167 lines
16 KiB
XML
167 lines
16 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.6231984" ID-GBIF-Dataset="80959f8f-aa73-499e-93ec-5c07458c7935" ID-GBIF-Taxon="160743189" ID-ISSN="0024-0672" ID-Zenodo-Dep="6231984" checkinTime="1576401033600" checkinUser="plazi" docAuthor="Schilthuizen, Menno & Perreau, Michel" docDate="2008" docId="F97A87E27404E6646688FEC5FC49FA6D" docLanguage="en" docName="ZM.82.19.189-210.pdf" docOrigin="Zool. Med. Leiden 82 (19)" docStyle="DocumentStyle{}" docTitle="Ptomaphaginus caroli Schilthuizen & Perreau 2008, spec. nov." docType="treatment" docVersion="5" lastPageNumber="193" masterDocId="0543FF9A7400E660674CFFA5FFD2FFE5" masterDocTitle="New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)" masterLastPageNumber="210" masterPageNumber="189" pageId="4" pageNumber="193" updateTime="1645572769617" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Schilthuizen, Menno</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Perreau, Michel</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zool. Med. Leiden</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2008</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2008-01-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>82</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>19</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>189</mods:start>
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<mods:end>210</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3576287</mods:identifier>
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<mods:identifier type="GBIF-Dataset">80959f8f-aa73-499e-93ec-5c07458c7935</mods:identifier>
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<mods:identifier type="ISSN">0024-0672</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3576287</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6231984" ID-GBIF-Taxon="160743189" ID-Zenodo-Dep="6231984" LSID="urn:lsid:plazi:treatment:F97A87E27404E6646688FEC5FC49FA6D" httpUri="http://treatment.plazi.org/id/F97A87E27404E6646688FEC5FC49FA6D" lastPageNumber="193" pageId="4" pageNumber="193">
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<subSubSection box="[452,818,352,376]" pageId="4" pageNumber="193" type="nomenclature">
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<paragraph blockId="4.[452,818,352,408]" box="[452,818,352,376]" pageId="4" pageNumber="193">
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<heading box="[452,818,352,376]" centered="true" fontSize="10" level="3" pageId="4" pageNumber="193" reason="1">
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<taxonomicName authority="Schilthuizen & Perreau, 2008" authorityName="Schilthuizen & Perreau" authorityYear="2008" box="[452,703,352,376]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="caroli" status="spec. nov.">
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<emphasis bold="true" box="[452,703,352,376]" italics="true" pageId="4" pageNumber="193">Ptomaphaginus caroli</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[709,818,353,376]" pageId="4" pageNumber="193" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[552,719,384,408]" pageId="4" pageNumber="193" type="description">
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<paragraph blockId="4.[452,818,352,408]" box="[552,719,384,408]" pageId="4" pageNumber="193">
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<heading box="[552,719,384,408]" centered="true" fontSize="10" level="3" pageId="4" pageNumber="193" reason="1">
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(
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<figureCitation box="[559,667,384,408]" captionStart="Figs 13-15. 13" captionStartId="19.[151,189,769,788]" captionTargetBox="[153,1083,201,739]" captionTargetId="figure@19.[459,777,172,750]" captionTargetPageId="19" captionText="Figs 13-15. 13, Ptomaphaginus bryanti Jeannel, aedeagus; reproduced from Jeannel (1936). 14, Ptomaphaginus caroli spec. nov., holotype; aedeagus dorsal view. 15, Ptomaphaginus caroli spec. nov., holotype; aedeagus lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576293" httpUri="https://zenodo.org/record/3576293/files/figure.png" pageId="4" pageNumber="193">figs 14-15</figureCitation>
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;
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<figureCitation box="[678,707,384,408]" captionStart="Figs 26-30. 26" captionStartId="21.[151,190,1138,1157]" captionTargetBox="[151,1120,192,1089]" captionTargetPageId="21" captionText="Figs 26-30. 26, Ptomaphaginus caroli spec. nov., holotype; habitus, dorsal view. 27, Ptomaphaginus similipes spec. nov., female paratype (from Crocker Range); protarsus. 28, Ptomaphaginus similipes spec. nov., male paratype (from Trus Madi); protarsus. 29, Ptomaphaginus kinabaluensis spec. nov., female paratype (from Gunung Kinabalu); elytral apices. 30, Ptomaphaginus kinabaluensis spec. nov., holotype; elytral apices." figureDoi="http://doi.org/10.5281/zenodo.3576297" httpUri="https://zenodo.org/record/3576297/files/figure.png" pageId="4" pageNumber="193">26</figureCitation>
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).
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="193" type="materials_examined">
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<paragraph blockId="4.[151,1119,447,495]" pageId="4" pageNumber="193">
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Type material.—
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<materialsCitation ID-GBIF-Occurrence="2512737959" collectingDate="2000-07-31" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" elevation="1350" location="Gunung Mas" municipality="Crocker Range Park" pageId="4" pageNumber="193" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
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<typeStatus box="[314,403,448,467]" pageId="4" pageNumber="193">Holotype</typeStatus>
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<specimenCount box="[409,427,447,467]" pageId="4" pageNumber="193" type="male">♂</specimenCount>
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:
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<collectingCountry box="[438,526,448,467]" name="Malaysia" pageId="4" pageNumber="193">Malaysia</collectingCountry>
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:
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<collectingRegion box="[535,594,448,467]" country="Malaysia" name="Sabah" pageId="4" pageNumber="193">Sabah</collectingRegion>
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:
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<collectingMunicipality box="[603,795,448,467]" pageId="4" pageNumber="193">Crocker Range Park</collectingMunicipality>
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:
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<location LSID="urn:lsid:plazi:treatment:F97A87E27404E6646688FEC5FC49FA6D:740C602F7404E6646468FE65FC73FE36" box="[804,929,448,467]" country="Malaysia" municipality="Crocker Range Park" name="Gunung Mas" pageId="4" pageNumber="193" stateProvince="Sabah">Gunung Mas</location>
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(
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<locationDeviation location="Kota Kinabalu" pageId="4" pageNumber="193">
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<quantity box="[941,1000,448,467]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="193" unit="km" value="30.0">30 km</quantity>
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ESE of Kota Kinabalu
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</locationDeviation>
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),
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<quantity box="[255,322,476,495]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="4" pageNumber="193" unit="m" value="1350.0">
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<elevation box="[255,322,476,495]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="4" pageNumber="193" unit="m" value="1350.0">1350 m</elevation>
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</quantity>
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alt.,
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<collectingMethod box="[367,423,476,495]" pageId="4" pageNumber="193">pitfall</collectingMethod>
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with carrion or dung,
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<date box="[636,737,476,495]" pageId="4" pageNumber="193" value="2000-07-31">
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<collectingDate box="[636,737,476,495]" pageId="4" pageNumber="193" value="2000-07-31">31.vii.2000</collectingDate>
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</date>
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, leg.
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<collectorName box="[785,933,476,495]" pageId="4" pageNumber="193">M. Schilthuizen</collectorName>
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(
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<collectionCode box="[945,993,476,495]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="4" pageNumber="193" type="Herbarium">BOR</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="193" type="diagnosis">
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<paragraph blockId="4.[151,1119,536,1417]" pageId="4" pageNumber="193">
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Diagnosis.— Habitus long and narrow, flat (
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<figureCitation box="[679,749,536,560]" captionStart="Figs 26-30. 26" captionStartId="21.[151,190,1138,1157]" captionTargetBox="[151,1120,192,1089]" captionTargetPageId="21" captionText="Figs 26-30. 26, Ptomaphaginus caroli spec. nov., holotype; habitus, dorsal view. 27, Ptomaphaginus similipes spec. nov., female paratype (from Crocker Range); protarsus. 28, Ptomaphaginus similipes spec. nov., male paratype (from Trus Madi); protarsus. 29, Ptomaphaginus kinabaluensis spec. nov., female paratype (from Gunung Kinabalu); elytral apices. 30, Ptomaphaginus kinabaluensis spec. nov., holotype; elytral apices." figureDoi="http://doi.org/10.5281/zenodo.3576297" httpUri="https://zenodo.org/record/3576297/files/figure.png" pageId="4" pageNumber="193">fig. 26</figureCitation>
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). Elytra 1.41 times as long as their combined width (length measured from the caudal tip of the scutellum to the apex of the elytra). Aedeagus (
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<figureCitation box="[406,513,599,623]" captionStart="Figs 13-15. 13" captionStartId="19.[151,189,769,788]" captionTargetBox="[153,1083,201,739]" captionTargetId="figure@19.[459,777,172,750]" captionTargetPageId="19" captionText="Figs 13-15. 13, Ptomaphaginus bryanti Jeannel, aedeagus; reproduced from Jeannel (1936). 14, Ptomaphaginus caroli spec. nov., holotype; aedeagus dorsal view. 15, Ptomaphaginus caroli spec. nov., holotype; aedeagus lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576293" httpUri="https://zenodo.org/record/3576293/files/figure.png" pageId="4" pageNumber="193">figs 14-15</figureCitation>
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) apically with two short ‘wings’ and a very small, indistinct terminal processus.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="193" type="description">
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<paragraph blockId="4.[151,1119,536,1417]" pageId="4" pageNumber="193">
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Description.— Habitus narrow and elongated, relatively flat (
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<figureCitation box="[895,969,663,687]" captionStart="Figs 26-30. 26" captionStartId="21.[151,190,1138,1157]" captionTargetBox="[151,1120,192,1089]" captionTargetPageId="21" captionText="Figs 26-30. 26, Ptomaphaginus caroli spec. nov., holotype; habitus, dorsal view. 27, Ptomaphaginus similipes spec. nov., female paratype (from Crocker Range); protarsus. 28, Ptomaphaginus similipes spec. nov., male paratype (from Trus Madi); protarsus. 29, Ptomaphaginus kinabaluensis spec. nov., female paratype (from Gunung Kinabalu); elytral apices. 30, Ptomaphaginus kinabaluensis spec. nov., holotype; elytral apices." figureDoi="http://doi.org/10.5281/zenodo.3576297" httpUri="https://zenodo.org/record/3576297/files/figure.png" pageId="4" pageNumber="193">fig. 26</figureCitation>
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). Moderately pigmented: mostly chestnut brown; only the legs and the antennal bases and apices lighter yellowish brown. Length
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<quantity box="[511,595,726,750]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="193" unit="mm" value="2.7">2.7 mm</quantity>
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. Antennae slender and relatively long, 1.1 times as long as the width of the head. Articles 6, 9, and 10 slightly wider than long. Male protarsi strongly dilated: the protibia at its largest width (excluding the lateral spines) is 1.2 times broader than the first article of the protarsus. There are no exceptionally long hairs on the ventral side of profemur and protibia. Thorax 1.62 times as wide as long, relatively flat, almost rectilinearly narrowing from caudal to rostral; the caudal corners sharp, narrowly, but distinctly drawn out. The mesosternal carina is strongly developed, its edge somewhat thickened. Elytra long, 1.41 times as long (measured from the caudal tip of the scutellum) as their combined width. Elytral apices distinctly truncate. Aedeagus (
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<figureCitation box="[204,310,1012,1036]" captionStart="Figs 13-15. 13" captionStartId="19.[151,189,769,788]" captionTargetBox="[153,1083,201,739]" captionTargetId="figure@19.[459,777,172,750]" captionTargetPageId="19" captionText="Figs 13-15. 13, Ptomaphaginus bryanti Jeannel, aedeagus; reproduced from Jeannel (1936). 14, Ptomaphaginus caroli spec. nov., holotype; aedeagus dorsal view. 15, Ptomaphaginus caroli spec. nov., holotype; aedeagus lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576293" httpUri="https://zenodo.org/record/3576293/files/figure.png" pageId="4" pageNumber="193">figs 14-15</figureCitation>
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) in dorsal view abruptly narrowing towards the apex, and there adorned with two very short lateral ‘wings’. The tip carries a short and obtuse triangular processus. In lateral view, the aedeagus is almost not curved, except for the apical quarter, which is bent ventrad under an obtuse angle. Male genital segment relatively broad, at the apex slightly triangular, 2.5 times as long as wide. Female unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="193" type="distribution">
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<paragraph blockId="4.[151,1119,536,1417]" pageId="4" pageNumber="193">
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Distribution.— Only the
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<typeStatus box="[475,574,1170,1194]" pageId="4" pageNumber="193">holotype</typeStatus>
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is known for this species. At the type locality, it was found syntopically with
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<taxonomicName authority="Schilthuizen & Perreau, 2008" authorityName="Schilthuizen & Perreau" authorityYear="2008" box="[477,592,1202,1226]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="similipes" status="spec. nov.">
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<emphasis box="[477,592,1202,1226]" italics="true" pageId="4" pageNumber="193">P. similipes</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[599,708,1203,1226]" pageId="4" pageNumber="193" rank="species">spec. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="193" type="discussion">
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<paragraph blockId="4.[151,1119,536,1417]" pageId="4" pageNumber="193">
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Remarks.— Based on aedeagal shape,
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<taxonomicName authorityName="Schilthuizen & Perreau" authorityYear="2008" box="[607,685,1234,1258]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="caroli">
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<emphasis box="[607,685,1234,1258]" italics="true" pageId="4" pageNumber="193">P. caroli</emphasis>
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</taxonomicName>
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is closely related to
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<taxonomicName authorityName="Jeannel" authorityYear="1936" box="[900,996,1234,1258]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="bryanti">
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<emphasis box="[900,996,1234,1258]" italics="true" pageId="4" pageNumber="193">P. bryanti</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Schilthuizen & Perreau" authorityYear="2008" box="[1008,1119,1234,1258]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="similipes">
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<emphasis box="[1008,1119,1234,1258]" italics="true" pageId="4" pageNumber="193">P. similipes</emphasis>
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</taxonomicName>
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and
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<taxonomicName box="[201,343,1266,1290]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="bryantoides">
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<emphasis box="[201,343,1266,1290]" italics="true" pageId="4" pageNumber="193">P. bryantoides</emphasis>
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</taxonomicName>
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. However, it differs from all these by the distinctly elongated habitus (elytral index of 1.41) and the apical ‘wings’ of the aedeagus, which are very short.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="193" type="etymology">
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<paragraph blockId="4.[151,1119,536,1417]" pageId="4" pageNumber="193">
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Etymology.— Named in honour of Carolus Linnaeus. The present paper appears in conjunction with the 250
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<superScript attach="left" box="[424,437,1360,1374]" fontSize="6" pageId="4" pageNumber="193">th</superScript>
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anniversary of the publication of Linnaeus’ Systema Naturae on
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<date box="[186,352,1392,1417]" pageId="4" pageNumber="193" value="1758-01-01">1 January 1758</date>
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, which marks the start of zoological nomenclature.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |