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<document id="0B37E3AF5A11945B7BB5CA46E1DEF304" ID-CLB-Dataset="21971" ID-DOI="10.5852/ejt.2016.189" ID-GBIF-Dataset="2e12b6f4-3106-47aa-8332-5ef9efa64219" ID-ISSN="2118-9773" ID-Zenodo-Dep="3837729" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590067049572" checkinUser="valdenar" docAuthor="Liu, Lan-Yu, Beaver, Roger A. &amp; Sanguansub, Sunisa" docDate="2016" docId="FB0E87A7FFE9083B10360C2E3A1CFDB5" docLanguage="en" docName="ejt-2016-189.pdf.imf" docOrigin="European Journal of Taxonomy 189 (189)" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Octomeristes Liu &amp; Beaver 2016, gen. nov." docType="treatment" docVersion="9" lastPageNumber="6" masterDocId="0737FFDFFFEB083E12570E233814FFD3" masterDocTitle="A new Oriental genus of bostrichid beetle (Coleoptera: Bostrichidae: Xyloperthini), a new synonym and a lectotype designation for Octodesmus episternalis (Lesne, 1901)" masterLastPageNumber="12" masterPageNumber="1" pageNumber="3" updateTime="1698840606035" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="BE689D1E16E2B6D2C42A0FDC626F05D5">A new Oriental genus of bostrichid beetle (Coleoptera: Bostrichidae: Xyloperthini), a new synonym and a lectotype designation for Octodesmus episternalis (Lesne, 1901)</mods:title>
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<mods:namePart id="28658E38587066356054CC2677F842DA">Liu, Lan-Yu</mods:namePart>
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<mods:affiliation id="030AB594116C67141C36659842EE82B9">Department of Science Communication, National Pingtung University, No. 4 - 18, Minsheng Rd, Pingtung City, Pingtung County 90049, Taiwan. &amp; urn: lsid: zoobank. org: author: 8 A 4 ECE 7 C- 2607 - 440 D-B 1 BC- 6 E 3 B 05 EF 02 BB &amp; Corresponding author: liulysky @ gmail. com</mods:affiliation>
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<mods:namePart id="F883DB3B09E36CA90C14E96EE70DFB08">Beaver, Roger A.</mods:namePart>
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<mods:affiliation id="693B9E639C29CE05B41EA7BD84ADD264">161 / 2 Mu 5, Soi Wat Pranon, T. Donkaew, A. Maerim, Chiangmai 50180, Thailand. &amp; rogerbeaver 6 @ gmail. com &amp; urn: lsid: zoobank. org: author: EEF 5 C 471 - ECFB- 4786 - 8 E 2 F- 13 C 5 B 5 EC 4 F 0 D</mods:affiliation>
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<mods:namePart id="A60C7EA2C057B6D4AAF4D8D8D932CB5D">Sanguansub, Sunisa</mods:namePart>
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<mods:affiliation id="424DB64E0394F68FB93D8D7DFF02FAAA">Department of Entomology, Faculty of Agriculture at Kamphaeng Saen, Kasetsart University, Kamphaeng Saen, Nakhonpathom 73140, Thailand. &amp; agrssss @ ku. ac. th &amp; urn: lsid: zoobank. org: author: 65 EBEA 67 - 515 A- 4 CC 7 - AE 93 - DAD 47765971 E</mods:affiliation>
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<mods:title id="26A256BB8D92C53BA2B75EB4D78FC23A">European Journal of Taxonomy</mods:title>
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<mods:date id="D59BDE160A57F753555568447C181324">2016</mods:date>
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<treatment id="FB0E87A7FFE9083B10360C2E3A1CFDB5" ID-DOI="http://doi.org/10.5281/zenodo.3852244" ID-GBIF-Taxon="164245969" ID-Zenodo-Dep="3852244" LSID="urn:lsid:zoobank.org:act:4D659D1D-561C-445C-A256-AFA66495D93C" httpUri="http://treatment.plazi.org/id/FB0E87A7FFE9083B10360C2E3A1CFDB5" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
<subSubSection id="3BBD653AFFE9083C10360C2E3CFAFD98" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="731836B1FFE9083C10360C2E3C0AFDFB" blockId="2.[401,1262,525,587]" box="[609,1054,525,552]" pageId="2" pageNumber="3">
<heading id="285081DDFFE9083C10360C2E3C0AFDFB" box="[609,1054,525,552]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
<taxonomicName id="B4A74D32FFE9083C10360C2E3BBCFDFB" authority="Liu &amp; Beaver" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[609,936,525,552]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="41D3EAA3FFE9083C10360C2E3AE9FDF4" bold="true" box="[609,765,525,551]" italics="true" pageId="2" pageNumber="3">Octomeristes</emphasis>
Liu &amp; Beaver
</taxonomicName>
,
<taxonomicNameLabel id="5AE057D8FFE9083C11E30C2D3C0AFDFB" box="[948,1054,526,552]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
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urn:lsid:zoobank.org:act:
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<paragraph id="731836B1FFE9083C12B40C4C396AFD5B" blockId="2.[227,382,622,649]" box="[227,382,622,649]" pageId="2" pageNumber="3">
<heading id="285081DDFFE9083C12B40C4C396AFD5B" bold="true" box="[227,382,622,649]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="41D3EAA3FFE9083C12B40C4C396AFD5B" bold="true" box="[227,382,622,649]" pageId="2" pageNumber="3">
<typeStatus id="AC1C8813FFE9083C12B40C4C3934FD5A" box="[227,288,623,649]" pageId="2" pageNumber="3">Type</typeStatus>
species
</emphasis>
</heading>
</paragraph>
<paragraph id="731836B1FFE9083C12B40CBD3C00FD64" blockId="2.[227,1044,669,696]" box="[227,1044,669,696]" pageId="2" pageNumber="3">
<taxonomicName id="B4A74D32FFE9083C12B40CBD3A9FFD6B" authority="Liu &amp; Beaver" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[227,651,669,696]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="pusillus" status="gen. et sp. nov.">
<emphasis id="41D3EAA3FFE9083C12B40CBD39F4FD64" box="[227,480,669,696]" italics="true" pageId="2" pageNumber="3">Octomeristes pusillus</emphasis>
Liu &amp; Beaver
</taxonomicName>
,
<taxonomicNameLabel id="5AE057D8FFE9083C10C10CBD3B5DFD6B" box="[662,841,670,696]" pageId="2" pageNumber="3" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, here designated.
</paragraph>
</subSubSection>
<subSubSection id="3BBD653AFFE9083C12B40CF83B99FB57" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="731836B1FFE9083C12B40CF83948FD26" blockId="2.[227,348,731,757]" box="[227,348,731,757]" pageId="2" pageNumber="3">
<heading id="285081DDFFE9083C12B40CF83948FD26" bold="true" box="[227,348,731,757]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="41D3EAA3FFE9083C12B40CF83948FD26" bold="true" box="[227,348,731,757]" pageId="2" pageNumber="3">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="731836B1FFE9083C12B40D293B99FB57" blockId="2.[227,1437,778,1156]" pageId="2" pageNumber="3">
A member of the tribe
<taxonomicName id="B4A74D32FFE9083C13A80D293A8CFCF7" authorityName="Lesne" authorityYear="1921" box="[511,664,778,804]" class="Insecta" family="Bostrichidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Xyloperthini">Xyloperthini</taxonomicName>
, characterized by the lamelliform intercoxal process of the first abdominal ventrite, and the mandibles crossed at the tips (
<bibRefCitation id="17364B40FFE9083C11C10D0D3C30FC9B" author="Lesne P." box="[918,1060,814,840]" pageId="2" pageNumber="3" pagination="473 - 639" refId="ref6385" refString="Lesne P. 1901. Revision des Coleopteres de la famille des Bostrychides 4. Annales de la Societe Entomologique de France 69: 473 - 639." type="journal article" year="1901">Lesne 1901</bibRefCitation>
;
<bibRefCitation id="17364B40FFE9083C16660D0E3CD5FC9B" author="Fisher W. S." box="[1073,1217,813,840]" pageId="2" pageNumber="3" refId="ref6331" refString="Fisher W. S. 1950. A revision of the North American species of beetles belonging to the family Bostrichidae. Miscellaneous Publication 698. United States Department of Agriculture, Washington, D. C. http: // dx. doi. org / 10.5962 / bhl. title. 65663" type="book" year="1950">Fisher 1950</bibRefCitation>
;
<bibRefCitation id="17364B40FFE9083C16990D0E3935FCB8" author="Liu L. Y. &amp; Schonitzer K." pageId="2" pageNumber="3" pagination="99 - 132" refId="ref6525" refString="Liu L. Y. &amp; Schonitzer K. 2011. Phylogenetic analysis of the family Bostrichidae auct. at suprageneric levels (Coleoptera: Bostrichidae). Mitteilungen der Munchner Entomologischen Gesellschaft 101: 99 - 132." type="journal article" year="2011">Liu &amp; Schönitzer 2011</bibRefCitation>
). Distinguished from other genera of
<taxonomicName id="B4A74D32FFE9083C10BC0D723B95FCB8" authorityName="Lesne" authorityYear="1921" box="[747,897,849,875]" class="Insecta" family="Bostrichidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Xyloperthini">Xyloperthini</taxonomicName>
by the following combination of characters: Frons weakly convex, without a crown of hairs on the head in either sex. Mandibles symmetrical, sharply pointed; antenna with eight antennomeres, the funicle and club each trimerous, antennomeres of club lacking stiff, erect hairs, with two indistinct c-shaped sensory impressions near the apex of antennomeres 6 and 7, but not on the last antennomere; pronotum without a lateral carina; posterior part of elytral disc with more than one pair of costae, each ending in a small process at the upper margin of the elytral declivity; protibiae normal, without small teeth on posterior face; female with third abdominal ventrite projecting over and concealing fourth ventrite, its posterior margin with lobes or teeth; fifth abdominal ventrite of male with pleural pieces.
</paragraph>
</subSubSection>
<subSubSection id="3BBD653AFFE9083C12B40A8A3C28FB21" box="[227,1084,1193,1266]" pageId="2" pageNumber="3" type="etymology">
<paragraph id="731836B1FFE9083C12B40A8A397EFB10" blockId="2.[227,362,1193,1219]" box="[227,362,1193,1219]" pageId="2" pageNumber="3">
<heading id="285081DDFFE9083C12B40A8A397EFB10" bold="true" box="[227,362,1193,1219]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="41D3EAA3FFE9083C12B40A8A397EFB10" bold="true" box="[227,362,1193,1219]" pageId="2" pageNumber="3">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="731836B1FFE9083C12B40AF43C28FB21" blockId="2.[227,1084,1239,1266]" box="[227,1084,1239,1266]" pageId="2" pageNumber="3">The genus name is masculine, and refers to the eight-segmented antenna.</paragraph>
</subSubSection>
<subSubSection id="3BBD653AFFE9083A12B40B363B18FD46" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="731836B1FFE9083C12B40B363960FAFC" blockId="2.[227,372,1301,1327]" box="[227,372,1301,1327]" pageId="2" pageNumber="3">
<heading id="285081DDFFE9083C12B40B363960FAFC" bold="true" box="[227,372,1301,1327]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="41D3EAA3FFE9083C12B40B363960FAFC" bold="true" box="[227,372,1301,1327]" pageId="2" pageNumber="3">Description</emphasis>
</heading>
</paragraph>
<paragraph id="731836B1FFE9083C12B40B6739FCF8D7" blockId="2.[227,1437,1348,1796]" pageId="2" pageNumber="3">Body elongate, cylindrical. Head deeply inserted in prothorax, not visible from above. Frons convex or slightly impressed, with scattered, fine, upwardly-directed hairs, which are denser and coarser laterally; fronto-clypeal suture indistinct at sides, impressed in middle; clypeus strongly transverse, anterior margin with a very short median lobe or projection, an emargination on each side of the lobe; labrum transverse with a fringe of hairs along anterior margin. Mandibles subequal, sharply pointed. Eyes small, oval, globose, strongly projecting. Antenna with 8 antennomeres, first antennomere elongate, about twice as long as the oval second, antennomeres 35 forming a loose funicle, each antennomere short, strongly transverse, the fifth widest, together slightly shorter than second antennomere; antennomeres 68 forming the elongate, compressed club, each antennomere with a dense covering of short, recumbent hairs, two indistinct c-shaped sensory impressions on the surface of antennomeres 6 and 7 near apex, but absent from antennomere 8; antennomeres 6 and 7 subquadrate to subtriangular, subequal in length, last antennomere elongate, oval, length variable from 1.51.9 times longer than the previous antennomere, but subequal in width.</paragraph>
<paragraph id="731836B1FFE9083C12B4090939ECF7C5" blockId="2.[227,1437,1833,2036]" lastBlockId="2.[291,504,2044,2070]" pageId="2" pageNumber="3">
Pronotum slightly wider than long, widest about one-third from base, anterior angles with a strong, upcurved tooth on margin, anterior margin between teeth straight, not depressed behind the margin; semicircular area above the anterior margin smooth and without teeth; sides broadly rounded, converging more strongly anteriorly, posterior angles broadly rounded, without a lateral carina, postero-lateral area with fine rugulosities; disc smooth, shining, very indistinctly punctured, glabrous; anterior slope with 3 or 4 large, upcurved teeth antero-laterally on each side behind the marginal tooth, with short, semi-erect hairs
<pageTitle id="3338EED6FFE9083C137409DF39ECF7C5" box="[291,504,2044,2070]" pageId="2" pageNumber="3">between the teeth.</pageTitle>
</paragraph>
<caption id="27D86639FFE8083D128C0C663D8DFDB2" ID-Table-UUID="27D86639FFE8083D128C0C663D8DFDB2" box="[219,1433,581,609]" httpUri="http://table.plazi.org/id/27D86639FFE8083D128C0C663D8DFDB2" pageId="3" pageNumber="4" targetBox="[225,1969,654,1781]" targetIsTable="true" targetPageId="3">
<paragraph id="731836B1FFE8083D128C0C663D8DFDB2" blockId="3.[219,1433,581,609]" box="[219,1433,581,609]" pageId="3" pageNumber="4">
<emphasis id="41D3EAA3FFE8083D128C0C66392AFDB2" bold="true" box="[219,318,581,609]" pageId="3" pageNumber="4">Table 1.</emphasis>
Comparison of characters of
<taxonomicName id="B4A74D32FFE8083D10CD0C663BC9FDB2" authority="Liu &amp; Beaver" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[666,989,581,609]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFE8083D10CD0C663B21FDB2" box="[666,821,581,609]" italics="true" pageId="3" pageNumber="4">Octomeristes</emphasis>
Liu &amp; Beaver
</taxonomicName>
with selected genera of
<taxonomicName id="B4A74D32FFE8083D16AC0C663D81FDB2" authorityName="Lesne" authorityYear="1921" box="[1275,1429,581,609]" class="Insecta" family="Bostrichidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Xyloperthini">Xyloperthini</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="731836B1FFE8083D13E10CAD3F4BF926" pageId="3" pageNumber="4">
<table id="01A7C411FFE8F7C112B60CAD3FA5F926" box="[225,1969,654,1781]" gridcols="6" gridrows="19" pageId="3" pageNumber="4">
<tr id="CD9734F3FFE8F7C112B60CAD3FA5FD76" box="[225,1969,654,677]" gridrow="0" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="8E465D8FFFE8F7C112B60CAD39ECFD76" box="[225,504,654,677]" gridcol="0" gridrow="0" pageId="3" pageNumber="4">
<emphasis id="41D3EAA3FFE8083D13E10CAD39E3FD76" bold="true" box="[438,503,654,677]" pageId="3" pageNumber="4">Genus</emphasis>
</th>
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<taxonomicName id="B4A74D32FFE8083D107F0C883ABCFD11" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[552,680,683,706]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="41D3EAA3FFE8083D107F0C883ABCFD11" bold="true" box="[552,680,683,706]" italics="true" pageId="3" pageNumber="4">Octomeristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AE057D8FFE8083D10F90C883B13FD11" box="[686,775,683,706]" pageId="3" pageNumber="4" rank="genus">gen. nov.</taxonomicNameLabel>
</td>
<td id="8E465D8FFFE8F7C1117A0C883C5CFD11" box="[813,1096,683,706]" gridcol="2" gridrow="1" pageId="3" pageNumber="4">
<taxonomicName id="B4A74D32FFE8083D11170C883C22FD11" authority="Lesne, 1901" authorityName=", Lesne" authorityYear="1901" box="[832,1078,683,706]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFE8083D11170C883BACFD11" bold="true" box="[832,952,683,706]" italics="true" pageId="3" pageNumber="4">Octodesmus</emphasis>
<bibRefCitation id="17364B40FFE8083D11E90C883C22FD11" author="Lesne P." box="[958,1078,683,706]" pageId="3" pageNumber="4" pagination="473 - 639" refId="ref6385" refString="Lesne P. 1901. Revision des Coleopteres de la famille des Bostrychides 4. Annales de la Societe Entomologique de France 69: 473 - 639." type="journal article" year="1901">Lesne, 1901</bibRefCitation>
</taxonomicName>
</td>
<td id="8E465D8FFFE8F7C116060C883D7FFD11" box="[1105,1387,683,706]" gridcol="3" gridrow="1" pageId="3" pageNumber="4">
<taxonomicName id="B4A74D32FFE8083D163E0C883D47FD11" authority="Lesne, 1901" authorityName="Lesne" authorityYear="1901" box="[1129,1363,683,706]" class="Insecta" family="Bostrichidae" genus="Xylionulus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFE8083D163E0C883CC1FD11" bold="true" box="[1129,1237,683,706]" italics="true" pageId="3" pageNumber="4">Xylionulus</emphasis>
<bibRefCitation id="17364B40FFE8083D168C0C883D47FD11" author="Lesne P." box="[1243,1363,683,706]" pageId="3" pageNumber="4" pagination="473 - 639" refId="ref6385" refString="Lesne P. 1901. Revision des Coleopteres de la famille des Bostrychides 4. Annales de la Societe Entomologique de France 69: 473 - 639." type="journal article" year="1901">Lesne, 1901</bibRefCitation>
</taxonomicName>
</td>
<td id="8E465D8FFFE8F7C117240C883E99FD11" box="[1395,1677,683,706]" gridcol="4" gridrow="1" pageId="3" pageNumber="4">
<taxonomicName id="B4A74D32FFE8083D17D80C883E66FD11" authority="Lesne, 1901" authorityName="Lesne" authorityYear="1901" box="[1423,1650,683,706]" class="Insecta" family="Bostrichidae" genus="Xylobosca" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFE8083D17D80C883DE0FD11" bold="true" box="[1423,1524,683,706]" italics="true" pageId="3" pageNumber="4">Xylobosca</emphasis>
<bibRefCitation id="17364B40FFE8083D17AD0C883E66FD11" author="Lesne P." box="[1530,1650,683,706]" pageId="3" pageNumber="4" pagination="473 - 639" refId="ref6385" refString="Lesne P. 1901. Revision des Coleopteres de la famille des Bostrychides 4. Annales de la Societe Entomologique de France 69: 473 - 639." type="journal article" year="1901">Lesne, 1901</bibRefCitation>
</taxonomicName>
</td>
<td id="8E465D8FFFE8F7C114C00C883FA5FD11" box="[1687,1969,683,706]" gridcol="5" gridrow="1" pageId="3" pageNumber="4">
<emphasis id="41D3EAA3FFE8083D14930C883F11FD11" bold="true" box="[1732,1797,683,706]" italics="true" pageId="3" pageNumber="4">Xylion</emphasis>
<bibRefCitation id="17364B40FFE8083D155C0C883F97FD11" author="Lesne P." box="[1803,1923,683,706]" pageId="3" pageNumber="4" pagination="473 - 639" refId="ref6385" refString="Lesne P. 1901. Revision des Coleopteres de la famille des Bostrychides 4. Annales de la Societe Entomologique de France 69: 473 - 639." type="journal article" year="1901">Lesne, 1901</bibRefCitation>
</td>
</tr>
<tr id="CD9734F3FFE8F7C112B60CE43FA5FD0D" box="[225,1969,711,734]" gridrow="2" pageId="3" pageNumber="4" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="8E465D8FFFE8F7C112B60CE439ECFD0D" box="[225,504,711,734]" gridcol="0" gridrow="2" pageId="3" pageNumber="4">
<taxonomicName id="B4A74D32FFE8083D12BB0CE43941FD0D" authority="Lesne, 1901 Character" authorityName="Lesne" authorityYear="1901" box="[236,341,711,734]" class="Insecta" family="Bostrichidae" genus="Xylion" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFE8083D12BB0CE43941FD0D" bold="true" box="[236,341,711,734]" pageId="3" pageNumber="4">Character</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="CD9734F3FFE8F7C112B60CD03FA5FCD9" box="[225,1969,755,778]" gridrow="3" pageId="3" pageNumber="4">
<th id="8E465D8FFFE8F7C112B60CD039ECFCD9" box="[225,504,755,778]" gridcol="0" gridrow="3" pageId="3" pageNumber="4">No. of antennal segments</th>
<td id="8E465D8FFFE8F7C1105D0CD03B37FCD9" box="[522,803,755,778]" gridcol="1" gridrow="3" pageId="3" pageNumber="4">8</td>
<td id="8E465D8FFFE8F7C1117A0CD03C5CFCD9" box="[813,1096,755,778]" gridcol="2" gridrow="3" pageId="3" pageNumber="4">8</td>
<td id="8E465D8FFFE8F7C116060CD03D7FFCD9" box="[1105,1387,755,778]" gridcol="3" gridrow="3" pageId="3" pageNumber="4">10</td>
<td id="8E465D8FFFE8F7C117240CD03E99FCD9" box="[1395,1677,755,778]" gridcol="4" gridrow="3" pageId="3" pageNumber="4">10</td>
<td id="8E465D8FFFE8F7C114C00CD03FA5FCD9" box="[1687,1969,755,778]" gridcol="5" gridrow="3" pageId="3" pageNumber="4">10</td>
</tr>
<tr id="CD9734F3FFE8F7C112B60D013FA5FC84" box="[225,1969,802,855]" gridrow="4" pageId="3" pageNumber="4">
<th id="8E465D8FFFE8F7C112B60D0139ECFC84" box="[225,504,802,855]" gridcol="0" gridrow="4" pageId="3" pageNumber="4">
Funicle length relative to 1
<superScript id="84D29BF9FFE8083D13B30D0139F9FCE3" attach="left" box="[484,493,802,816]" fontSize="5" pageId="3" pageNumber="4">st</superScript>
antennomere of club
</th>
<td id="8E465D8FFFE8F7C1105D0D013B37FC84" box="[522,803,802,855]" gridcol="1" gridrow="4" pageId="3" pageNumber="4">Much shorter</td>
<td id="8E465D8FFFE8F7C1117A0D013C5CFC84" box="[813,1096,802,855]" gridcol="2" gridrow="4" pageId="3" pageNumber="4">Subequal</td>
<td id="8E465D8FFFE8F7C116060D013D7FFC84" box="[1105,1387,802,855]" gridcol="3" gridrow="4" pageId="3" pageNumber="4">A little longer</td>
<td id="8E465D8FFFE8F7C117240D013E99FC84" box="[1395,1677,802,855]" gridcol="4" gridrow="4" pageId="3" pageNumber="4">More than half the length</td>
<td id="8E465D8FFFE8F7C114C00D013FA5FC84" box="[1687,1969,802,855]" gridcol="5" gridrow="4" pageId="3" pageNumber="4">Equal in length</td>
</tr>
<tr id="CD9734F3FFE8F7C112B60D433FA5FC1D" box="[225,1969,864,974]" gridrow="5" pageId="3" pageNumber="4">
<th id="8E465D8FFFE8F7C112B60D4339ECFC1D" box="[225,504,864,974]" gridcol="0" gridrow="5" pageId="3" pageNumber="4">Sensory impressions on club</th>
<td id="8E465D8FFFE8F7C1105D0D433B37FC1D" box="[522,803,864,974]" gridcol="1" gridrow="5" pageId="3" pageNumber="4">
Two indistinct c-shaped impressions on 1
<superScript id="84D29BF9FFE8083D10970DA93ADDFC4B" attach="left" box="[704,713,906,920]" fontSize="5" pageId="3" pageNumber="4">st</superScript>
and 2
<superScript id="84D29BF9FFE8083D11530DA93B06FC4B" attach="left" box="[772,786,906,920]" fontSize="5" pageId="3" pageNumber="4">nd</superScript>
antennomeres of club
</td>
<td id="8E465D8FFFE8F7C1117A0D433C5CFC1D" box="[813,1096,864,974]" gridcol="2" gridrow="5" pageId="3" pageNumber="4">Two distinct circular sensory impressions on each anten- nomere of club including apical</td>
<td id="8E465D8FFFE8F7C116060D433D7FFC1D" box="[1105,1387,864,974]" gridcol="3" gridrow="5" pageId="3" pageNumber="4">
Two large distinct impres- sions on each face of 1
<superScript id="84D29BF9FFE8083D17650DA93D2FFC4B" attach="left" box="[1330,1339,906,920]" fontSize="5" pageId="3" pageNumber="4">st</superScript>
and 2
<superScript id="84D29BF9FFE8083D16250D843C94FC66" attach="left" box="[1138,1152,935,949]" fontSize="5" pageId="3" pageNumber="4">nd</superScript>
antennomeres of club
</td>
<td id="8E465D8FFFE8F7C117240D433E99FC1D" box="[1395,1677,864,974]" gridcol="4" gridrow="5" pageId="3" pageNumber="4">
Two zones of concentration of sensory pores on 1
<subScript id="EF2334F4FFE8083D14190DA93E43FC4B" attach="right" box="[1614,1623,906,920]" fontSize="5" pageId="3" pageNumber="4">
<superScript id="84D29BF9FFE8083D14190DA93E43FC4B" attach="left" box="[1614,1623,906,920]" fontSize="5" pageId="3" pageNumber="4">st</superScript>
</subScript>
and 2
<subScript id="EF2334F4FFE8083D17C20D843DB7FC66" attach="left" box="[1429,1443,935,949]" fontSize="5" pageId="3" pageNumber="4">
<superScript id="84D29BF9FFE8083D17C20D843DB7FC66" attach="left" box="[1429,1443,935,949]" fontSize="5" pageId="3" pageNumber="4">nd</superScript>
</subScript>
antennomeres of club
</td>
<td id="8E465D8FFFE8F7C114C00D433FA5FC1D" box="[1687,1969,864,974]" gridcol="5" gridrow="5" pageId="3" pageNumber="4">Without distinct impressions, only areas with denser pores</td>
</tr>
<tr id="CD9734F3FFE8F7C112B60DF23FA5FBED" box="[225,1969,977,1086]" gridrow="6" pageId="3" pageNumber="4">
<th id="8E465D8FFFE8F7C112B60DF239ECFBED" box="[225,504,977,1086]" gridcol="0" gridrow="6" pageId="3" pageNumber="4">Fronto-clypeal suture</th>
<td id="8E465D8FFFE8F7C1105D0DF23B37FBED" box="[522,803,977,1086]" gridcol="1" gridrow="6" pageId="3" pageNumber="4">Indistinct at sides, impressed in middle</td>
<td id="8E465D8FFFE8F7C1117A0DF23C5CFBED" box="[813,1096,977,1086]" gridcol="2" gridrow="6" pageId="3" pageNumber="4">Obsolete or scarcely visible at side, strongly deepened and foveolate in middle</td>
<td id="8E465D8FFFE8F7C116060DF23D7FFBED" box="[1105,1387,977,1086]" gridcol="3" gridrow="6" pageId="3" pageNumber="4">Indistinct at sides, impressed in middle</td>
<td id="8E465D8FFFE8F7C117240DF23E99FBED" box="[1395,1677,977,1086]" gridcol="4" gridrow="6" pageId="3" pageNumber="4">Male: distinct, foveiform in middle Female: very fine or indis- tinct</td>
<td id="8E465D8FFFE8F7C114C00DF23FA5FBED" box="[1687,1969,977,1086]" gridcol="5" gridrow="6" pageId="3" pageNumber="4">Very distinct</td>
</tr>
<tr id="CD9734F3FFE8F7C112B60A6D3FA5FBB6" box="[225,1969,1102,1125]" gridrow="7" pageId="3" pageNumber="4">
<th id="8E465D8FFFE8F7C112B60A6D39ECFBB6" box="[225,504,1102,1125]" gridcol="0" gridrow="7" pageId="3" pageNumber="4">Posterior angles of pronotum</th>
<td id="8E465D8FFFE8F7C1105D0A6D3B37FBB6" box="[522,803,1102,1125]" gridcol="1" gridrow="7" pageId="3" pageNumber="4">Without lateral carina</td>
<td id="8E465D8FFFE8F7C1117A0A6D3C5CFBB6" box="[813,1096,1102,1125]" gridcol="2" gridrow="7" pageId="3" pageNumber="4">Without lateral carina</td>
<td id="8E465D8FFFE8F7C116060A6D3D7FFBB6" box="[1105,1387,1102,1125]" gridcol="3" gridrow="7" pageId="3" pageNumber="4">Without lateral carina</td>
<td id="8E465D8FFFE8F7C117240A6D3E99FBB6" box="[1395,1677,1102,1125]" gridcol="4" gridrow="7" pageId="3" pageNumber="4">Without lateral carina</td>
<td id="8E465D8FFFE8F7C114C00A6D3FA5FBB6" box="[1687,1969,1102,1125]" gridcol="5" gridrow="7" pageId="3" pageNumber="4">With lateral carina</td>
</tr>
<tr id="CD9734F3FFE8F7C112B60A553FA5FB5E" box="[225,1969,1142,1165]" gridrow="8" pageId="3" pageNumber="4" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-5="1">
<td id="8E465D8FFFE8F7C117240A553E99FB5E" box="[1395,1677,1142,1165]" gridcol="4" gridrow="8" pageId="3" pageNumber="4">Male: elytral declivity with</td>
</tr>
<tr id="CD9734F3FFE8F7C112B60AB13FA5FB30" box="[225,1969,1170,1251]" gridrow="9" pageId="3" pageNumber="4">
<th id="8E465D8FFFE8F7C112B60AB139ECFB30" box="[225,504,1170,1251]" gridcol="0" gridrow="9" pageId="3" pageNumber="4">Elytral sculpture</th>
<td id="8E465D8FFFE8F7C1105D0AB13B37FB30" box="[522,803,1170,1251]" gridcol="1" gridrow="9" pageId="3" pageNumber="4">Each elytron with two or three costae on upper margin of declivity</td>
<td id="8E465D8FFFE8F7C1117A0AB13C5CFB30" box="[813,1096,1170,1251]" gridcol="2" gridrow="9" pageId="3" pageNumber="4">Elytral declivity with a single large spine on each elytron</td>
<td id="8E465D8FFFE8F7C116060AB13D7FFB30" box="[1105,1387,1170,1251]" gridcol="3" gridrow="9" pageId="3" pageNumber="4">Elytral declivity bidentate on each side at the upper margin</td>
<td id="8E465D8FFFE8F7C117240AB13E99FB30" box="[1395,1677,1170,1251]" gridcol="4" gridrow="9" pageId="3" pageNumber="4">or without a pair of strong spines Female: elytral declivity</td>
<td id="8E465D8FFFE8F7C114C00AB13FA5FB30" box="[1687,1969,1170,1251]" gridcol="5" gridrow="9" pageId="3" pageNumber="4">Apical declivity with 4 apophyses or marginal tubercles on each side</td>
</tr>
<tr id="CD9734F3FFE8F7C112B60ACA3FA5FAD3" box="[225,1969,1257,1280]" gridrow="10" pageId="3" pageNumber="4" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<td id="8E465D8FFFE8F7C117240ACA3FA5FAD3" box="[1395,1969,1257,1280]" colspan="2" colspanRight="1" gridcol="4" gridrow="10" pageId="3" pageNumber="4">without distinct upper margin</td>
</tr>
<tr id="CD9734F3FFE8F7C112B60B203FA5FAE5" box="[225,1969,1283,1334]" gridrow="11" pageId="3" pageNumber="4">
<th id="8E465D8FFFE8F7C112B60B2039ECFAE5" box="[225,504,1283,1334]" gridcol="0" gridrow="11" pageId="3" pageNumber="4">Margin of elytra</th>
<td id="8E465D8FFFE8F7C1105D0B203B37FAE5" box="[522,803,1283,1334]" gridcol="1" gridrow="11" pageId="3" pageNumber="4">Emarginate</td>
<td id="8E465D8FFFE8F7C1117A0B203C5CFAE5" box="[813,1096,1283,1334]" gridcol="2" gridrow="11" pageId="3" pageNumber="4">Not emarginate</td>
<td id="8E465D8FFFE8F7C116060B203D7FFAE5" box="[1105,1387,1283,1334]" gridcol="3" gridrow="11" pageId="3" pageNumber="4">Emarginate or not</td>
<td id="8E465D8FFFE8F7C117240B203E99FAE5" box="[1395,1677,1283,1334]" gridcol="4" gridrow="11" pageId="3" pageNumber="4">Male: not emarginate Female: emarginate</td>
<td id="8E465D8FFFE8F7C114C00B203FA5FAE5" box="[1687,1969,1283,1334]" gridcol="5" gridrow="11" pageId="3" pageNumber="4">Emarginate</td>
</tr>
<tr id="CD9734F3FFE8F7C112B60B1B3FA5FA83" box="[225,1969,1336,1360]" gridrow="12" pageId="3" pageNumber="4" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-5="1">
<td id="8E465D8FFFE8F7C1105D0B1B3B37FA83" box="[522,803,1336,1360]" gridcol="1" gridrow="12" pageId="3" pageNumber="4">
3
<superScript id="84D29BF9FFE8083D107D0B1B3A22FA95" attach="left" box="[554,566,1336,1350]" fontSize="5" pageId="3" pageNumber="4">rd</superScript>
ventrite thickened and
</td>
<td id="8E465D8FFFE8F7C117240B1B3E99FA83" box="[1395,1677,1336,1360]" gridcol="4" gridrow="12" pageId="3" pageNumber="4">
1
<superScript id="84D29BF9FFE8083D17DF0B1B3D85FA95" attach="left" box="[1416,1425,1336,1350]" fontSize="5" pageId="3" pageNumber="4">st</superScript>
ventrite enlarged; 4
<superScript id="84D29BF9FFE8083D14050B1B3E49FA95" attach="left" box="[1618,1629,1336,1350]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
and
</td>
</tr>
<tr id="CD9734F3FFE8F7C112B60B773FA5FA75" box="[225,1969,1364,1446]" gridrow="13" pageId="3" pageNumber="4">
<th id="8E465D8FFFE8F7C112B60B7739ECFA75" box="[225,504,1364,1446]" gridcol="0" gridrow="13" pageId="3" pageNumber="4">Modified ventrites of female abdomen</th>
<td id="8E465D8FFFE8F7C1105D0B773B37FA75" box="[522,803,1364,1446]" gridcol="1" gridrow="13" pageId="3" pageNumber="4">
modified, overlapping and concealing 4
<subScript id="EF2334F4FFE8083D10C60B523A88FAAC" attach="left" box="[657,668,1393,1407]" fontSize="5" pageId="3" pageNumber="4">th</subScript>
ventrite; lat- ter with a pair of spines on
</td>
<td id="8E465D8FFFE8F7C1117A0B773C5CFA75" box="[813,1096,1364,1446]" gridcol="2" gridrow="13" pageId="3" pageNumber="4">
4
<superScript id="84D29BF9FFE8083D11070B403B4FFAA2" attach="left" box="[848,859,1379,1393]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
ventrite with a pair of spines on posterior margin
</td>
<td id="8E465D8FFFE8F7C116060B773D7FFA75" box="[1105,1387,1364,1446]" gridcol="3" gridrow="13" pageId="3" pageNumber="4">
4
<subScript id="EF2334F4FFE8083D16080B403CA4FAA2" attach="left" box="[1119,1200,1379,1403]" fontSize="5" pageId="3" pageNumber="4">
<superScript id="84D29BF9FFE8083D16080B403C7EFAA2" attach="left" box="[1119,1130,1379,1393]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
and 5
<superScript id="84D29BF9FFE8083D16F20B403CA4FAA2" attach="left" box="[1189,1200,1379,1393]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
</subScript>
ventrites variously modified
</td>
<td id="8E465D8FFFE8F7C117240B773E99FA75" box="[1395,1677,1364,1446]" gridcol="4" gridrow="13" pageId="3" pageNumber="4">
5
<superScript id="84D29BF9FFE8083D17C70B773D8FFAB1" attach="left" box="[1424,1435,1364,1378]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
ventrites emarginate in middle posteriorly; 4
<subScript id="EF2334F4FFE8083D141E0B523E40FAAC" attach="left" box="[1609,1620,1393,1407]" fontSize="5" pageId="3" pageNumber="4">th</subScript>
ven- trite with a pair of spines in
</td>
<td id="8E465D8FFFE8F7C114C00B773FA5FA75" box="[1687,1969,1364,1446]" gridcol="5" gridrow="13" pageId="3" pageNumber="4">
1
<superScript id="84D29BF9FFE8083D14F20B403EBAFAA2" attach="left" box="[1701,1710,1379,1393]" fontSize="5" pageId="3" pageNumber="4">st</superScript>
and 2
<superScript id="84D29BF9FFE8083D14BE0B403EE3FAA2" attach="left" box="[1769,1783,1379,1393]" fontSize="5" pageId="3" pageNumber="4">nd</superScript>
ventrites enlarged; 2
<subScript id="EF2334F4FFE8083D14FC0BA33EADFA5D" attach="right" box="[1707,1721,1408,1422]" fontSize="5" pageId="3" pageNumber="4">
<superScript id="84D29BF9FFE8083D14FC0BA33EADFA5D" attach="left" box="[1707,1721,1408,1422]" fontSize="5" pageId="3" pageNumber="4">nd</superScript>
</subScript>
to 5
<subScript id="EF2334F4FFE8083D14B40BA33EFAFA5D" attach="left" box="[1763,1774,1408,1422]" fontSize="5" pageId="3" pageNumber="4">
<superScript id="84D29BF9FFE8083D14B40BA33EFAFA5D" attach="left" box="[1763,1774,1408,1422]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
</subScript>
variously modified
</td>
</tr>
<tr id="CD9734F3FFE8F7C112B60B8F3FA5FA10" box="[225,1969,1452,1475]" gridrow="14" pageId="3" pageNumber="4" rowspan-0="1" rowspan-2="1" rowspan-3="1" rowspan-5="1">
<td id="8E465D8FFFE8F7C1105D0B8F3B37FA10" box="[522,803,1452,1475]" gridcol="1" gridrow="14" pageId="3" pageNumber="4">posterior margin</td>
<td id="8E465D8FFFE8F7C117240B8F3E99FA10" box="[1395,1677,1452,1475]" gridcol="4" gridrow="14" pageId="3" pageNumber="4">some species</td>
</tr>
<tr id="CD9734F3FFE8F7C112B60BE43FA5FA2F" box="[225,1969,1479,1532]" gridrow="15" pageId="3" pageNumber="4">
<th id="8E465D8FFFE8F7C112B60BE439ECFA2F" box="[225,504,1479,1532]" gridcol="0" gridrow="15" pageId="3" pageNumber="4">Male abdomen</th>
<td id="8E465D8FFFE8F7C1105D0BE43B37FA2F" box="[522,803,1479,1532]" gridcol="1" gridrow="15" pageId="3" pageNumber="4">
5
<superScript id="84D29BF9FFE8083D10490BE43A3DFA06" attach="left" box="[542,553,1479,1493]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
ventrite with well-devel- oped pleural pieces
</td>
<td id="8E465D8FFFE8F7C1117A0BE43C5CFA2F" box="[813,1096,1479,1532]" gridcol="2" gridrow="15" pageId="3" pageNumber="4">
5
<superScript id="84D29BF9FFE8083D11100BE43B46FA06" attach="left" box="[839,850,1479,1493]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
ventrite without pleural pieces
</td>
<td id="8E465D8FFFE8F7C116060BE43D7FFA2F" box="[1105,1387,1479,1532]" gridcol="3" gridrow="15" pageId="3" pageNumber="4">
5
<superScript id="84D29BF9FFE8083D16320BE43C64FA06" attach="left" box="[1125,1136,1479,1493]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
ventrite with well-devel- oped pleural pieces
</td>
<td id="8E465D8FFFE8F7C117240BE43E99FA2F" box="[1395,1677,1479,1532]" gridcol="4" gridrow="15" pageId="3" pageNumber="4">
5
<superScript id="84D29BF9FFE8083D17D40BE43D9AFA06" attach="left" box="[1411,1422,1479,1493]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
ventrite with wide pleural pieces
</td>
<td id="8E465D8FFFE8F7C114C00BE43FA5FA2F" box="[1687,1969,1479,1532]" gridcol="5" gridrow="15" pageId="3" pageNumber="4">
5
<superScript id="84D29BF9FFE8083D14F00BE43EA6FA06" attach="left" box="[1703,1714,1479,1493]" fontSize="5" pageId="3" pageNumber="4">th</superScript>
ventrite with more or less developed pleural pieces
</td>
</tr>
<tr id="CD9734F3FFE8F7C112B608223FA5F981" box="[225,1969,1537,1618]" gridrow="16" pageId="3" pageNumber="4">
<th id="8E465D8FFFE8F7C112B6082239ECF981" box="[225,504,1537,1618]" gridcol="0" gridrow="16" pageId="3" pageNumber="4">Protibia</th>
<td id="8E465D8FFFE8F7C1105D08223D7FF981" box="[522,1387,1537,1618]" colspan="3" colspanRight="2" gridcol="1" gridrow="16" pageId="3" pageNumber="4">Wider at apex than in Wider at apex than in middle, Wider at apex than in middle, middle, external face more or external face flattened external face flattened less flattened</td>
<td id="8E465D8FFFE8F7C1172408223FA5F981" box="[1395,1969,1537,1618]" colspan="2" colspanRight="1" gridcol="4" gridrow="16" pageId="3" pageNumber="4">Wider at apex than in Widest towards middle, middle, external face more or without a flat face externally less flattened</td>
</tr>
<tr id="CD9734F3FFE8F7C112B608763FA5F901" box="[225,1969,1621,1746]" gridrow="17" pageId="3" pageNumber="4">
<th id="8E465D8FFFE8F7C112B6087639ECF901" box="[225,504,1621,1746]" gridcol="0" gridrow="17" pageId="3" pageNumber="4">Protarsi</th>
<td id="8E465D8FFFE8F7C1105D08763B37F901" box="[522,803,1621,1746]" gridcol="1" gridrow="17" pageId="3" pageNumber="4">Male: with shorter and straighter hairs. Female: with long, curving hairs on ventral side</td>
<td id="8E465D8FFFE8F7C1117A08763C5CF901" box="[813,1096,1621,1746]" gridcol="2" gridrow="17" pageId="3" pageNumber="4">Male: with shorter and straighter hairs. Female: with long, curving hairs on ventral side</td>
<td id="8E465D8FFFE8F7C1160608763D7FF901" box="[1105,1387,1621,1746]" gridcol="3" gridrow="17" pageId="3" pageNumber="4">A fringe of short hairs on ventral side in both sexes</td>
<td id="8E465D8FFFE8F7C1172408763E99F901" box="[1395,1677,1621,1746]" gridcol="4" gridrow="17" pageId="3" pageNumber="4">Male: without modified hairs. Female: with long erect hairs on inner side</td>
<td id="8E465D8FFFE8F7C114C008763FA5F901" box="[1687,1969,1621,1746]" gridcol="5" gridrow="17" pageId="3" pageNumber="4">Male: with fairly long but straight hairs. Female: with longer, curving hairs on ventral side</td>
</tr>
<tr id="CD9734F3FFE8F7C112B608FD3FA5F926" box="[225,1969,1758,1781]" gridrow="18" pageId="3" pageNumber="4">
<th id="8E465D8FFFE8F7C112B608FD39ECF926" box="[225,504,1758,1781]" gridcol="0" gridrow="18" pageId="3" pageNumber="4">Geographical distribution</th>
<td id="8E465D8FFFE8F7C1105D08FD3B37F926" box="[522,803,1758,1781]" gridcol="1" gridrow="18" pageId="3" pageNumber="4">Oriental</td>
<td id="8E465D8FFFE8F7C1117A08FD3C5CF926" box="[813,1096,1758,1781]" gridcol="2" gridrow="18" pageId="3" pageNumber="4">Oriental</td>
<td id="8E465D8FFFE8F7C1160608FD3D7FF926" box="[1105,1387,1758,1781]" gridcol="3" gridrow="18" pageId="3" pageNumber="4">Afrotropical</td>
<td id="8E465D8FFFE8F7C1172408FD3E99F926" box="[1395,1677,1758,1781]" gridcol="4" gridrow="18" pageId="3" pageNumber="4">Australian</td>
<td id="8E465D8FFFE8F7C114C008FD3FA5F926" box="[1687,1969,1758,1781]" gridcol="5" gridrow="18" pageId="3" pageNumber="4">Afrotropical</td>
</tr>
</table>
</paragraph>
<paragraph id="731836B1FFEF083A12B40F123CADFE66" blockId="4.[227,1436,305,437]" pageId="4" pageNumber="5">Scutellum, small, punctate. Elytra subequal to pronotum in width, strongly convex, smooth, shining, indistinctly punctured anteriorly, the punctures larger and deeper near the declivity, posterior part of disc with more than one pair of costae, each costa ending in a small tooth or projection at upper margin of declivity; declivity weakly convex, glabrous, the postero-lateral margin emarginate.</paragraph>
<paragraph id="731836B1FFEF083A12B40FF43D60FDC0" blockId="4.[227,1436,470,532]" pageId="4" pageNumber="5">Legs subequal in length, procoxae contiguous, mesocoxae narrowly separated; tibiae expanded toward apices, protarsi equal in length to protibiae, meso- and meta-tarsi longer than their respective tibiae.</paragraph>
<paragraph id="731836B1FFEF083A12B40C163B18FD46" blockId="4.[227,1437,565,661]" pageId="4" pageNumber="5">Intercoxal process of first abdominal ventrite lamelliform. Female with third abdominal ventrite projecting over and concealing fourth ventrite, its posterior margin with lobes or teeth. Last visible abdominal ventrite of male with pleural pieces.</paragraph>
</subSubSection>
<subSubSection id="3BBD653AFFEF083B12B40C943AA8FE25" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="discussion">
<paragraph id="731836B1FFEF083A12B40C943941FD02" blockId="4.[227,341,695,721]" box="[227,341,695,721]" pageId="4" pageNumber="5">
<heading id="285081DDFFEF083A12B40C943941FD02" bold="true" box="[227,341,695,721]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="41D3EAA3FFEF083A12B40C943941FD02" bold="true" box="[227,341,695,721]" pageId="4" pageNumber="5">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="731836B1FFEF083A12B40CC639A5FAE0" blockId="4.[227,1437,741,1331]" pageId="4" pageNumber="5">
The genus
<taxonomicName id="B4A74D32FFEF083A13310CC53A15FCD3" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[358,513,742,768]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="41D3EAA3FFEF083A13310CC53A15FCD3" box="[358,513,742,768]" italics="true" pageId="4" pageNumber="5">Octomeristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AE057D8FFEF083A105E0CC53A61FCD3" box="[521,629,742,768]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
(abbreviated here as
<emphasis id="41D3EAA3FFEF083A11230CC53BB2FCD3" box="[884,934,742,768]" italics="true" pageId="4" pageNumber="5">Om.</emphasis>
) is noteworthy because the antennae have only eight segments. The only other genus of
<taxonomicName id="B4A74D32FFEF083A11510D2B3B88FCF1" authorityName="Lesne" authorityYear="1921" box="[774,924,776,802]" class="Insecta" family="Bostrichidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Xyloperthini">Xyloperthini</taxonomicName>
with 8-segmented antennae is
<taxonomicName id="B4A74D32FFEF083A175A0D2B393DFC95" authority="Lesne" authorityName=", Lesne" authorityYear="1901" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A175A0D2B3D88FCF1" box="[1293,1436,776,802]" italics="true" pageId="4" pageNumber="5">Octodesmus</emphasis>
Lesne
</taxonomicName>
(abbreviated here as
<emphasis id="41D3EAA3FFEF083A107C0D083A42FC96" box="[555,598,811,837]" italics="true" pageId="4" pageNumber="5">Od.</emphasis>
). Other xyloperthine genera have 9- to 11-segmented antennae. There are presently four species in
<taxonomicName id="B4A74D32FFEF083A10780D6C3AAAFCBA" authorityName=", Lesne" authorityYear="1901" box="[559,702,847,873]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A10780D6C3AAAFCBA" box="[559,702,847,873]" italics="true" pageId="4" pageNumber="5">Octodesmus</emphasis>
</taxonomicName>
: the type species,
<taxonomicName id="B4A74D32FFEF083A11C40D6C3CEDFCBA" authority="(Lesne, 1901)" baseAuthorityName="Lesne" baseAuthorityYear="1901" box="[915,1273,847,873]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="episternalis">
<emphasis id="41D3EAA3FFEF083A11C40D6C3C5AFCBA" box="[915,1102,847,873]" italics="true" pageId="4" pageNumber="5">Od. episternalis</emphasis>
(
<bibRefCitation id="17364B40FFEF083A160B0D6C3CE4FCBA" author="Lesne P." box="[1116,1264,847,873]" pageId="4" pageNumber="5" pagination="473 - 639" refId="ref6385" refString="Lesne P. 1901. Revision des Coleopteres de la famille des Bostrychides 4. Annales de la Societe Entomologique de France 69: 473 - 639." type="journal article" year="1901">Lesne, 1901</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="B4A74D32FFEF083A17530D6C3993FC5F" authority="(Lesne, 1897)" baseAuthorityName="Lesne" baseAuthorityYear="1897" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="41D3EAA3FFEF083A17530D6C3D89FCBA" box="[1284,1437,847,873]" italics="true" pageId="4" pageNumber="5">Od. parvulus</emphasis>
(Lesne, 1897)
</taxonomicName>
,
<taxonomicName id="B4A74D32FFEF083A13C60D513B14FC5F" authority="(Lesne, 1932)" baseAuthorityName="Lesne" baseAuthorityYear="1932" box="[401,768,882,908]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="41D3EAA3FFEF083A13C60D513A4EFC5F" box="[401,602,882,908]" italics="true" pageId="4" pageNumber="5">Od. minutissimus</emphasis>
(
<bibRefCitation id="17364B40FFEF083A103F0D513AECFC5F" author="Lesne P." box="[616,760,882,908]" pageId="4" pageNumber="5" pagination="651 - 663" refId="ref6470" refString="Lesne P. 1932. Notes sur les Coleopteres Terediles 22. Diagnoses de Bostrychides nouveaux de l'Asie orientale. Bulletin du Museum national d'Histoire naturelle 1932: 651 - 663." type="journal article" year="1932">Lesne, 1932</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="B4A74D32FFEF083A11610D513C5FFC5F" authority="Chujo, 1964" authorityName="Chujo" authorityYear="1964" box="[822,1099,882,908]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kamoli">
<emphasis id="41D3EAA3FFEF083A11610D513BA2FC5F" box="[822,950,882,908]" italics="true" pageId="4" pageNumber="5">Od. kamoli</emphasis>
<bibRefCitation id="17364B40FFEF083A11EB0D513C5FFC5F" author="Chujo M." box="[956,1099,882,908]" pageId="4" pageNumber="5" pagination="201 - 215" refId="ref6301" refString="Chujo M. 1964. Coleoptera from Southeast Asia (III). 17. Family Bostrichidae. Nature and Life in Southeast Asia 3: 201 - 215." type="journal article" year="1964">Chûjô, 1964</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="17364B40FFEF083A160F0D513937FC7C" author="Borowski J. &amp; Wegrzynowicz P." pageId="4" pageNumber="5" refId="ref6279" refString="Borowski J. &amp; Wegrzynowicz P. 2007. World catalogue of Bostrichidae (Coleoptera). Wydawnictwo Mantis, Olsztyn." type="book" year="2007">Borowski &amp; Wegrzynowicz 2007</bibRefCitation>
). The senior author has examined
<typeStatus id="AC1C8813FFEF083A109B0DB53B27FC63" box="[716,819,918,944]" pageId="4" pageNumber="5" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="B4A74D32FFEF083A11360DB63C35FC7C" baseAuthorityName="Lesne" baseAuthorityYear="1901" box="[865,1057,917,943]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="episternalis">
<emphasis id="41D3EAA3FFEF083A11360DB63C35FC7C" box="[865,1057,917,943]" italics="true" pageId="4" pageNumber="5">Od. episternalis</emphasis>
</taxonomicName>
(MNHN), the
<typeStatus id="AC1C8813FFEF083A168C0DB63D56FC7C" box="[1243,1346,917,943]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
of
<taxonomicName id="B4A74D32FFEF083A17260DB6395EFC01" baseAuthorityName="Lesne" baseAuthorityYear="1897" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="parvulus">
<emphasis id="41D3EAA3FFEF083A17260DB6395EFC01" italics="true" pageId="4" pageNumber="5">Od. parvulus</emphasis>
</taxonomicName>
(MNHN), and the
<typeStatus id="AC1C8813FFEF083A106B0D9B3AB7FC01" box="[572,675,952,978]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
of
<taxonomicName id="B4A74D32FFEF083A10840D9B3B5AFC01" authorityName="Chujo" authorityYear="1964" box="[723,846,952,978]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="kamoli">
<emphasis id="41D3EAA3FFEF083A10840D9B3B5AFC01" box="[723,846,952,978]" italics="true" pageId="4" pageNumber="5">Od.kamoli</emphasis>
</taxonomicName>
(KUM). We have been unable to examine type material of
<taxonomicName id="B4A74D32FFEF083A13250DF83A55FC26" baseAuthorityName="Lesne" baseAuthorityYear="1932" box="[370,577,987,1013]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="41D3EAA3FFEF083A13250DF83A55FC26" box="[370,577,987,1013]" italics="true" pageId="4" pageNumber="5">Od. minutissimus</emphasis>
</taxonomicName>
. However, it has been possible to show (see discussion below) that it should be transferred to
<taxonomicName id="B4A74D32FFEF083A105F0DDC3AB7FBCA" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[520,675,1023,1049]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="41D3EAA3FFEF083A105F0DDC3AB7FBCA" box="[520,675,1023,1049]" italics="true" pageId="4" pageNumber="5">Octomeristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AE057D8FFEF083A10FB0DDC3B0FFBCA" box="[684,795,1023,1049]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
, and it is considered here as a member of that genus. When
<bibRefCitation id="17364B40FFEF083A13640A0139C5FBEF" author="Lesne P." box="[307,465,1058,1084]" pageId="4" pageNumber="5" pagination="651 - 663" refId="ref6470" refString="Lesne P. 1932. Notes sur les Coleopteres Terediles 22. Diagnoses de Bostrychides nouveaux de l'Asie orientale. Bulletin du Museum national d'Histoire naturelle 1932: 651 - 663." type="journal article" year="1932">Lesne (1932)</bibRefCitation>
described
<taxonomicName id="B4A74D32FFEF083A10010A013B9CFBEF" baseAuthorityName="Lesne" baseAuthorityYear="1932" box="[598,904,1058,1084]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="minutissimus">
<emphasis id="41D3EAA3FFEF083A10010A013B9CFBEF" box="[598,904,1058,1084]" italics="true" pageId="4" pageNumber="5">Octodesmus minutissimus</emphasis>
</taxonomicName>
, he suggested that it formed a link between
<taxonomicName id="B4A74D32FFEF083A12B40A663966FB8C" authorityName=", Lesne" authorityYear="1901" box="[227,370,1093,1119]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A12B40A663966FB8C" box="[227,370,1093,1119]" italics="true" pageId="4" pageNumber="5">Octodesmus</emphasis>
</taxonomicName>
on the one hand, and the genera
<taxonomicName id="B4A74D32FFEF083A10BC0A663BDBFB8C" authority="Lesne, 1901" authorityName="Lesne" authorityYear="1901" box="[747,975,1093,1120]" class="Insecta" family="Bostrichidae" genus="Xylion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A10BC0A663B2CFB8C" box="[747,824,1093,1119]" italics="true" pageId="4" pageNumber="5">Xylion</emphasis>
<bibRefCitation id="17364B40FFEF083A11690A653BDBFB8C" author="Lesne P." box="[830,975,1093,1120]" pageId="4" pageNumber="5" pagination="473 - 639" refId="ref6385" refString="Lesne P. 1901. Revision des Coleopteres de la famille des Bostrychides 4. Annales de la Societe Entomologique de France 69: 473 - 639." type="journal article" year="1901">Lesne, 1901</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="B4A74D32FFEF083A118D0A663CF8FB8C" authority="Lesne, 1901" authorityName="Lesne" authorityYear="1901" box="[986,1260,1093,1120]" class="Insecta" family="Bostrichidae" genus="Xylionulus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A118D0A663C43FB8C" box="[986,1111,1093,1119]" italics="true" pageId="4" pageNumber="5">Xylionulus</emphasis>
<bibRefCitation id="17364B40FFEF083A160A0A653CF8FB8C" author="Lesne P." box="[1117,1260,1093,1120]" pageId="4" pageNumber="5" pagination="473 - 639" refId="ref6385" refString="Lesne P. 1901. Revision des Coleopteres de la famille des Bostrychides 4. Annales de la Societe Entomologique de France 69: 473 - 639." type="journal article" year="1901">Lesne, 1901</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="B4A74D32FFEF083A17750A663967FB51" authority="Lesne, 1901" authorityName="Lesne" authorityYear="1901" class="Insecta" family="Bostrichidae" genus="Xylobosca" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A17750A663D88FB8C" box="[1314,1436,1093,1119]" italics="true" pageId="4" pageNumber="5">Xylobosca</emphasis>
<bibRefCitation id="17364B40FFEF083A12B40A4A3967FB51" author="Lesne P." box="[227,371,1128,1155]" pageId="4" pageNumber="5" pagination="473 - 639" refId="ref6385" refString="Lesne P. 1901. Revision des Coleopteres de la famille des Bostrychides 4. Annales de la Societe Entomologique de France 69: 473 - 639." type="journal article" year="1901">Lesne, 1901</bibRefCitation>
</taxonomicName>
on the other. Since then, the genus
<taxonomicName id="B4A74D32FFEF083A114B0A4B3B7DFB51" authorityName="Lesne" authorityYear="1901" box="[796,873,1128,1154]" class="Insecta" family="Bostrichidae" genus="Xylion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A114B0A4B3B7DFB51" box="[796,873,1128,1154]" italics="true" pageId="4" pageNumber="5">Xylion</emphasis>
</taxonomicName>
has been split up (
<bibRefCitation id="17364B40FFEF083A161F0A4B3CC3FB51" author="Lesne P." box="[1096,1239,1128,1154]" pageId="4" pageNumber="5" pagination="137 - 152" refId="ref6502" refString="Lesne P. 1940. Sur quelques Bostrychides Indiens. Annales de la Societe Entomologique de France 109: 137 - 152." type="journal article" year="1940">Lesne 1940</bibRefCitation>
;
<bibRefCitation id="17364B40FFEF083A16B40A4B3D9AFB50" author="Vrydagh J. M." box="[1251,1422,1128,1155]" pageId="4" pageNumber="5" pagination="1 - 23" refId="ref6697" refString="Vrydagh J. M. 1955. Contribution a l'etude des Bostrychidae Coleoptera Teredilia 3. Les Bostrychides de Mozambique. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 31 (16): 1 - 23." type="journal article" year="1955">Vrydagh 1955</bibRefCitation>
), and the genus restricted to a more closely related group of seven species (
<bibRefCitation id="17364B40FFEF083A16370AA83D19FB75" author="Vrydagh J. M." box="[1120,1293,1163,1190]" pageId="4" pageNumber="5" pagination="1 - 23" refId="ref6697" refString="Vrydagh J. M. 1955. Contribution a l'etude des Bostrychidae Coleoptera Teredilia 3. Les Bostrychides de Mozambique. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 31 (16): 1 - 23." type="journal article" year="1955">Vrydagh 1955</bibRefCitation>
). Following
<bibRefCitation id="17364B40FFEF083A12B40A8C3981FB1A" author="Lesne P." box="[227,405,1199,1225]" pageId="4" pageNumber="5" pagination="651 - 663" refId="ref6470" refString="Lesne P. 1932. Notes sur les Coleopteres Terediles 22. Diagnoses de Bostrychides nouveaux de l'Asie orientale. Bulletin du Museum national d'Histoire naturelle 1932: 651 - 663." type="journal article" year="1932">Lesnes (1932)</bibRefCitation>
suggestion, we compare
<taxonomicName id="B4A74D32FFEF083A10980A8C3B7EFB1A" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[719,874,1199,1225]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="41D3EAA3FFEF083A10980A8C3B7EFB1A" box="[719,874,1199,1225]" italics="true" pageId="4" pageNumber="5">Octomeristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AE057D8FFEF083A11220A8C3BF0FB1A" box="[885,996,1199,1225]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
(including
<taxonomicName id="B4A74D32FFEF083A16250A8C3D5CFB1A" baseAuthorityName="Lesne" baseAuthorityYear="1932" box="[1138,1352,1199,1225]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="minutissimus" status="comb. nov.">
<emphasis id="41D3EAA3FFEF083A16250A8C3D5CFB1A" box="[1138,1352,1199,1225]" italics="true" pageId="4" pageNumber="5">Om. minutissimus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AE057D8FFEF083A17030A8C3902FB3F" pageId="4" pageNumber="5" rank="species">comb. nov.</taxonomicNameLabel>
) to the four genera listed by him, using
<taxonomicName id="B4A74D32FFEF083A10B90AF13B2FFB3F" authorityName="Lesne" authorityYear="1901" box="[750,827,1234,1260]" class="Insecta" family="Bostrichidae" genus="Xylion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A10B90AF13B2FFB3F" box="[750,827,1234,1260]" italics="true" pageId="4" pageNumber="5">Xylion</emphasis>
</taxonomicName>
in the restricted sense of
<bibRefCitation id="17364B40FFEF083A163F0AF13D37FB3E" author="Vrydagh J. M." box="[1128,1315,1234,1261]" pageId="4" pageNumber="5" pagination="1 - 23" refId="ref6697" refString="Vrydagh J. M. 1955. Contribution a l'etude des Bostrychidae Coleoptera Teredilia 3. Les Bostrychides de Mozambique. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 31 (16): 1 - 23." type="journal article" year="1955">Vrydagh (1955)</bibRefCitation>
(
<tableCitation id="3E25030AFFEF083A17650AF13D9EFB3F" box="[1330,1418,1234,1260]" captionStart="Table 1" captionStartId="3.[219,288,581,609]" captionTargetBox="[225,1969,654,1781]" captionTargetId="graphics@3.[219,1975,644,1790]" captionTargetPageId="3" captionText="Table 1. Comparison of characters of Octomeristes Liu &amp; Beaver with selected genera of Xyloperthini." httpUri="http://table.plazi.org/id/27D86639FFE8083D128C0C663D8DFDB2" pageId="4" pageNumber="5" tableUuid="27D86639FFE8083D128C0C663D8DFDB2">Table 1</tableCitation>
). A more wide-ranging study of the genera of
<taxonomicName id="B4A74D32FFEF083A10BE0AD63B6BFADC" authorityName="Lesne" authorityYear="1921" box="[745,895,1269,1295]" class="Insecta" family="Bostrichidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Xyloperthini">Xyloperthini</taxonomicName>
would be valuable, but is beyond the scope of the present paper.
</paragraph>
<paragraph id="731836B1FFEF083A12B40B773B01F95B" blockId="4.[227,1437,1364,1672]" pageId="4" pageNumber="5">
The genus
<taxonomicName id="B4A74D32FFEF083A13390B773A1DFABD" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[366,521,1364,1390]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="41D3EAA3FFEF083A13390B773A1DFABD" box="[366,521,1364,1390]" italics="true" pageId="4" pageNumber="5">Octomeristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AE057D8FFEF083A10410B773A92FABD" box="[534,646,1364,1390]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
can be distinguished from all other xyloperthine genera by the combination of the 8-segmented antenna, with the funicle length much shorter than the first antennomere of the antennal club, and the thickened and modified third abdominal ventrite, which overlaps and conceals the fourth ventrite.
<taxonomicName id="B4A74D32FFEF083A106B0B9D3AC3FA0B" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[572,727,1470,1496]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="41D3EAA3FFEF083A106B0B9D3AC3FA0B" box="[572,727,1470,1496]" italics="true" pageId="4" pageNumber="5">Octomeristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AE057D8FFEF083A10B60B9D3B5AFA0B" box="[737,846,1470,1496]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
is further distinguished from
<taxonomicName id="B4A74D32FFEF083A16EF0B9E3D53FA04" authorityName=", Lesne" authorityYear="1901" box="[1208,1351,1469,1495]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A16EF0B9E3D53FA04" box="[1208,1351,1469,1495]" italics="true" pageId="4" pageNumber="5">Octodesmus</emphasis>
</taxonomicName>
by the absence of sensory impressions on the apical segment of the antennal club (present in
<taxonomicName id="B4A74D32FFEF083A16860BC33D74FA29" authorityName=", Lesne" authorityYear="1901" box="[1233,1376,1504,1530]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A16860BC33D74FA29" box="[1233,1376,1504,1530]" italics="true" pageId="4" pageNumber="5">Octodesmus</emphasis>
</taxonomicName>
), the sculpture of the elytra (
<tableCitation id="3E25030AFFEF083A13A208273A59F9CD" box="[501,589,1540,1566]" captionStart="Table 1" captionStartId="3.[219,288,581,609]" captionTargetBox="[225,1969,654,1781]" captionTargetId="graphics@3.[219,1975,644,1790]" captionTargetPageId="3" captionText="Table 1. Comparison of characters of Octomeristes Liu &amp; Beaver with selected genera of Xyloperthini." httpUri="http://table.plazi.org/id/27D86639FFE8083D128C0C663D8DFDB2" pageId="4" pageNumber="5" tableUuid="27D86639FFE8083D128C0C663D8DFDB2">Table 1</tableCitation>
), and the presence in the male of well-developed pleural pieces (absent in
<taxonomicName id="B4A74D32FFEF083A135508043985F992" authorityName=", Lesne" authorityYear="1901" box="[258,401,1575,1601]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A135508043985F992" box="[258,401,1575,1601]" italics="true" pageId="4" pageNumber="5">Octodesmus</emphasis>
</taxonomicName>
). It is likely that the reduction in the number of antennal segments in the two genera is related to the small size of the species, and is not an indication of a close relationship.
<emphasis id="41D3EAA3FFEF083A169208693D88F9B7" box="[1221,1436,1610,1636]" italics="true" pageId="4" pageNumber="5">
<taxonomicName id="B4A74D32FFEF083A169208693D01F9B7" authorityName="Lesne" authorityYear="1901" box="[1221,1301,1610,1636]" class="Insecta" family="Bostrichidae" genus="Xylion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Xylion</taxonomicName>
,
<taxonomicName id="B4A74D32FFEF083A174808693D88F9B7" authorityName="Lesne" authorityYear="1901" box="[1311,1436,1610,1636]" class="Insecta" family="Bostrichidae" genus="Xylionulus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Xylionulus</taxonomicName>
</emphasis>
and
<taxonomicName id="B4A74D32FFEF083A1343084E399AF954" authorityName="Lesne" authorityYear="1901" box="[276,398,1645,1671]" class="Insecta" family="Bostrichidae" genus="Xylobosca" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A1343084E399AF954" box="[276,398,1645,1671]" italics="true" pageId="4" pageNumber="5">Xylobosca</emphasis>
</taxonomicName>
all have 10-segmented antennae.
</paragraph>
<paragraph id="731836B1FFEF083A12B4088A3B19F86A" blockId="4.[227,1437,1705,1977]" pageId="4" pageNumber="5">
The genus
<taxonomicName id="B4A74D32FFEF083A1335088A39C8F910" authorityName="Lesne" authorityYear="1901" box="[354,476,1705,1731]" class="Insecta" family="Bostrichidae" genus="Xylobosca" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A1335088A39C8F910" box="[354,476,1705,1731]" italics="true" pageId="4" pageNumber="5">Xylobosca</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="B4A74D32FFEF083A1088088A3B6EF910" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[735,890,1705,1731]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="41D3EAA3FFEF083A1088088A3B6EF910" box="[735,890,1705,1731]" italics="true" pageId="4" pageNumber="5">Octomeristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AE057D8FFEF083A11D6088A3BFFF910" box="[897,1003,1705,1731]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
and the other genera considered here by the form of the protibia, which is atypical for the
<taxonomicName id="B4A74D32FFEF083A111308EF3BC9F935" authorityName="Lesne" authorityYear="1921" box="[836,989,1740,1766]" class="Insecta" family="Bostrichidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="tribe" tribe="Xyloperthini">Xyloperthini</taxonomicName>
, and is widest towards the middle (not at the apex), and lacks a flat external face. The sexual dimorphism of the elytra of
<taxonomicName id="B4A74D32FFEF083A16C508CC3D18F8DA" authorityName="Lesne" authorityYear="1901" box="[1170,1292,1775,1801]" class="Insecta" family="Bostrichidae" genus="Xylobosca" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A16C508CC3D18F8DA" box="[1170,1292,1775,1801]" italics="true" pageId="4" pageNumber="5">Xylobosca</emphasis>
</taxonomicName>
is also quite different from
<taxonomicName id="B4A74D32FFEF083A13C709303A3FF8FE" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[400,555,1811,1837]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="41D3EAA3FFEF083A13C709303A3FF8FE" box="[400,555,1811,1837]" italics="true" pageId="4" pageNumber="5">Octomeristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AE057D8FFEF083A106509303A8AF8FE" box="[562,670,1811,1837]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
However, it is interesting to note the presence of a pair of spines on the fourth ventrite of the female in some species of
<taxonomicName id="B4A74D32FFEF083A11D409163BE9F89C" authorityName="Lesne" authorityYear="1901" box="[899,1021,1845,1871]" class="Insecta" family="Bostrichidae" genus="Xylobosca" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A11D409163BE9F89C" box="[899,1021,1845,1871]" italics="true" pageId="4" pageNumber="5">Xylobosca</emphasis>
</taxonomicName>
, as in
<taxonomicName id="B4A74D32FFEF083A160409153CFAF883" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[1107,1262,1846,1872]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="41D3EAA3FFEF083A160409153CFAF883" box="[1107,1262,1846,1872]" italics="true" pageId="4" pageNumber="5">Octomeristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AE057D8FFEF083A16AF09153D71F883" box="[1272,1381,1846,1872]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
The genus
<taxonomicName id="B4A74D32FFEF083A1379097A396FF8A0" authorityName="Lesne" authorityYear="1901" box="[302,379,1881,1907]" class="Insecta" family="Bostrichidae" genus="Xylion" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A1379097A396FF8A0" box="[302,379,1881,1907]" italics="true" pageId="4" pageNumber="5">Xylion</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="B4A74D32FFEF083A1028097A3B0EF8A0" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[639,794,1881,1907]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="41D3EAA3FFEF083A1028097A3B0EF8A0" box="[639,794,1881,1907]" italics="true" pageId="4" pageNumber="5">Octomeristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AE057D8FFEF083A1176097A3B9FF8A0" box="[801,907,1881,1907]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
and the other three genera by the presence of a lateral carina at the posterior angles of the pronotum, and the remarkable enlargement and modification of the second abdominal ventrite of the female.
</paragraph>
<paragraph id="731836B1FFEF083A12B409F93D88F7CB" blockId="4.[227,1436,2010,2037]" lastBlockId="4.[1244,1436,2046,2072]" pageId="4" pageNumber="5">
The genus that is morphologically most similar to
<taxonomicName id="B4A74D32FFEF083A117209F83BD4F826" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[805,960,2011,2037]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="41D3EAA3FFEF083A117209F83BD4F826" box="[805,960,2011,2037]" italics="true" pageId="4" pageNumber="5">Octomeristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AE057D8FFEF083A119209F83C3AF826" box="[965,1070,2011,2037]" pageId="4" pageNumber="5" rank="genus">gen. nov.</taxonomicNameLabel>
is
<taxonomicName id="B4A74D32FFEF083A161A09F93CDEF827" authorityName="Lesne" authorityYear="1901" box="[1101,1226,2010,2036]" class="Insecta" family="Bostrichidae" genus="Xylionulus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEF083A161A09F93CDEF827" box="[1101,1226,2010,2036]" italics="true" pageId="4" pageNumber="5">Xylionulus</emphasis>
</taxonomicName>
, with very similar distinguished by
</paragraph>
<paragraph id="731836B1FFEF083B12B409DD3B36FEBD" blockId="4.[227,1237,2046,2072]" lastBlockId="5.[227,1436,305,367]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
characters of the frons, pronotum and elytra (
<tableCitation id="3E25030AFFEF083A10B809DD3B53F7CB" box="[751,839,2046,2072]" captionStart="Table 1" captionStartId="3.[219,288,581,609]" captionTargetBox="[225,1969,654,1781]" captionTargetId="graphics@3.[219,1975,644,1790]" captionTargetPageId="3" captionText="Table 1. Comparison of characters of Octomeristes Liu &amp; Beaver with selected genera of Xyloperthini." httpUri="http://table.plazi.org/id/27D86639FFE8083D128C0C663D8DFDB2" pageId="4" pageNumber="5" tableUuid="27D86639FFE8083D128C0C663D8DFDB2">Table 1</tableCitation>
). However, the two genera can be characters of the antennae (
<tableCitation id="3E25030AFFEE083B10730F123A68FE98" box="[548,636,305,331]" captionStart="Table 1" captionStartId="3.[219,288,581,609]" captionTargetBox="[225,1969,654,1781]" captionTargetId="graphics@3.[219,1975,644,1790]" captionTargetPageId="3" captionText="Table 1. Comparison of characters of Octomeristes Liu &amp; Beaver with selected genera of Xyloperthini." httpUri="http://table.plazi.org/id/27D86639FFE8083D128C0C663D8DFDB2" pageId="5" pageNumber="6" tableUuid="27D86639FFE8083D128C0C663D8DFDB2">Table 1</tableCitation>
), and the modification of the fourth and fifth ventrites of the female abdomen in
<taxonomicName id="B4A74D32FFEE083B13230F7739E5FEBD" authorityName="Lesne" authorityYear="1901" box="[372,497,340,366]" class="Insecta" family="Bostrichidae" genus="Xylionulus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEE083B13230F7739E5FEBD" box="[372,497,340,366]" italics="true" pageId="5" pageNumber="6">Xylionulus</emphasis>
</taxonomicName>
and not the third ventrite.
</paragraph>
<paragraph id="731836B1FFEE083B12B40FB53AA8FE25" blockId="5.[227,1436,406,503]" pageId="5" pageNumber="6">
<taxonomicName id="B4A74D32FFEE083B12B40FB53924FE63" authorityName="Lesne" authorityYear="1901" box="[227,304,406,432]" class="Insecta" family="Bostrichidae" genus="Xylion" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEE083B12B40FB53924FE63" box="[227,304,406,432]" italics="true" pageId="5" pageNumber="6">Xylion</emphasis>
</taxonomicName>
and
<taxonomicName id="B4A74D32FFEE083B13310FB539F7FE63" authorityName="Lesne" authorityYear="1901" box="[358,483,406,432]" class="Insecta" family="Bostrichidae" genus="Xylionulus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEE083B13310FB539F7FE63" box="[358,483,406,432]" italics="true" pageId="5" pageNumber="6">Xylionulus</emphasis>
</taxonomicName>
are Afrotropical genera,
<taxonomicName id="B4A74D32FFEE083B11540FB53B69FE63" authorityName="Lesne" authorityYear="1901" box="[771,893,406,432]" class="Insecta" family="Bostrichidae" genus="Xylobosca" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEE083B11540FB53B69FE63" box="[771,893,406,432]" italics="true" pageId="5" pageNumber="6">Xylobosca</emphasis>
</taxonomicName>
is an Australian genus, whilst
<taxonomicName id="B4A74D32FFEE083B168B0FB53D7FFE63" authorityName=", Lesne" authorityYear="1901" box="[1244,1387,406,432]" class="Insecta" family="Bostrichidae" genus="Octodesmus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="41D3EAA3FFEE083B168B0FB53D7FFE63" box="[1244,1387,406,432]" italics="true" pageId="5" pageNumber="6">Octodesmus</emphasis>
</taxonomicName>
and
<taxonomicName id="B4A74D32FFEE083B12B40F9A396AFE00" authorityName="Liu &amp; Beaver" authorityYear="2016" box="[227,382,441,467]" class="Insecta" family="Bostrichidae" genus="Octomeristes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="41D3EAA3FFEE083B12B40F9A396AFE00" box="[227,382,441,467]" italics="true" pageId="5" pageNumber="6">Octomeristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5AE057D8FFEE083B13D10F9A39E5FE00" box="[390,497,441,467]" pageId="5" pageNumber="6" rank="genus">gen. nov.</taxonomicNameLabel>
are Oriental genera, but the morphological similarities of the five genera might indicate a common origin in Gondwana.
</paragraph>
</subSubSection>
<subSubSection id="3BBD653AFFEE083B12B40C3E3A1CFDB5" box="[227,520,541,614]" pageId="5" pageNumber="6" type="distribution">
<paragraph id="731836B1FFEE083B12B40C3E3968FDE4" blockId="5.[227,380,541,567]" box="[227,380,541,567]" pageId="5" pageNumber="6">
<heading id="285081DDFFEE083B12B40C3E3968FDE4" bold="true" box="[227,380,541,567]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="41D3EAA3FFEE083B12B40C3E3968FDE4" bold="true" box="[227,380,541,567]" pageId="5" pageNumber="6">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="731836B1FFEE083B12B40C6F3A1CFDB5" blockId="5.[227,520,588,614]" box="[227,520,588,614]" pageId="5" pageNumber="6">
<collectingCountry id="0BB07621FFEE083B12B40C6F3936FDB5" box="[227,290,588,614]" name="India" pageId="5" pageNumber="6">India</collectingCountry>
, northern
<collectingCountry id="0BB07621FFEE083B13CF0C6F3A10FDB5" box="[408,516,588,614]" name="Thailand" pageId="5" pageNumber="6">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>