, Frankfurt, Cat No. 2783). Accordingly, with respect to the original description the species diagnosis should be modified as follows:
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</paragraph>
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<paragraph id="7509369F9A51FF8A1DBCAABCFD26FA0D" blockId="19.[151,1436,1039,1461]" pageId="19" pageNumber="568">
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One gizzard in segment
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<emphasis id="47C2EA8D9A51FF8A1CAFAABCFE39FB64" bold="true" box="[468,482,1220,1245]" pageId="19" pageNumber="568">5</emphasis>
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(not in
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<emphasis id="47C2EA8D9A51FF8A1F43AABCFD9EFB64" bold="true" box="[568,581,1220,1245]" pageId="19" pageNumber="568">6</emphasis>
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as in the original description). Calciferous glands in segments
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<emphasis id="47C2EA8D9A51FF8A187BAABCFAE3FB64" bold="true" box="[1280,1336,1220,1245]" pageId="19" pageNumber="568">8–12</emphasis>
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(
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<quantity id="B24E9B7A9A51FF8A1833AABCFA47FB64" box="[1352,1436,1220,1245]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.302" metricValueMin="2.286" pageId="19" pageNumber="568" unit="in" value="11.0" valueMax="13.0" valueMin="9.0">
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<emphasis id="47C2EA8D9A51FF8A1833AABCFA5BFB64" bold="true" box="[1352,1408,1220,1245]" pageId="19" pageNumber="568">9–13</emphasis>
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in
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</quantity>
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o.d.). Only one pair of seminal vesicles in segment
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<emphasis id="47C2EA8D9A51FF8A1F8EAA9FFCCAFAB9" bold="true" box="[757,785,1255,1280]" pageId="19" pageNumber="568">12</emphasis>
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(doubtful in
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<quantity id="B24E9B7A9A51FF8A1ED5AA9FFC32FAB9" box="[942,1001,1255,1280]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="19" pageNumber="568" unit="in" value="11.0">
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<emphasis id="47C2EA8D9A51FF8A1ED5AA9FFC12FAB9" bold="true" box="[942,969,1255,1280]" pageId="19" pageNumber="568">11</emphasis>
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in
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</quantity>
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o.d.) and one pair of iridescent male funnels in
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<emphasis id="47C2EA8D9A51FF8A1C6AAB74FEF0FA9C" bold="true" box="[273,299,1292,1317]" pageId="19" pageNumber="568">11</emphasis>
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(not seen in o.d.). It was not possible to assert the position and number of hearts because they had been removed; however, since the position of the gizzard is in
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<emphasis id="47C2EA8D9A51FF8A1E1DAB57FCAFFAF1" bold="true" box="[870,884,1327,1352]" pageId="19" pageNumber="568">5</emphasis>
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and not in
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<emphasis id="47C2EA8D9A51FF8A1E82AB57FBDCFAF1" bold="true" box="[1017,1031,1327,1352]" pageId="19" pageNumber="568">6</emphasis>
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as originally described, it is highly probable that also the hearts are located one segment more anteriorly, i.e. not in
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<emphasis id="47C2EA8D9A51FF8A1963AB2CFB8BFAD4" bold="true" box="[1048,1104,1364,1389]" pageId="19" pageNumber="568">9–13</emphasis>
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but in
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<emphasis id="47C2EA8D9A51FF8A19E5AB2CFB0DFAD4" bold="true" box="[1182,1238,1364,1389]" pageId="19" pageNumber="568">8–12</emphasis>
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. As
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<bibRefCitation id="11274B6E9A51FF8A1871AB2CFA47FAD4" author="Gates" box="[1290,1436,1364,1389]" pageId="19" pageNumber="568" refString="Gates, G. (1962) On some earthworms of Eisen's collections. Proceedings of the California Academy of Sciences, 4 (31), 185 - 225." type="journal article" year="1962">Gates (1962)</bibRefCitation>
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already suggested, we found that
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<taxonomicName id="B2B64D1C9A51FF8A1F69AB01FD5AFA29" box="[530,641,1399,1424]" class="Clitellata" family="Octochaetidae" genus="Ramiellona" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="19" pageNumber="568" phylum="Annelida" rank="species" species="lasiura">
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<emphasis id="47C2EA8D9A51FF8A1F69AB01FD5AFA29" box="[530,641,1399,1424]" italics="true" pageId="19" pageNumber="568">R. lasiura</emphasis>
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</taxonomicName>
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is a metandric species with its gizzard placed in the usual position for this genus and with last hearts probably in segment
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<emphasis id="47C2EA8D9A51FF8A1FA1ABE4FD2DFA0C" bold="true" box="[730,758,1436,1461]" pageId="19" pageNumber="568">12</emphasis>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |