307 lines
30 KiB
XML
307 lines
30 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6231986" ID-GBIF-Dataset="80959f8f-aa73-499e-93ec-5c07458c7935" ID-GBIF-Taxon="160743188" ID-ISSN="0024-0672" ID-Zenodo-Dep="6231986" checkinTime="1576401033600" checkinUser="plazi" docAuthor="Schilthuizen, Menno & Perreau, Michel" docDate="2008" docId="F97A87E27404E66566FEFA15FC5AF99C" docLanguage="en" docName="ZM.82.19.189-210.pdf" docOrigin="Zool. Med. Leiden 82 (19)" docStyle="DocumentStyle{}" docTitle="Ptomaphaginus similipes Schilthuizen & Perreau 2008, spec. nov." docType="treatment" docVersion="5" lastPageId="5" lastPageNumber="194" masterDocId="0543FF9A7400E660674CFFA5FFD2FFE5" masterDocTitle="New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)" masterLastPageNumber="210" masterPageNumber="189" pageId="4" pageNumber="193" updateTime="1645572769617" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schilthuizen, Menno</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Perreau, Michel</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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||
<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zool. Med. Leiden</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2008</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2008-01-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>82</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>189</mods:start>
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<mods:end>210</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3576287</mods:identifier>
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<mods:identifier type="GBIF-Dataset">80959f8f-aa73-499e-93ec-5c07458c7935</mods:identifier>
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<mods:identifier type="ISSN">0024-0672</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3576287</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6231986" ID-GBIF-Taxon="160743188" ID-Zenodo-Dep="6231986" LSID="urn:lsid:plazi:treatment:F97A87E27404E66566FEFA15FC5AF99C" httpUri="http://treatment.plazi.org/id/F97A87E27404E66566FEFA15FC5AF99C" lastPageId="5" lastPageNumber="194" pageId="4" pageNumber="193">
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||
<subSubSection box="[434,837,1456,1480]" pageId="4" pageNumber="193" type="nomenclature">
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||
<paragraph blockId="4.[434,837,1456,1512]" box="[434,837,1456,1480]" pageId="4" pageNumber="193">
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<heading box="[434,837,1456,1480]" centered="true" fontSize="10" level="3" pageId="4" pageNumber="193" reason="1">
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<taxonomicName authority="Schilthuizen & Perreau, 2008" authorityName="Schilthuizen & Perreau" authorityYear="2008" box="[434,722,1456,1480]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="similipes" status="spec. nov.">
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<emphasis bold="true" box="[434,722,1456,1480]" italics="true" pageId="4" pageNumber="193">Ptomaphaginus similipes</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[728,837,1456,1479]" pageId="4" pageNumber="193" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[535,736,1488,1512]" pageId="4" pageNumber="193" type="description">
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||
<paragraph blockId="4.[434,837,1456,1512]" box="[535,736,1488,1512]" pageId="4" pageNumber="193">
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<heading box="[535,736,1488,1512]" centered="true" fontSize="10" level="3" pageId="4" pageNumber="193" reason="1">
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(
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<figureCitation box="[542,651,1488,1512]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="4" pageNumber="193">figs 18-19</figureCitation>
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;
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<figureCitation box="[661,725,1488,1512]" captionStart="Figs 26-30. 26" captionStartId="21.[151,190,1138,1157]" captionTargetBox="[151,1120,192,1089]" captionTargetPageId="21" captionText="Figs 26-30. 26, Ptomaphaginus caroli spec. nov., holotype; habitus, dorsal view. 27, Ptomaphaginus similipes spec. nov., female paratype (from Crocker Range); protarsus. 28, Ptomaphaginus similipes spec. nov., male paratype (from Trus Madi); protarsus. 29, Ptomaphaginus kinabaluensis spec. nov., female paratype (from Gunung Kinabalu); elytral apices. 30, Ptomaphaginus kinabaluensis spec. nov., holotype; elytral apices." figureDoi="http://doi.org/10.5281/zenodo.3576297" httpUri="https://zenodo.org/record/3576297/files/figure.png" pageId="4" pageNumber="193">27-28</figureCitation>
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).
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="5" lastPageNumber="194" pageId="4" pageNumber="193" type="materials_examined">
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<paragraph blockId="4.[151,1119,1550,1654]" lastBlockId="5.[151,1119,193,464]" lastPageId="5" lastPageNumber="194" pageId="4" pageNumber="193">
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Type material.—
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<materialsCitation ID-GBIF-Occurrence="2512737984" collectingDate="2000-10-31" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" latitude="5.78465" location="Crocker Range Park" longLatPrecision="1" longitude="116.34153" pageId="4" pageNumber="193" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
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||
<typeStatus box="[312,401,1551,1570]" pageId="4" pageNumber="193">Holotype</typeStatus>
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||
<specimenCount box="[407,425,1550,1570]" pageId="4" pageNumber="193" type="male">♂</specimenCount>
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||
:
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<collectingCountry box="[435,523,1551,1570]" name="Malaysia" pageId="4" pageNumber="193">Malaysia</collectingCountry>
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||
:
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<collectingRegion box="[531,590,1551,1570]" country="Malaysia" name="Sabah" pageId="4" pageNumber="193">Sabah</collectingRegion>
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||
:
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<location LSID="urn:lsid:plazi:treatment:F97A87E27404E66566FEFA15FC5AF99C:740C602F7404E664651AF9AAFCC6F9C7" box="[598,788,1551,1570]" country="Malaysia" latitude="5.78465" longLatPrecision="1" longitude="116.34153" name="Crocker Range Park" pageId="4" pageNumber="193" stateProvince="Sabah">Crocker Range Park</location>
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,
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<geoCoordinate box="[797,896,1551,1570]" degrees="5" direction="north" minutes="47.079" orientation="latitude" pageId="4" pageNumber="193" precision="1" value="5.78465">5º47.079’N</geoCoordinate>
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<geoCoordinate box="[902,1020,1551,1570]" degrees="116" direction="east" minutes="20.492" orientation="longitude" pageId="4" pageNumber="193" precision="1" value="116.34153">116º20.492’E</geoCoordinate>
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||
,
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||
<date box="[1028,1115,1551,1570]" pageId="4" pageNumber="193" value="2000-10-31">
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<collectingDate box="[1028,1115,1551,1570]" pageId="4" pageNumber="193" value="2000-10-31">31.x.2000</collectingDate>
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</date>
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,
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<collectingMethod box="[151,207,1579,1598]" pageId="4" pageNumber="193">pitfall</collectingMethod>
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with carrion or dung, leg.
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<collectorName box="[464,612,1579,1598]" pageId="4" pageNumber="193">M. Schilthuizen</collectorName>
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(
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<collectionCode box="[625,673,1579,1598]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="4" pageNumber="193" type="Herbarium">BOR</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="2512737994" collectingDate="2000-10-31" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" latitude="5.78465" location="Crocker Range Park" longLatPrecision="1" longitude="116.34153" pageId="4" pageNumber="193" specimenCount="10" specimenCount-male="3" stateProvince="Sabah" typeStatus="paratype">
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<typeStatus box="[689,783,1579,1598]" pageId="4" pageNumber="193">Paratypes</typeStatus>
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same collection data as
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<typeStatus box="[1013,1099,1579,1598]" pageId="4" pageNumber="193">holotype</typeStatus>
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:
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<specimenCount pageId="4" pageNumber="193" type="male">3 ♂♂</specimenCount>
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,
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<specimenCount box="[198,313,1607,1626]" pageId="4" pageNumber="193" type="generic">7 specimens</specimenCount>
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of undetermined sex (
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<collectionCode box="[528,576,1607,1626]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="4" pageNumber="193" type="Herbarium">BOR</collectionCode>
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)
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||
</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="2512737992" collectingDate="2000-07-31" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" elevation="1350" lastPageId="5" lastPageNumber="194" location="Gunung Mas" municipality="Crocker Range Park" pageId="4" pageNumber="193" specimenCount="2" stateProvince="Sabah" typeStatus="paratype">
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<collectingCountry box="[592,680,1607,1626]" name="Malaysia" pageId="4" pageNumber="193">Malaysia</collectingCountry>
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:
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<collectingRegion box="[689,748,1607,1626]" country="Malaysia" name="Sabah" pageId="4" pageNumber="193">Sabah</collectingRegion>
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||
:
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||
<collectingMunicipality box="[757,950,1607,1626]" pageId="4" pageNumber="193">Crocker Range Park</collectingMunicipality>
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||
:
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||
<location LSID="urn:lsid:plazi:treatment:F97A87E27404E66566FEFA15FC5AF99C:740C602F7404E66464F3F9E2FBEEF9BF" box="[959,1084,1607,1626]" country="Malaysia" municipality="Crocker Range Park" name="Gunung Mas" pageId="4" pageNumber="193" stateProvince="Sabah">Gunung Mas</location>
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||
(
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||
<locationDeviation location="Kota Kinabalu" pageId="4" pageNumber="193">
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||
<quantity metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="193" unit="km" value="30.0">30 km</quantity>
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||
ESE of Kota Kinabalu
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||
</locationDeviation>
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||
),
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||
<quantity box="[411,479,1635,1654]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="4" pageNumber="193" unit="m" value="1350.0">
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||
<elevation box="[411,479,1635,1654]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="4" pageNumber="193" unit="m" value="1350.0">1350 m</elevation>
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||
</quantity>
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||
alt.,
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||
<date box="[525,626,1635,1654]" pageId="4" pageNumber="193" value="2000-07-31">
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||
<collectingDate box="[525,626,1635,1654]" pageId="4" pageNumber="193" value="2000-07-31">31.vii.2000</collectingDate>
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||
</date>
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||
,
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||
<collectingMethod box="[636,692,1635,1654]" pageId="4" pageNumber="193">pitfall</collectingMethod>
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with carrion or dung, leg.
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<collectorName box="[948,1099,1635,1654]" pageId="4" pageNumber="193">M. Schilthuizen</collectorName>
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||
:
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<specimenCount lastPageId="5" lastPageNumber="194" pageId="4" pageNumber="193" type="generic">2 specimens</specimenCount>
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||
of unidentified sex (
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||
<collectionCode box="[441,489,193,212]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="5" pageNumber="194" type="Herbarium">BOR</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
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||
<materialsCitation ID-GBIF-Occurrence="2512737976" collectingDate="2001-09-16" collectingDateMax="2001-09-20" collectingDateMin="2001-09-16" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" elevation="200" location="Sugud Forest Reserve" pageId="5" pageNumber="194" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
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||
<collectingCountry box="[504,592,193,212]" name="Malaysia" pageId="5" pageNumber="194">Malaysia</collectingCountry>
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||
:
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||
<collectingRegion box="[599,658,193,212]" country="Malaysia" name="Sabah" pageId="5" pageNumber="194">Sabah</collectingRegion>
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||
:
|
||
<location LSID="urn:lsid:plazi:treatment:F97A87E27404E66566FEFA15FC5AF99C:740C602F7405E66565D6FF64FCBAFF31" box="[666,872,193,212]" country="Malaysia" name="Sugud Forest Reserve" pageId="5" pageNumber="194" stateProvince="Sabah">Sugud Forest Reserve</location>
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||
,
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||
<locationDeviation box="[881,1118,193,212]" location="Kota Kinabalu" pageId="5" pageNumber="194">
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||
<quantity box="[881,938,193,212]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="194" unit="km" value="15.0">15 km</quantity>
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S of Kota Kinabalu
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||
</locationDeviation>
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||
(
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<collectingRegion box="[158,218,221,240]" country="Malaysia" name="Sabah" pageId="5" pageNumber="194">Sabah</collectingRegion>
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||
),
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||
<quantity box="[233,289,221,240]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="194" unit="m" value="200.0">
|
||
<elevation box="[233,289,221,240]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="194" unit="m" value="200.0">200 m</elevation>
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||
</quantity>
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||
alt.,
|
||
<date box="[333,455,221,240]" pageId="5" pageNumber="194" value="2001-09-16" valueMax="2001-09-20" valueMin="2001-09-16">
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||
<collectingDate box="[333,455,221,240]" pageId="5" pageNumber="194" value="2001-09-16" valueMax="2001-09-20" valueMin="2001-09-16">16-20.ix.2001</collectingDate>
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||
</date>
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||
,
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||
<collectingMethod box="[464,520,221,240]" pageId="5" pageNumber="194">pitfall</collectingMethod>
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||
with carrion, leg.
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||
<collectorName box="[690,839,221,240]" pageId="5" pageNumber="194">M. Schilthuizen</collectorName>
|
||
,
|
||
<specimenCount box="[848,882,220,240]" pageId="5" pageNumber="194" type="male">1 ♂</specimenCount>
|
||
, 4 individuals of unidentified sex (
|
||
<collectionCode box="[246,294,249,268]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="5" pageNumber="194" type="Herbarium">BOR</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2512737982" collectingDate="2001-07-02" collectingDateMax="2001-07-05" collectingDateMin="2001-07-02" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" elevation="1017" location="Mahua Waterfall" pageId="5" pageNumber="194" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
|
||
<collectingCountry box="[308,396,249,268]" name="Malaysia" pageId="5" pageNumber="194">Malaysia</collectingCountry>
|
||
:
|
||
<collectingRegion box="[403,462,249,268]" country="Malaysia" name="Sabah" pageId="5" pageNumber="194">Sabah</collectingRegion>
|
||
:
|
||
<location LSID="urn:lsid:plazi:treatment:F97A87E27404E66566FEFA15FC5AF99C:740C602F7405E665669AFF5CFDA1FEE9" box="[470,627,249,268]" country="Malaysia" name="Mahua Waterfall" pageId="5" pageNumber="194" stateProvince="Sabah">Mahua Waterfall</location>
|
||
(5˚47’52.3”N 116˚24’19.0”E),
|
||
<quantity box="[897,964,249,268]" metricMagnitude="3" metricUnit="m" metricValue="1.017" pageId="5" pageNumber="194" unit="m" value="1017.0">
|
||
<elevation box="[897,964,249,268]" metricMagnitude="3" metricUnit="m" metricValue="1.017" pageId="5" pageNumber="194" unit="m" value="1017.0">1017 m</elevation>
|
||
</quantity>
|
||
alt.,
|
||
<date box="[1007,1115,249,268]" pageId="5" pageNumber="194" value="2001-07-02" valueMax="2001-07-05" valueMin="2001-07-02">
|
||
<collectingDate box="[1007,1115,249,268]" pageId="5" pageNumber="194" value="2001-07-02" valueMax="2001-07-05" valueMin="2001-07-02">2-5.vii.2001</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod box="[151,207,277,296]" pageId="5" pageNumber="194">pitfall</collectingMethod>
|
||
with carrion or dung, leg.
|
||
<collectorName box="[484,638,277,296]" pageId="5" pageNumber="194">M. Schilthuizen</collectorName>
|
||
,
|
||
<specimenCount box="[652,690,276,296]" pageId="5" pageNumber="194" type="male">1 ♂</specimenCount>
|
||
(
|
||
<collectionCode box="[705,753,277,296]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="5" pageNumber="194" type="Herbarium">BOR</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation ID-GBIF-Occurrence="2512737986" collectingDate="2001-10-23" collectingDateMax="2001-10-28" collectingDateMin="2001-10-23" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" elevation="1400" location="Mount Trus Madi" pageId="5" pageNumber="194" specimenCount="12" specimenCount-female="2" specimenCount-male="3" stateProvince="Sabah" typeStatus="paratype">
|
||
<collectingCountry box="[772,860,277,296]" name="Malaysia" pageId="5" pageNumber="194">Malaysia</collectingCountry>
|
||
:
|
||
<collectingRegion box="[873,932,277,296]" country="Malaysia" name="Sabah" pageId="5" pageNumber="194">Sabah</collectingRegion>
|
||
:
|
||
<location LSID="urn:lsid:plazi:treatment:F97A87E27404E66566FEFA15FC5AF99C:740C602F7405E66564FCFEB0FB8DFECD" box="[944,1119,277,296]" country="Malaysia" name="Mount Trus Madi" pageId="5" pageNumber="194" stateProvince="Sabah">Mount Trus Madi</location>
|
||
(5˚34.152’N 116˚29.638’E),
|
||
<quantity box="[397,464,305,324]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="194" unit="m" value="1400.0">
|
||
<elevation box="[397,464,305,324]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="194" unit="m" value="1400.0">1400 m</elevation>
|
||
</quantity>
|
||
alt.,
|
||
<collectingMethod box="[509,565,305,324]" pageId="5" pageNumber="194">pitfall</collectingMethod>
|
||
with lamb,
|
||
<date box="[677,793,305,324]" pageId="5" pageNumber="194" value="2001-10-23" valueMax="2001-10-28" valueMin="2001-10-23">
|
||
<collectingDate box="[677,793,305,324]" pageId="5" pageNumber="194" value="2001-10-23" valueMax="2001-10-28" valueMin="2001-10-23">23-28.x.2001</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName box="[841,991,305,324]" pageId="5" pageNumber="194">M. Schilthuizen</collectorName>
|
||
,
|
||
<specimenCount box="[1000,1052,304,324]" pageId="5" pageNumber="194" type="male">3 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount box="[1062,1114,304,324]" pageId="5" pageNumber="194" type="female">2 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount box="[151,265,333,352]" pageId="5" pageNumber="194" type="generic">7 specimens</specimenCount>
|
||
of unidentified sex in 70% ethanol (
|
||
<collectionCode box="[603,651,333,352]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="5" pageNumber="194" type="Herbarium">BOR</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1119,193,464]" pageId="5" pageNumber="194">
|
||
<typeStatus box="[151,337,361,380]" isOtherTypeStatus="true" pageId="5" pageNumber="194">Additional material</typeStatus>
|
||
(not included in the type series).— Type locality,
|
||
<specimenCount box="[800,914,361,380]" pageId="5" pageNumber="194" type="generic">4 specimens</specimenCount>
|
||
of unidentified sex in 70% ethanol,
|
||
<specimenCount box="[278,393,389,408]" pageId="5" pageNumber="194" type="generic">2 specimens</specimenCount>
|
||
of unidentified sex in 100% ethanol; Sugud,
|
||
<specimenCount box="[818,865,388,408]" pageId="5" pageNumber="194" type="male">7♂♂</specimenCount>
|
||
in 100% ethanol; Kinabalu Park Headquarters,
|
||
<date box="[343,450,417,436]" pageId="5" pageNumber="194" value="2003-07-05" valueMax="2003-07-08" valueMin="2003-07-05">5-8.vii.2003</date>
|
||
, in pitfall with pork,
|
||
<specimenCount box="[653,687,416,436]" pageId="5" pageNumber="194" type="male">1 ♂</specimenCount>
|
||
(DNA isolate). (For full collection details, see Type material above.)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="194" type="diagnosis">
|
||
<paragraph blockId="5.[151,1119,504,1657]" pageId="5" pageNumber="194">
|
||
Diagnosis.— Habitus relatively slender, ovoid. Elytra on average 1.2 times as long as their combined width (length measured from the caudal tip of the scutellum to the elytral apices). Aedeagus (
|
||
<figureCitation box="[449,557,567,591]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="5" pageNumber="194">figs 18-19</figureCitation>
|
||
) with two short apical lateral ‘wings’ and a short terminal processus. Male forelegs usually with long hairs on the ventral side of the profemur and protibia. Male protarsus completely undilated, of the same width as in the female (
|
||
<figureCitation box="[281,392,661,685]" captionStart="Figs 26-30. 26" captionStartId="21.[151,190,1138,1157]" captionTargetBox="[151,1120,192,1089]" captionTargetPageId="21" captionText="Figs 26-30. 26, Ptomaphaginus caroli spec. nov., holotype; habitus, dorsal view. 27, Ptomaphaginus similipes spec. nov., female paratype (from Crocker Range); protarsus. 28, Ptomaphaginus similipes spec. nov., male paratype (from Trus Madi); protarsus. 29, Ptomaphaginus kinabaluensis spec. nov., female paratype (from Gunung Kinabalu); elytral apices. 30, Ptomaphaginus kinabaluensis spec. nov., holotype; elytral apices." figureDoi="http://doi.org/10.5281/zenodo.3576297" httpUri="https://zenodo.org/record/3576297/files/figure.png" pageId="5" pageNumber="194">figs 27-28</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="194" type="description">
|
||
<paragraph blockId="5.[151,1119,504,1657]" pageId="5" pageNumber="194">
|
||
Description.— Habitus relatively slender, ovoid. Moderately pigmented: mostly chestnut brown; only the legs, the elytral apices, the tips and the bases of the antennae lighter brown. Length 2.4-3.0 mm (n = 18). Antennae slender and relatively long, 1.1-1.2 times as long as the width of the head (n = 2). Articles 6, 9, and 10 slightly longer than wide. Male and female protarsi completely undilated (
|
||
<figureCitation box="[784,898,817,841]" captionStart="Figs 26-30. 26" captionStartId="21.[151,190,1138,1157]" captionTargetBox="[151,1120,192,1089]" captionTargetPageId="21" captionText="Figs 26-30. 26, Ptomaphaginus caroli spec. nov., holotype; habitus, dorsal view. 27, Ptomaphaginus similipes spec. nov., female paratype (from Crocker Range); protarsus. 28, Ptomaphaginus similipes spec. nov., male paratype (from Trus Madi); protarsus. 29, Ptomaphaginus kinabaluensis spec. nov., female paratype (from Gunung Kinabalu); elytral apices. 30, Ptomaphaginus kinabaluensis spec. nov., holotype; elytral apices." figureDoi="http://doi.org/10.5281/zenodo.3576297" httpUri="https://zenodo.org/record/3576297/files/figure.png" pageId="5" pageNumber="194">figs 27-28</figureCitation>
|
||
): the protibia at its largest width (excluding the lateral spines) is 2.5 times broader than the first article of the protarsus (n = 1). In the male, the ventral side of profemur and protibia carry long hairs, some of which are as long as the tarsus. (In one
|
||
<typeStatus box="[761,861,912,936]" pageId="5" pageNumber="194">paratype</typeStatus>
|
||
specimen from Mount Trus Madi, these hairs were absent.) Thorax 1.69-1.80 times as wide as long (n = 3), the caudal corners distinctly drawn out. The mesosternal carina is strongly developed, its edge somewhat thickened. Elytra of medium length, 1.19-1.23 times as long (measured from the caudal tip of the scutellum) as their combined width. Elytral apices distinctly truncate, identical in both sexes. The 3
|
||
<superScript attach="none" box="[581,595,1067,1081]" fontSize="6" pageId="5" pageNumber="194">rd-</superScript>
|
||
5
|
||
<superScript attach="left" box="[617,630,1067,1081]" fontSize="6" pageId="5" pageNumber="194">th</superScript>
|
||
visible abdominal sternite in the male each have a broad and relatively distinct central notch and are depressed in the vicinity of these notches. Aedeagus (
|
||
<figureCitation box="[447,557,1131,1155]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="5" pageNumber="194">figs 18-19</figureCitation>
|
||
) in dorsal view abruptly narrowing towards the apex, and there adorned with two short lateral ‘wings’. The tip is composed of a short, obtuse, triangular processus. In lateral view, the aedeagus is almost not curved, except for the apical quarter, which is bent ventrad under an obtuse angle. Male genital segment relatively broad, at the apex slightly triangular, 2.5 times as long as wide.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="194" type="distribution">
|
||
<paragraph blockId="5.[151,1119,504,1657]" pageId="5" pageNumber="194">
|
||
Distribution.— This species is distributed in the Crocker and Trus Madi mountain ranges of western
|
||
<collectingRegion box="[355,425,1319,1343]" country="Malaysia" name="Sabah" pageId="5" pageNumber="194">Sabah</collectingRegion>
|
||
, where it was found in lower montane forest, but also (Sugud) in lowland forest in the foot hills.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="194" type="discussion">
|
||
<paragraph blockId="5.[151,1119,504,1657]" pageId="5" pageNumber="194">
|
||
Remarks.— Based on aedeagal shape,
|
||
<taxonomicName authorityName="Schilthuizen & Perreau" authorityYear="2008" box="[624,740,1381,1405]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="similipes">
|
||
<emphasis box="[624,740,1381,1405]" italics="true" pageId="5" pageNumber="194">P. similipes</emphasis>
|
||
</taxonomicName>
|
||
is closely related to
|
||
<taxonomicName authorityName="Schilthuizen & Perreau" authorityYear="2008" box="[969,1051,1381,1405]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="caroli">
|
||
<emphasis box="[969,1051,1381,1405]" italics="true" pageId="5" pageNumber="194">P. caroli</emphasis>
|
||
</taxonomicName>
|
||
, from which it differs strongly in the habitus, which is very slender in
|
||
<taxonomicName authorityName="Schilthuizen & Perreau" authorityYear="2008" box="[868,949,1413,1437]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="caroli">
|
||
<emphasis box="[868,949,1413,1437]" italics="true" pageId="5" pageNumber="194">P. caroli</emphasis>
|
||
</taxonomicName>
|
||
. It may also be related to
|
||
<taxonomicName authorityName="Szymczakowski" authorityYear="1974" box="[267,394,1445,1468]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="coronatus">
|
||
<emphasis box="[267,394,1445,1468]" italics="true" pageId="5" pageNumber="194">P. coronatus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion box="[465,564,1444,1468]" country="Philippines" name="Palawan" pageId="5" pageNumber="194">Palawan</collectingRegion>
|
||
, which has a similar habitus. The undilated protarsi make females and males difficult to distinguish externally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="5" pageNumber="194" type="etymology">
|
||
<paragraph blockId="5.[151,1119,504,1657]" pageId="5" pageNumber="194">
|
||
Etymology.—
|
||
<taxonomicName authorityName="Schilthuizen & Perreau" authorityYear="2008" box="[354,444,1507,1531]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="194" phylum="Arthropoda" rank="species" species="similipes">
|
||
<emphasis box="[354,444,1507,1531]" italics="true" pageId="5" pageNumber="194">similipes</emphasis>
|
||
</taxonomicName>
|
||
, a Latin composite noun in apposition, meaning ‘identical feet’. The name refers to the fact that in this species males and females both have undilated tarsi. The name is based on a suggestion by Jan-Willem van Velzen, a visitor to an exhibition on Linnaean taxonomy in the National Museum of Natural History ‘Naturalis’, in which the naming of this new species was an exhibit feature.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |