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<document id="6A56BAC2136E5BC4DF2789B5AA46812D" ID-DOI="10.5281/zenodo.185321" ID-GBIF-Dataset="849ca01a-84e7-470f-b75a-24617a9990b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="185321" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459981305554" checkinUser="plazi" docAuthor="Frick, Holger &amp; Muff, Patrick" docDate="2009" docId="FB3EE035970CFFD9ECA92EA0FACEF8D0" docLanguage="en" docName="zt01982p037.pdf" docOrigin="Zootaxa 1982" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Caracladus tsurusakii Saito 1988" docType="treatment" docVersion="14" lastPageNumber="18" masterDocId="0707984D971DFFC8EC3E2E37FFA5FF8C" masterDocTitle="Revision of the genus Caracladus with the description of Caracladus zamoniensis spec. nov. (Araneae, Linyphiidae, Erigoninae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="18" updateTime="1698231856956" updateUser="plazi">
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<mods:title id="10F0D79F4DE634888226754B9CD5C62D">Revision of the genus Caracladus with the description of Caracladus zamoniensis spec. nov. (Araneae, Linyphiidae, Erigoninae)</mods:title>
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<mods:namePart id="139044928273737A03B55697D4987C26">Frick, Holger</mods:namePart>
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<mods:namePart id="0599F5EEE5F87CFB4A68B6656E180CC0">Muff, Patrick</mods:namePart>
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<mods:date id="E5D7EC7FC9DCEE89D73123BE88777322">2009</mods:date>
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<mods:number id="CA235B6CEC42BEBCE2BC2D42636BEB5D">1982</mods:number>
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<treatment id="FB3EE035970CFFD9ECA92EA0FACEF8D0" ID-DOI="http://doi.org/10.5281/zenodo.4426012" ID-GBIF-Taxon="119378667" ID-Zenodo-Dep="4426012" LSID="urn:lsid:plazi:treatment:FB3EE035970CFFD9ECA92EA0FACEF8D0" httpUri="http://treatment.plazi.org/id/FB3EE035970CFFD9ECA92EA0FACEF8D0" lastPageNumber="18" pageId="17" pageNumber="18">
<subSubSection id="3B8D02A8970CFFD9ECA92EA0FE88FF58" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="73285123970CFFD9ECA92EA0FD94FF3D" blockId="17.[151,561,151,212]" box="[151,561,151,177]" pageId="17" pageNumber="18">
<heading id="2860E64F970CFFD9ECA92EA0FD94FF3D" bold="true" box="[151,561,151,177]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
<taxonomicName id="B4972AA0970CFFD9ECA92EA0FD94FF3D" ID-CoL="QZSD" authority="Saito, 1988" authorityName="Saito" authorityYear="1988" box="[151,561,151,177]" class="Arachnida" family="Linyphiidae" genus="Caracladus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="tsurusakii">
<emphasis id="41E38D31970CFFD9ECA92EA0FD94FF3D" bold="true" box="[151,561,151,177]" pageId="17" pageNumber="18">
<emphasis id="41E38D31970CFFD9ECA92EA0FE04FF3D" bold="true" box="[151,417,151,177]" italics="true" pageId="17" pageNumber="18">Caracladus tsurusakii</emphasis>
<bibRefCitation id="17062CD2970CFFD9ED962EA0FD94FF3D" author="Saito" box="[424,561,151,177]" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">Saito, 1988</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="73285123970CFFD9ECA92E8DFE88FF58" blockId="17.[151,561,151,212]" box="[151,301,186,212]" pageId="17" pageNumber="18">
(
<figureCitation id="EBAC4DA6970CFFD9ECA12E8DFE83FF58" box="[159,294,186,212]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Figs 4046</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3B8D02A8970CFFD9ECA92F3DFD15FEAD" box="[151,688,266,289]" pageId="17" pageNumber="18" type="reference_group">
<paragraph id="73285123970CFFD9ECA92F3DFD15FEAD" blockId="17.[151,688,266,289]" box="[151,688,266,289]" pageId="17" pageNumber="18">
<treatmentCitationGroup id="5387760D970CFFD9ECA92F3DFD15FEAD" box="[151,688,266,289]" pageId="17" pageNumber="18">
<taxonomicName id="B4972AA0970CFFD9ECA92F3DFD8BFEAD" ID-CoL="QZSD" authority="Saito, 1988: 21" authorityName="Saito" authorityPageNumber="21" authorityYear="1988" box="[151,558,266,289]" class="Arachnida" family="Linyphiidae" genus="Caracladus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="tsurusakii">
<emphasis id="41E38D31970CFFD9ECA92F3DFE26FEAD" box="[151,387,266,289]" italics="true" pageId="17" pageNumber="18">Caracladus tsurusakii</emphasis>
<treatmentCitation id="F2367732970CFFD9EDB42F3DFD8BFEAD" author="Saito" box="[394,558,266,289]" page="21" pageId="17" pageNumber="18" year="1988">
<bibRefCitation id="17062CD2970CFFD9EDB42F3DFDA2FEAD" author="Saito" box="[394,519,266,289]" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">Saito, 1988</bibRefCitation>
: 21
</treatmentCitation>
</taxonomicName>
, figs 1115.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="3B8D02A8970CFFD9ECA92F65FD0EFDB8" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="73285123970CFFD9ECA92F65FD0EFDB8" blockId="17.[151,1437,338,1884]" pageId="17" pageNumber="18">
<emphasis id="41E38D31970CFFD9ECA92F65FEE9FEE0" bold="true" box="[151,332,338,364]" pageId="17" pageNumber="18">
<typeStatus id="AC2CEF81970CFFD9ECA92F65FF71FEE0" box="[151,212,338,364]" pageId="17" pageNumber="18">Type</typeStatus>
material.
</emphasis>
<materialsCitation id="C3FF5B7E970CFFD9ED6D2F65FF46FE30" pageId="17" pageNumber="18" typeStatus="holotype">
<emphasis id="41E38D31970CFFD9ED6D2F65FDFAFEE0" bold="true" box="[339,607,338,364]" pageId="17" pageNumber="18">
<typeStatus id="AC2CEF81970CFFD9ED6D2F65FE5BFEE0" box="[339,510,338,364]" pageId="17" pageNumber="18" type="holotype">HOLOTYPE</typeStatus>
:
<collectingCountry id="0B8011B3970CFFD9EE342F65FDFEFEE0" box="[522,603,338,364]" name="Japan" pageId="17" pageNumber="18">Japan</collectingCountry>
:
</emphasis>
<emphasis id="41E38D31970CFFD9EE592F65FD7CFEE7" box="[615,729,338,363]" italics="true" pageId="17" pageNumber="18">
<collectingRegion id="B1539FC1970CFFD9EE592F65FD7CFEE7" box="[615,729,338,363]" country="Japan" name="Hokkaido" pageId="17" pageNumber="18">Hokkaido</collectingRegion>
</emphasis>
:
<location id="764807F8970CFFD9EED62F65FABDFEE0" LSID="urn:lsid:plazi:treatment:FB3EE035970CFFD9ECA92EA0FACEF8D0:764807F8970CFFD9EED62F65FABDFEE0" box="[744,1304,338,364]" country="Japan" latitude="42.875" longLatPrecision="19" longitude="140.65833" name="Kuttyuan-cho, Abuta-gun, Youtei-zan mountain" pageId="17" pageNumber="18" stateProvince="Hokkaido">Kuttyuan-cho, Abuta-gun, Youtei-zan mountain</location>
[
<geoCoordinate id="16A337E4970CFFD9E9182F65FF0AFE18" degrees="42" direction="north" minutes="52" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="30" value="42.875">42°52'30'' N</geoCoordinate>
,
<geoCoordinate id="16A337E4970CFFD9EC822F4DFEF9FE18" box="[188,348,378,404]" degrees="140" direction="east" minutes="39" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="30" value="140.65833">140°39'30'' E</geoCoordinate>
],
<specimenCount id="65919AAA970CFFD9ED4C2F4DFE30FE02" box="[370,405,378,404]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
<date id="072977E3970CFFD9EDA32F4DFDB9FE18" box="[413,540,378,404]" pageId="17" pageNumber="18" value="1981-09-25">
<collectingDate id="176D8E0B970CFFD9EDA32F4DFDB9FE18" box="[413,540,378,404]" pageId="17" pageNumber="18" value="1981-09-25">25.ix.1981</collectingDate>
</date>
, leg.
<collectorName id="DE6234F5970CFFD9EE602F4DFC27FE18" box="[606,898,378,404]" pageId="17" pageNumber="18">N. Tsurusaki, S. Nakano</collectorName>
,
<collectorName id="DE6234F5970CFFD9EFB12F4DFC59FE18" box="[911,1020,378,404]" pageId="17" pageNumber="18">S. Sasaki</collectorName>
(
<collectionCode id="1586C9E6970CFFD9E8332F4DFBC5FE18" box="[1037,1120,378,404]" collectionName="Japan, Tokyo, National Science Museum (Natural History)" pageId="17" pageNumber="18">NSMT</collectionCode>
<specimenCode id="2331F958970CFFD9E8572F4DFB60FE18" box="[1129,1221,378,404]" pageId="17" pageNumber="18">Ar5249</specimenCode>
, examined) (
<bibRefCitation id="17062CD2970CFFD9E9602F4DFF73FE30" author="Saito" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">Saito 1988</bibRefCitation>
).
</materialsCitation>
<materialsCitation id="C3FF5B7E970CFFD9ECCF2F95FF46FD80" ID-GBIF-Occurrence="3901401349" pageId="17" pageNumber="18" typeStatus="paratype">
<emphasis id="41E38D31970CFFD9ECCF2F95FDAFFE30" bold="true" box="[241,522,418,444]" pageId="17" pageNumber="18">
<typeStatus id="AC2CEF81970CFFD9ECCF2F95FE04FE30" box="[241,417,418,444]" pageId="17" pageNumber="18" type="paratype">PARATYPES</typeStatus>
:
<collectingCountry id="0B8011B3970CFFD9ED8A2F95FDA0FE30" box="[436,517,418,444]" name="Japan" pageId="17" pageNumber="18">Japan</collectingCountry>
:
</emphasis>
<emphasis id="41E38D31970CFFD9EE262F95FD2FFE37" box="[536,650,418,443]" italics="true" pageId="17" pageNumber="18">
<collectingRegion id="B1539FC1970CFFD9EE262F95FD2FFE37" box="[536,650,418,443]" country="Japan" name="Hokkaido" pageId="17" pageNumber="18">Hokkaido</collectingRegion>
</emphasis>
: Kuttyuan-cho, Abuta-gun, Youtei-zan mountain [
<geoCoordinate id="16A337E4970CFFD9E8C52F95FA32FE30" box="[1275,1431,418,444]" degrees="42" direction="north" minutes="52" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="30" value="42.875">42°52'30'' N</geoCoordinate>
,
<geoCoordinate id="16A337E4970CFFD9ECA92FFDFE9FFE68" box="[151,314,458,484]" degrees="140" direction="east" minutes="39" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="30" value="140.65833">140°39'30'' E</geoCoordinate>
],
<specimenCount id="65919AAA970CFFD9ED6C2FFDFED0FE52" box="[338,373,458,484]" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
<date id="072977E3970CFFD9EDBE2FFDFE5AFE68" box="[384,511,458,484]" pageId="17" pageNumber="18" value="1981-09-25">25.ix.1981</date>
, leg. N. Tsurusaki, S. Nakano, S. Sasaki (
<collectionCode id="1586C9E6970CFFD9E83A2FFDFBF2FE68" box="[1028,1111,458,484]" collectionName="Japan, Tokyo, National Science Museum (Natural History)" pageId="17" pageNumber="18">NSMT</collectionCode>
Ar5250, examined) (
<bibRefCitation id="17062CD2970CFFD9E9602FFDFF73FD80" author="Saito" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">Saito 1988</bibRefCitation>
);
</materialsCitation>
<materialsCitation id="C3FF5B7E970CFFD9ECD32FC5FD0DFDB8" ID-GBIF-Occurrence="3901401336" pageId="17" pageNumber="18">
Kuttyuan-cho, Abuta-gun, Youtei-zan mountain [
<geoCoordinate id="16A337E4970CFFD9EF0A2FC5FC69FD80" box="[820,972,498,524]" degrees="42" direction="north" minutes="52" orientation="latitude" pageId="17" pageNumber="18" precision="15" seconds="30" value="42.875">42°52'30'' N</geoCoordinate>
,
<geoCoordinate id="16A337E4970CFFD9EFE42FC5FBDEFD80" box="[986,1147,498,524]" degrees="140" direction="east" minutes="39" orientation="longitude" pageId="17" pageNumber="18" precision="15" seconds="30" value="140.65833">140°39'30'' E</geoCoordinate>
],
<specimenCount id="65919AAA970CFFD9E8AF2FC5FB11FD8A" box="[1169,1204,498,524]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
<date id="072977E3970CFFD9E8832FC5FA98FD80" box="[1213,1341,498,524]" pageId="17" pageNumber="18" value="1981-09-25">25.ix.1981</date>
, leg. N. Tsurusaki, S. Nakano, S. Sasaki (
<bibRefCitation id="17062CD2970CFFD9EE242C2DFD3BFDB8" author="Saito" box="[538,670,538,564]" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">Saito 1988</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B8D02A8970CFFD9ECFB2C75FC69FCA8" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="73285123970CFFD9ECFB2C75FAFFFD58" blockId="17.[151,1437,338,1884]" pageId="17" pageNumber="18">
<emphasis id="41E38D31970CFFD9ECFB2C75FE3CFDD0" bold="true" box="[197,409,578,604]" pageId="17" pageNumber="18">
Diagnosis.
<emphasis id="41E38D31970CFFD9ED6E2C75FE3CFDD0" bold="true" box="[336,409,578,604]" italics="true" pageId="17" pageNumber="18">Males</emphasis>
</emphasis>
: Most similar to
<taxonomicName id="B4972AA0970CFFD9EE4F2C75FD49FDD7" box="[625,748,578,603]" class="Arachnida" family="Linyphiidae" genus="Caracladus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="avicula">
<emphasis id="41E38D31970CFFD9EE4F2C75FD49FDD7" box="[625,748,578,603]" italics="true" pageId="17" pageNumber="18">C. avicula</emphasis>
</taxonomicName>
with differences in the structure of the male palp (
<bibRefCitation id="17062CD2970CFFD9E9602C75FF73FD08" author="Saito" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">Saito 1988</bibRefCitation>
). Cephalic lobe similar to
<taxonomicName id="B4972AA0970CFFD9EE2B2C5DFD2AFD0F" box="[533,655,618,643]" class="Arachnida" family="Linyphiidae" genus="Caracladus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="avicula">
<emphasis id="41E38D31970CFFD9EE2B2C5DFD2AFD0F" box="[533,655,618,643]" italics="true" pageId="17" pageNumber="18">C. avicula</emphasis>
</taxonomicName>
but more robust and lacking a neck. Embolus whip-like as in
<taxonomicName id="B4972AA0970CFFD9E9BF2C5DFF48FD27" class="Arachnida" family="Linyphiidae" genus="Caracladus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="avicula">
<emphasis id="41E38D31970CFFD9E9BF2C5DFF48FD27" italics="true" pageId="17" pageNumber="18">C. avicula</emphasis>
</taxonomicName>
but much longer and with a loop back, radical tailpiece long and bent (
<figureCitation id="EBAC4DA6970CFFD9E8752CA5FB00FD20" box="[1099,1189,658,684]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Fig. 41</figureCitation>
). Distal suprategular apophysis blunt and nearly transparent (
<figureCitation id="EBAC4DA6970CFFD9EE552C8DFD66FD58" box="[619,707,698,724]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Fig. 41</figureCitation>
). Palpal tibia with two well defined apophyses (
<figureCitation id="EBAC4DA6970CFFD9E8CB2C8DFAEBFD58" box="[1269,1358,698,724]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Fig. 42</figureCitation>
).
</paragraph>
<paragraph id="73285123970CFFD9ECFB2CD5FC69FCA8" blockId="17.[151,1437,338,1884]" pageId="17" pageNumber="18">
<emphasis id="41E38D31970CFFD9ECFB2CD5FE8EFD70" bold="true" box="[197,299,738,764]" italics="true" pageId="17" pageNumber="18">Females</emphasis>
: Nearly triangular dorsal plate with two copulatory pouches anterior and ventral to it and together not broader than the anterior end of the dorsal plate (
<figureCitation id="EBAC4DA6970CFFD9EF592D3DFC65FCA8" box="[871,960,778,804]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Fig. 40</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="3B8D02A8970CFFD9ECFB2D05FD65F9E0" pageId="17" pageNumber="18" type="description">
<paragraph id="73285123970CFFD9ECFB2D05FE1FFAF0" blockId="17.[151,1437,338,1884]" pageId="17" pageNumber="18">
<emphasis id="41E38D31970CFFD9ECFB2D05FE03FCC0" bold="true" box="[197,422,818,844]" pageId="17" pageNumber="18">
Description.
<emphasis id="41E38D31970CFFD9ED562D05FE03FCC0" bold="true" box="[360,422,818,844]" italics="true" pageId="17" pageNumber="18">Male</emphasis>
</emphasis>
(
<typeStatus id="AC2CEF81970CFFD9ED872D05FD81FCC0" box="[441,548,818,844]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
, NSMT Ar5249): Total length:
<quantity id="B46FFCC6970CFFD9EF962D05FBBDFCC0" box="[936,1048,818,844]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="18" unit="mm" value="2.5">2.50 mm</quantity>
. Cephalothorax: chestnut brown with darker radiating lines, cervical grooves black (
<bibRefCitation id="17062CD2970CFFD9EED22D6DFCD5FCF8" author="Saito" box="[748,880,858,884]" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">Saito 1988</bibRefCitation>
);
<quantity id="B46FFCC6970CFFD9EFBA2D6DFC49FCF8" box="[900,1004,858,884]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="17" pageNumber="18" unit="mm" value="1.03">1.03 mm</quantity>
long without cephalic lobe (
<figureCitation id="EBAC4DA6970CFFD9E9092D6DFA2BFCF8" box="[1335,1422,858,884]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Fig. 45</figureCitation>
),
<quantity id="B46FFCC6970CFFD9ECA92DB5FEA4FC10" box="[151,257,898,924]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="17" pageNumber="18" unit="mm" value="1.35">1.35 mm</quantity>
long with cephalic lobe (
<figureCitation id="EBAC4DA6970CFFD9EE0B2DB5FD2BFC10" box="[565,654,898,924]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Fig. 45</figureCitation>
);
<quantity id="B46FFCC6970CFFD9EE982DB5FCB5FC10" box="[678,784,898,924]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="17" pageNumber="18" unit="mm" value="0.78">0.78 mm</quantity>
wide. Cephalic lobe: projecting forward, thick at base and sharply pointed tip with sparse stout hairs; at thinnest part below the eye-field
<quantity id="B46FFCC6970CFFD9E8BE2D9DFB4EFC48" box="[1152,1259,938,964]" metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="18" unit="mm" value="0.14">0.14 mm</quantity>
wide laterally,
<quantity id="B46FFCC6970CFFD9ECA92DE5FEA5FC60" box="[151,256,978,1004]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="17" pageNumber="18" unit="mm" value="0.23">0.23 mm</quantity>
wide dorsally (
<figureCitation id="EBAC4DA6970CFFD9ED862DE5FDE7FC60" box="[440,578,978,1004]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Figs 43, 45</figureCitation>
); sulcus absent (
<figureCitation id="EBAC4DA6970CFFD9EF382DE5FCC5FC60" box="[774,864,978,1004]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Fig. 44</figureCitation>
). Eyes: PME upmost on the cephalic lobe, separated by slightly less than 1.5 times their diameter; position of the PLE adjacent to and above the ALE; one long macroseta projecting forward between AME (
<figureCitation id="EBAC4DA6970CFFD9EED72A15FCE5FBB0" box="[745,832,1058,1084]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Fig. 45</figureCitation>
); anterior row slightly procurved, eyes separated by their radius. Clypeus: wide, slightly convex. Chelicerae: dark brown (
<bibRefCitation id="17062CD2970CFFD9EFF32A7DFBF7FBE8" author="Saito" box="[973,1106,1098,1124]" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">Saito 1988</bibRefCitation>
); promargin with five teeth; retromargin with six denticles; many broad stridulatory striae. Sternum: yellowish brown, strongly darker at the margins (
<bibRefCitation id="17062CD2970CFFD9ED0A2AADFE1FFB38" author="Saito" box="[308,442,1178,1204]" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">Saito 1988</bibRefCitation>
). Legs: tibia IIV with one dorsal proximal macroseta (1-1-1-1); Tm I: 0.63. Pedipalp: prolateral tibial apophysis glabrous, facing forward; retrolateral tibial apophysis hairy, facing forward (
<bibRefCitation id="17062CD2970CFFD9ECA12ADDFE29FA88" author="Saito" box="[159,396,1258,1284]" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">
Saito 1988:
<figureCitation id="EBAC4DA6970CFFD9ED0A2ADDFE29FA88" box="[308,396,1258,1284]" captionStart="FIGURES 9 16" captionStartId="5.[151,269,1938,1962]" captionTargetBox="[182,1400,184,1912]" captionTargetId="figure@5.[182,1407,184,1915]" captionTargetPageId="5" captionText="FIGURES 9 16. Caracladus avicula (L. Koch, 1869). Male: cephalothorax lateral (9), frontal (10), cephalic lobe lateral (11); thick hair on cephalic lobe anterior to the PME (12); chelicerae lateral (13); sulci frontal (14); booklung cover ventral (15); left spinnerets (16). Scale: 100 Μm if not stated differently." httpUri="https://zenodo.org/record/185323/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
</bibRefCitation>
); suprategular apophysis semi circular; marginal suprategular apophysis minute; distal suprategular apophysis thin, blunt (
<figureCitation id="EBAC4DA6970CFFD9EE062B25FD34FAA0" box="[568,657,1298,1324]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Fig. 41</figureCitation>
); embolic membrane with papillae; radix simple without any processes other than the long radical tailpiece and the very long, whip-like embolus that is strongly curved towards the tip (
<figureCitation id="EBAC4DA6970CFFD9ED6B2B55FE08FAF0" box="[341,429,1378,1404]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Fig. 41</figureCitation>
).
</paragraph>
<paragraph id="73285123970CFFD9ECFB2BBDFD65F9E0" blockId="17.[151,1437,338,1884]" pageId="17" pageNumber="18">
<emphasis id="41E38D31970CFFD9ECFB2BBDFE85FA28" bold="true" box="[197,288,1418,1444]" italics="true" pageId="17" pageNumber="18">Female</emphasis>
(
<typeStatus id="AC2CEF81970CFFD9ED112BBDFE3DFA28" box="[303,408,1418,1444]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
, NSMT Ar5250): Total length:
<quantity id="B46FFCC6970CFFD9EF372BBDFCD0FA28" box="[777,885,1418,1444]" metricMagnitude="-3" metricUnit="m" metricValue="2.58" pageId="17" pageNumber="18" unit="mm" value="2.58">2.58 mm</quantity>
. Cephalothorax:
<quantity id="B46FFCC6970CFFD9E8052BBDFB07FA28" box="[1083,1186,1418,1444]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="17" pageNumber="18" unit="mm" value="1.08">1.08 mm</quantity>
long;
<quantity id="B46FFCC6970CFFD9E8D22BBDFAF1FA28" box="[1260,1364,1418,1444]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="17" pageNumber="18" unit="mm" value="0.83">0.83 mm</quantity>
wide; general appearance similar to males, but without cephalic lobe (
<bibRefCitation id="17062CD2970CFFD9EF8F2B85FB9EFA40" author="Saito" box="[945,1083,1458,1484]" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">Saito 1988</bibRefCitation>
). Eyes: anterior row slightly recurved, separated by about the radius of the AME; posterior eyes separated by at least an eye diameter (
<bibRefCitation id="17062CD2970CFFD9ECA12835FE81F990" author="Saito" box="[159,292,1538,1564]" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">Saito 1988</bibRefCitation>
). Epigyne: dorsal plate visible in ventral view; receptacula lateral to the dorsal plate, barely visible in transparency trough ventral plate (
<figureCitation id="EBAC4DA6970CFFD9EEBD281DFD7AF9C8" box="[643,735,1578,1604]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Fig. 40</figureCitation>
); very small copulatory pouches (
<figureCitation id="EBAC4DA6970CFFD9E846281DFB76F9C8" box="[1144,1235,1578,1604]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Fig. 40</figureCitation>
). Vulva: without copulatory duct (
<figureCitation id="EBAC4DA6970CFFD9ED602865FE13F9E0" box="[350,438,1618,1644]" captionStart="FIGURES 40 46" captionStartId="18.[151,269,1931,1955]" captionTargetBox="[212,1365,209,1895]" captionTargetId="figure@18.[180,1405,197,1903]" captionTargetPageId="18" captionText="FIGURES 40 46. Caracladus tsurusakii Saito, 1988. Female: epigyne ventral (40). Male: palp prolateral (41); palpal tibia dorsal (42); cephalothorax dorsal (43), frontal (44) and lateral (45); right leg tibia I retrolateral (46). Scale: 200 Μm. Specimens: Paratype, NSMT Ar 5250 (female); holotype, NSMT Ar 5250 (male)." httpUri="https://zenodo.org/record/185328/files/figure.png" pageId="17" pageNumber="18">Fig. 40</figureCitation>
); receptacula globular.
</paragraph>
</subSubSection>
<subSubSection id="3B8D02A8970CFFD9ECFB284DFB3DF968" pageId="17" pageNumber="18" type="distribution">
<paragraph id="73285123970CFFD9ECFB284DFB31F918" blockId="17.[151,1437,338,1884]" box="[197,1172,1658,1684]" pageId="17" pageNumber="18">
<emphasis id="41E38D31970CFFD9ECFB284DFEC3F918" bold="true" box="[197,358,1658,1684]" pageId="17" pageNumber="18">Distribution.</emphasis>
Only known from the
<typeStatus id="AC2CEF81970CFFD9EE4F284DFD06F918" box="[625,675,1658,1684]" pageId="17" pageNumber="18">type</typeStatus>
locality on
<collectingRegion id="B1539FC1970CFFD9EF11284DFC03F918" box="[815,934,1658,1684]" country="Japan" name="Hokkaido" pageId="17" pageNumber="18">Hokkaido</collectingRegion>
,
<collectingCountry id="0B8011B3970CFFD9EF8F284DFC51F918" box="[945,1012,1658,1684]" name="Japan" pageId="17" pageNumber="18">Japan</collectingCountry>
(
<bibRefCitation id="17062CD2970CFFD9E83D284DFB22F918" author="Saito" box="[1027,1159,1658,1684]" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">Saito 1988</bibRefCitation>
).
</paragraph>
<paragraph id="73285123970CFFD9ECFB2895FB3DF968" blockId="17.[151,1437,338,1884]" pageId="17" pageNumber="18">
<emphasis id="41E38D31970CFFD9ECFB2895FE88F930" bold="true" box="[197,301,1698,1724]" pageId="17" pageNumber="18">Habitat.</emphasis>
The specimens were collected under leaf litter by sifting or with pitfall traps (
<bibRefCitation id="17062CD2970CFFD9E8FB2895FAEFF930" author="Saito" box="[1221,1354,1698,1724]" pageId="17" pageNumber="36" refString="Saito, H. (1988) Four new erigonine spiders (Araneae: Linyphiidae) from Japan. Edaphologia, 39, 17 - 24." type="journal article" year="1988">Saito 1988</bibRefCitation>
). If we correctly located the locus typicus, this species occurs on a volcano on ca.
<quantity id="B46FFCC6970CFFD9E83F28FDFBFFF968" box="[1025,1114,1738,1764]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="17" pageNumber="18" unit="m" value="1300.0">1300 m</quantity>
a.s.l.
</paragraph>
</subSubSection>
<subSubSection id="3B8D02A8970CFFD9ECFB28C5FCB3F880" box="[197,790,1778,1804]" pageId="17" pageNumber="18" type="description">
<paragraph id="73285123970CFFD9ECFB28C5FCB3F880" blockId="17.[151,1437,338,1884]" box="[197,790,1778,1804]" pageId="17" pageNumber="18">
<emphasis id="41E38D31970CFFD9ECFB28C5FEE8F880" bold="true" box="[197,333,1778,1804]" pageId="17" pageNumber="18">Phenology.</emphasis>
One record from late September, only.
</paragraph>
</subSubSection>
<subSubSection id="3B8D02A8970CFFD9ECFB292DFACEF8D0" pageId="17" pageNumber="18" type="discussion">
<paragraph id="73285123970CFFD9ECFB292DFACEF8D0" blockId="17.[151,1437,338,1884]" pageId="17" pageNumber="18">
<emphasis id="41E38D31970CFFD9ECFB292DFE9BF8B8" bold="true" box="[197,318,1818,1844]" pageId="17" pageNumber="18">Remarks.</emphasis>
<taxonomicName id="B4972AA0970CFFD9ED78292CFE79F8B8" box="[326,476,1819,1844]" class="Arachnida" family="Linyphiidae" genus="Caracladus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="tsurusakii">
<emphasis id="41E38D31970CFFD9ED78292CFE79F8B8" box="[326,476,1819,1844]" italics="true" pageId="17" pageNumber="18">C. tsurusakii</emphasis>
</taxonomicName>
lacks a copulatory duct. The insertion of sperm is assumed to take place through a space between the ventral and the dorsal plates which are supposed to be pressed apart during copulation.
</paragraph>
</subSubSection>
</treatment>
</document>