161 lines
20 KiB
XML
161 lines
20 KiB
XML
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<mods:title id="0AF0AACA9EE8B5A73F9A8D6227DF7DF1">Revision of the Early Ordovician (late Tremadocian; Stairsian) cheirurid trilobite Tesselacauda Ross, with species from the Great Basin, western USA</mods:title>
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<paragraph id="73433695F6658D10FF139BE1FD8DF991" blockId="16.[151,639,153,215]" box="[151,639,153,180]" pageId="16" pageNumber="217">
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<heading id="280B81F9F6658D10FF139BE1FD8DF991" bold="true" box="[151,639,153,180]" fontSize="11" level="1" pageId="16" pageNumber="217" reason="1">
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<emphasis id="4188EA87F6658D10FF139BE1FD8DF991" bold="true" box="[151,639,153,180]" pageId="16" pageNumber="217">
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<emphasis id="4188EA87F6658D10FF139BE1FEB4F996" bold="true" box="[151,326,153,180]" italics="true" pageId="16" pageNumber="217">
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<taxonomicName id="B4FC4D16F6658D10FF139BE1FEC8F996" ID-CoL="646XF" authorityName="Ross" authorityYear="1951" box="[151,314,153,180]" class="Trilobita" family="Pliomeridae" genus="Tesselacauda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Phacopida" pageId="16" pageNumber="217" phylum="Arthropoda" rank="genus">Tesselacauda</taxonomicName>
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?
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</emphasis>
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<taxonomicName id="B4FC4D16F6658D10FEC99BE1FD8DF991" authority="Kobayashi, 1955" authorityName="Kobayashi" authorityYear="1955" box="[333,639,153,180]" class="Trilobita" family="Pliomeridae" genus="Tesselacauda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Phacopida" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="flabella">
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<emphasis id="4188EA87F6658D10FEC99BE1FE5AF996" bold="true" box="[333,424,153,180]" italics="true" pageId="16" pageNumber="217">flabella</emphasis>
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<bibRefCitation id="176D4B64F6658D10FE349BE1FD8DF991" author="Kobayashi, T." box="[432,639,153,179]" pageId="16" pageNumber="217" pagination="355 - 493" refId="ref15931" refString="Kobayashi, T. (1955) The Ordovician fossils of the McKay Group in British Columbia western Canada, with a note on the early Ordovician palaeogeography. Journal of the Faculty of Science, Tokyo University, Section 2, 9, 355 - 493." type="journal article" year="1955">Kobayashi, 1955</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="73433695F6658D10FF139BC5FE46F9F5" blockId="16.[151,639,153,215]" box="[151,436,189,215]" pageId="16" pageNumber="217">
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(
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<figureCitation id="EBC72A10F6658D10FF1B9BC5FF0CF9F5" box="[159,254,189,215]" captionStart="PLATE 14" captionStartId="46.[151,242,1268,1295]" captionTargetBox="[153,886,1215,1228]" captionTargetId="graphics@46.[153,886,1215,1229]" captionText="PLATE 14. 1–16, 19. Tesselacauda kriegerae n. sp., from Section G 26.6–27.0 m and Section C 111.6 m, both Fillmore Formation (upper Tremadocian; Stairsian; Bearriverops alsacharovi Zone), Ibex area, Millard County, western Utah, USA. 1, 2, 4. Cranidium, SUI 147602, dorsal, right lateral, and anterior views, x17 (G 27.0 m). 3. Right librigena, SUI 147603, external view, x15 (G 26.6 m). 5, 8, 10. Right librigena, SUI 147604, internal, external, and ventrolateral views, x12 (G 26.6 m). 6. Left librigena, SUI 147605, external view, x12 (G 26.6 m). 7. Right librigena, SUI 147606, external view, x15 (C 111.6 m). 9. Left librigena, SUI 147607, external view, x15 (G 26.6 m). 11. Left librigena, SUI 147608, external view, x15 (G 26.6 m). 12, 14, 16. Left librigena, SUI 147609, external, internal, and ventrolateral views, x12 (G 26.6 m). 13, 15. Right librigena, SUI 147610, external and ventrolateral views, x15 (G 26.6 m). 19. Left librigena, SUI 147611, external view, x12 (G 26.6 m). 17, 18, 20. Tesselacauda? flabella Kobayashi, 1955, from Section G 26.6 m, Fillmore Formation (upper Tremadocian; Stairsian; Bearriverops alsacharovi Zone), Ibex area, Millard County, western Utah, USA. 17, 18. Pygidium, SUI 147612, dorsal and left lateral views, x6. 20. Pygidium, SUI 147613, dorsal view, x12." figureDoi="http://doi.org/10.5281/zenodo.3446133" httpUri="https://zenodo.org/record/3446133/files/figure.png" pageId="16" pageNumber="217">Plate 14</figureCitation>
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, figs 17, 18, 20)
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</paragraph>
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</subSubSection>
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<subSubSection id="3BE6651EF6658D10FF439B82FAF6F810" pageId="16" pageNumber="217" type="description">
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<paragraph id="73433695F6658D10FF439B82FC6EF831" blockId="16.[199,1284,250,306]" box="[199,924,250,275]" pageId="16" pageNumber="217">
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1955
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<taxonomicName id="B4FC4D16F6658D10FED39B83FD5AF831" authority="Kobayashi" authorityName="Kobayashi" authorityYear="1955" box="[343,680,250,275]" class="Trilobita" family="Pliomeridae" genus="Tesselacauda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Phacopida" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="flabella">
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<emphasis id="4188EA87F6658D10FED39B83FDC0F831" box="[343,562,251,275]" italics="true" pageId="16" pageNumber="217">Tesselacauda flabella</emphasis>
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Kobayashi
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</taxonomicName>
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, p. 417, pl. 2, figs 8a, b.
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</paragraph>
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<paragraph id="73433695F6658D10FF439A62FAF6F810" blockId="16.[199,1284,250,306]" box="[199,1284,281,306]" pageId="16" pageNumber="217">
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1989
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<taxonomicName id="B4FC4D16F6658D10FED39A62FD5BF810" authority="Kobayashi" authorityName="Kobayashi" authorityYear="1955" box="[343,681,281,306]" class="Trilobita" family="Pliomeridae" genus="Tesselacauda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Phacopida" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="flabella">
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<emphasis id="4188EA87F6658D10FED39A62FDC0F810" box="[343,562,282,306]" italics="true" pageId="16" pageNumber="217">Tesselacauda flabella</emphasis>
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Kobayashi
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</taxonomicName>
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; Dean, p. 17, pl. 2, fig. 12 (
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<emphasis id="4188EA87F6658D10FC3B9A62FC17F813" box="[959,997,282,305]" italics="true" pageId="16" pageNumber="217">non</emphasis>
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pl. 2, figs 2, 4, 7, 8, 10, 11).
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</paragraph>
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</subSubSection>
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<subSubSection id="3BE6651EF6658D10FF139A20FC54FDE4" pageId="16" pageNumber="217" type="materials_examined">
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<paragraph id="73433695F6658D10FF139A20FEC1F898" blockId="16.[151,1437,344,1222]" pageId="16" pageNumber="217">
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<emphasis id="4188EA87F6658D10FF139A20FEF4F851" bold="true" box="[151,262,344,371]" pageId="16" pageNumber="217">Material.</emphasis>
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Assigned specimens
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<collectionCode id="15EDAE50F6658D10FD8C9A21FDC7F851" box="[520,565,345,371]" country="USA" httpUri="http://grbio.org/cool/fx78-973d" name="The University of Iowa (formerly State University of Iowa)" pageId="16" pageNumber="217" type="University or college">SUI</collectionCode>
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147612 and 147613, from Section G 26.6 m, Fillmore Formation (upper Tremadocian; Stairsian;
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<taxonomicName id="B4FC4D16F6658D10FE2B9A05FCE4F8B5" authority="Zone" authorityName="Zone" box="[431,790,380,407]" class="Trilobita" family="Dimeropygidae" genus="Bearriverops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Proetida" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="alsacharovi">
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<emphasis id="4188EA87F6658D10FE2B9A05FD3CF8B4" box="[431,718,380,406]" italics="true" pageId="16" pageNumber="217">Bearriverops alsacharovi</emphasis>
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Zone
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</taxonomicName>
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), southern Confusion Range, Ibex area, Millard County, western
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<collectingRegion id="B138F877F6658D10FF719AD8FEDDF898" box="[245,303,416,442]" country="United States of America" name="Utah" pageId="16" pageNumber="217">Utah</collectingRegion>
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.
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</paragraph>
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<paragraph id="73433695F6658D10FF439ABAFC1CFBDB" blockId="16.[151,1437,344,1222]" pageId="16" pageNumber="217">
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<emphasis id="4188EA87F6658D10FF439ABAFEBEF8FF" bold="true" box="[199,332,450,477]" pageId="16" pageNumber="217">Discussion.</emphasis>
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<bibRefCitation id="176D4B64F6658D10FED19ABBFDEBF8FF" author="Kobayashi, T." box="[341,537,450,477]" pageId="16" pageNumber="217" pagination="355 - 493" refId="ref15931" refString="Kobayashi, T. (1955) The Ordovician fossils of the McKay Group in British Columbia western Canada, with a note on the early Ordovician palaeogeography. Journal of the Faculty of Science, Tokyo University, Section 2, 9, 355 - 493." type="journal article" year="1955">Kobayashi (1955</bibRefCitation>
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, p. 417, pl. 2, figs 8a, b) established this species on the basis of a single illustrated pygidium from the McKay Group of southeastern
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<collectingRegion id="B138F877F6658D10FCA79A9FFC18FB23" box="[803,1002,487,513]" country="Canada" name="British Columbia" pageId="16" pageNumber="217">British Columbia</collectingRegion>
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,
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<collectingCountry id="0BEB7605F6658D10FC7C9A9FFBA2FB23" box="[1016,1104,487,513]" name="Canada" pageId="16" pageNumber="217">Canada</collectingCountry>
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.
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<bibRefCitation id="176D4B64F6658D10FBD89A9FFB16FB23" author="Dean, W. T." box="[1116,1252,487,513]" pageId="16" pageNumber="217" pagination="1 - 141" refId="ref15210" refString="Dean, W. T. (1989) Trilobites from the Survey Peak, Outram and Skoki formations (Upper Cambrian-Lower Ordovician) at Wilcox Pass, Jasper National Park, Alberta. Geological Survey of Canada Bulletin, 389, 1 - 141. https: // doi. org / 10.4095 / 127615" type="journal article" year="1989">Dean (1989</bibRefCitation>
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, pl. 2, figs 2, 4, 7, 8, 10–12) reillustrated this
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<typeStatus id="AC478837F6658D10FE619972FDB5FB06" box="[485,583,522,548]" pageId="16" pageNumber="217" type="holotype">holotype</typeStatus>
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and assigned three pygidia (two fragmentary) and a fragmentary cranidium from the middle member of the Survey Peak Formation, Wilcox Pass, Jasper National Park,
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<collectingRegion id="B138F877F6658D10FB109956FB1EFB6A" box="[1172,1260,558,584]" country="Canada" name="Alberta" pageId="16" pageNumber="217">Alberta</collectingRegion>
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. While possibly representing a related species, Dean’s specimens are obviously much less effaced than Kobayashi’s
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<typeStatus id="AC478837F6658D10FB609929FAB4FB49" box="[1252,1350,593,619]" pageId="16" pageNumber="217" type="holotype">holotype</typeStatus>
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and are unlikely to be conspecific. In fact, the Survey Peak specimens are not effaced at all, and the effacement of both the posterior axial and ring furrows and most of the pleural and interpleural furrows is the most striking feature of the
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<typeStatus id="AC478837F6658D10FF1399C3FF0FFBF7" box="[151,253,699,725]" pageId="16" pageNumber="217" type="holotype">holotype</typeStatus>
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. The affinity of the cranidial fragment associated by Dean is impossible to assess; it appears to represent a cheirurid, but consists only of part of a glabella and part of an occipital ring.
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</paragraph>
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<paragraph id="73433695F6658D10FF43987BFCFDFAD3" blockId="16.[151,1437,344,1222]" pageId="16" pageNumber="217">
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Two pygidia from the
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<taxonomicName id="B4FC4D16F6658D10FE39987BFCEAFA3F" authority="Zone" authorityName="Zone" box="[445,792,770,797]" class="Trilobita" family="Dimeropygidae" genus="Bearriverops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Proetida" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="alsacharovi">
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<emphasis id="4188EA87F6658D10FE39987BFD2AFA3E" box="[445,728,770,796]" italics="true" pageId="16" pageNumber="217">Bearriverops alsacharovi</emphasis>
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Zone
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</taxonomicName>
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appear much closer in morphology to
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<bibRefCitation id="176D4B64F6658D10FB45987AFA6EFA3F" author="Kobayashi, T." box="[1217,1436,770,797]" pageId="16" pageNumber="217" pagination="355 - 493" refId="ref15931" refString="Kobayashi, T. (1955) The Ordovician fossils of the McKay Group in British Columbia western Canada, with a note on the early Ordovician palaeogeography. Journal of the Faculty of Science, Tokyo University, Section 2, 9, 355 - 493." type="journal article" year="1955">Kobayashi’s (1955)</bibRefCitation>
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<typeStatus id="AC478837F6658D10FF13985EFF0FFA62" box="[151,253,806,832]" pageId="16" pageNumber="217" type="holotype">holotype</typeStatus>
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. A smaller specimen preserving the anterior part of the axis and left pleural region (Pl. 14, fig. 20) is nearly identical to the
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<typeStatus id="AC478837F6658D10FEC79831FE5BFA41" box="[323,425,841,867]" pageId="16" pageNumber="217" type="holotype">holotype</typeStatus>
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. A specimen more than twice the size preserving most of the left pleural region (Pl. 14, figs 17, 18) seems conspecific, and, if so, shows that the pygidium became increasingly effaced with size. Unfortunately the species is extremely rare, as these are the only specimens found and no prospective cranidia were recovered. They are of some potential importance, however, in that they may anchor some of
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<bibRefCitation id="176D4B64F6658D10FBA298CCFAF3FAEC" author="Kobayashi, T." box="[1062,1281,947,974]" pageId="16" pageNumber="217" pagination="355 - 493" refId="ref15931" refString="Kobayashi, T. (1955) The Ordovician fossils of the McKay Group in British Columbia western Canada, with a note on the early Ordovician palaeogeography. Journal of the Faculty of Science, Tokyo University, Section 2, 9, 355 - 493." type="journal article" year="1955">Kobayashi’s (1955)</bibRefCitation>
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species to the trilobite zonal scheme presented by
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<bibRefCitation id="176D4B64F6658D10FDAD98AFFCF9FAD3" author="Adrain, J. M. & Karim, T. S. & Westrop, S. R. & Landing, E." box="[553,779,983,1009]" pageId="16" pageNumber="217" pagination="167 - 214" refId="ref14639" refString="Adrain, J. M., Karim, T. S., Westrop, S. R. & Landing, E. (2014) Trilobite biostratigraphy of the Stairsian Stage (upper Tremadocian) of the Ibexian Series, Lower Ordovician, western United States. Memoirs of the Association of Australasian Palaeontologists, 45, 167 - 214." type="journal article" year="2014">
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Adrain
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<emphasis id="4188EA87F6658D10FDFA98A0FD44FAD3" box="[638,694,983,1009]" italics="true" pageId="16" pageNumber="217">et al.</emphasis>
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(2014)
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</bibRefCitation>
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.
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</paragraph>
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<paragraph id="73433695F6658D10FF439883FC54FDE4" blockId="16.[151,1437,344,1222]" pageId="16" pageNumber="217">
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So little is known of the species that its affinities are impossible to determine with any confidence. The pygidia are broadly comparable to those of species of
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<taxonomicName id="B4FC4D16F6658D10FD1B9F66FCC7FD1A" authorityName="Ross" authorityYear="1951" box="[671,821,1054,1080]" class="Trilobita" family="Pliomeridae" genus="Tesselacauda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Phacopida" pageId="16" pageNumber="217" phylum="Arthropoda" rank="genus">
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||
<emphasis id="4188EA87F6658D10FD1B9F66FCC7FD1A" box="[671,821,1054,1080]" italics="true" pageId="16" pageNumber="217">Tesselacauda</emphasis>
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||
</taxonomicName>
|
||
, but the smaller silicified example (Pl. 14, fig. 20) appears to show the pleural region of the third segment with an expressed pleural furrow. This is absent from all large pygidia assigned to
|
||
<taxonomicName id="B4FC4D16F6658D10FEF29F1DFDFEFD5D" authorityName="Ross" authorityYear="1951" box="[374,524,1125,1151]" class="Trilobita" family="Pliomeridae" genus="Tesselacauda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Phacopida" pageId="16" pageNumber="217" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4188EA87F6658D10FEF29F1DFDFEFD5D" box="[374,524,1125,1151]" italics="true" pageId="16" pageNumber="217">Tesselacauda</emphasis>
|
||
</taxonomicName>
|
||
herein. It is expressed on a single specimen, a very small pygidium assigned to
|
||
<taxonomicName id="B4FC4D16F6658D10FA0C9F1EFEF0FD81" authorityName="Adrain & Karim" authorityYear="2019" class="Trilobita" family="Pliomeridae" genus="Tesselacauda" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Phacopida" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="kriegerae">
|
||
<emphasis id="4188EA87F6658D10FA0C9F1EFEF0FD81" italics="true" pageId="16" pageNumber="217">T. kriegerae</emphasis>
|
||
</taxonomicName>
|
||
(Pl. 18, fig. 28). Much more information would be required to meaningfully evaluate the taxon, but it is documented here because of its possible biostratigraphic significance.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |