treatments-xml/data/F9/7A/87/F97A87E27403E66466D1FF65FC98FEDD.xml
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<document ID-DOI="http://doi.org/10.5281/zenodo.6231982" ID-GBIF-Dataset="80959f8f-aa73-499e-93ec-5c07458c7935" ID-GBIF-Taxon="160743190" ID-ISSN="0024-0672" ID-Zenodo-Dep="6231982" checkinTime="1576401033600" checkinUser="plazi" docAuthor="Schilthuizen, Menno &amp; Perreau, Michel" docDate="2008" docId="F97A87E27403E66466D1FF65FC98FEDD" docLanguage="en" docName="ZM.82.19.189-210.pdf" docOrigin="Zool. Med. Leiden 82 (19)" docStyle="DocumentStyle{}" docTitle="Ptomaphaginus bryantioides Schilthuizen &amp; Perreau 2008, spec. nov." docType="treatment" docVersion="5" lastPageId="4" lastPageNumber="193" masterDocId="0543FF9A7400E660674CFFA5FFD2FFE5" masterDocTitle="New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)" masterLastPageNumber="210" masterPageNumber="189" pageId="3" pageNumber="192" updateTime="1645572769617" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New species and new records of Ptomaphaginus Portevin from the Sunda region, Southeast Asia (Coleoptera: Leiodidae: Cholevinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Schilthuizen, Menno</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Perreau, Michel</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zool. Med. Leiden</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2008</mods:date>
<mods:detail type="pubDate">
<mods:number>2008-01-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>82</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>19</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>189</mods:start>
<mods:end>210</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3576287</mods:identifier>
<mods:identifier type="GBIF-Dataset">80959f8f-aa73-499e-93ec-5c07458c7935</mods:identifier>
<mods:identifier type="ISSN">0024-0672</mods:identifier>
<mods:identifier type="Zenodo-Dep">3576287</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6231982" ID-GBIF-Taxon="160743190" ID-Zenodo-Dep="6231982" LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD" httpUri="http://treatment.plazi.org/id/F97A87E27403E66466D1FF65FC98FEDD" lastPageId="4" lastPageNumber="193" pageId="3" pageNumber="192">
<subSubSection box="[413,857,192,216]" pageId="3" pageNumber="192" type="nomenclature">
<paragraph blockId="3.[413,857,192,248]" box="[413,857,192,216]" pageId="3" pageNumber="192">
<heading box="[413,857,192,216]" centered="true" fontSize="10" level="3" pageId="3" pageNumber="192" reason="1">
<taxonomicName authority="Schilthuizen &amp; Perreau, 2008" authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[413,742,192,216]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="bryantioides" status="spec. nov.">
<emphasis bold="true" box="[413,742,192,216]" italics="true" pageId="3" pageNumber="192">Ptomaphaginus bryantioides</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[748,857,193,216]" pageId="3" pageNumber="192" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[571,700,224,248]" pageId="3" pageNumber="192" type="description">
<paragraph blockId="3.[413,857,192,248]" box="[571,700,224,248]" pageId="3" pageNumber="192">
<heading box="[571,700,224,248]" centered="true" fontSize="10" level="3" pageId="3" pageNumber="192" reason="1">
(
<figureCitation box="[578,688,224,248]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="3" pageNumber="192">figs 20-21</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="192" type="materials_examined">
<paragraph blockId="3.[151,1120,287,727]" pageId="3" pageNumber="192">
Type material.—
<materialsCitation ID-GBIF-Occurrence="2512737995" collectingDate="2000-05-15" collectionCode="BOR" collectorName="J. de Roode" country="Malaysia" county="Danum Valley Field Centre" latitude="5.0" location="Valley Field Centre" longLatPrecision="1305" longitude="117.833336" municipality="Danum Valley" pageId="3" pageNumber="192" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus box="[319,408,288,307]" pageId="3" pageNumber="192">Holotype</typeStatus>
<specimenCount box="[417,435,287,307]" pageId="3" pageNumber="192" type="male"></specimenCount>
:
<collectingCountry box="[448,536,288,307]" name="Malaysia" pageId="3" pageNumber="192">Malaysia</collectingCountry>
:
<collectingRegion box="[548,607,288,307]" country="Malaysia" name="Sabah" pageId="3" pageNumber="192">Sabah</collectingRegion>
:
<collectingCounty box="[618,885,288,307]" pageId="3" pageNumber="192">
<collectingMunicipality box="[618,757,288,307]" pageId="3" pageNumber="192">
Danum
<location LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD:740C602F7403E66365F6FE85FD27FED6" box="[698,757,288,307]" country="Malaysia" county="Danum Valley Field Centre" latitude="5.0" longLatPrecision="1305" longitude="117.833336" municipality="Danum Valley" name="Valley Field Centre" pageId="3" pageNumber="192" stateProvince="Sabah">Valley</location>
</collectingMunicipality>
Field Centre
</collectingCounty>
(
<locationDeviation location="Lahad Datu" pageId="3" pageNumber="192">
<quantity box="[899,961,288,307]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="192" unit="km" value="60.0">60 km</quantity>
WSW of Lahad Datu
</locationDeviation>
),
<geoCoordinate box="[217,285,316,335]" degrees="5" direction="north" minutes="00" orientation="latitude" pageId="3" pageNumber="192" precision="925" value="5.0">5 º00N</geoCoordinate>
<geoCoordinate box="[292,372,316,335]" degrees="117" direction="east" minutes="50" orientation="longitude" pageId="3" pageNumber="192" precision="925" value="117.833336">117º50E</geoCoordinate>
,
<date box="[382,468,316,335]" pageId="3" pageNumber="192" value="2000-05-15">
<collectingDate box="[382,468,316,335]" pageId="3" pageNumber="192" value="2000-05-15">15.v.2000</collectingDate>
</date>
,
<collectingMethod box="[478,534,316,335]" pageId="3" pageNumber="192">pitfall</collectingMethod>
with chicken, leg.
<collectorName box="[713,822,316,335]" pageId="3" pageNumber="192">J. de Roode</collectorName>
(
<collectionCode box="[835,883,316,335]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
.
<materialsCitation ID-GBIF-Occurrence="2512737998" collectingDate="2000-05-15" collectionCode="BOR" collectorName="J. de Roode" country="Malaysia" county="Danum Valley Field Centre" latitude="5.0" location="Valley Field Centre" longLatPrecision="1305" longitude="117.833336" municipality="Danum Valley" pageId="3" pageNumber="192" specimenCount="11" specimenCount-female="9" specimenCount-male="2" stateProvince="Sabah" typeStatus="paratype">
<typeStatus box="[898,993,316,335]" pageId="3" pageNumber="192">Paratypes</typeStatus>
, same collection data as
<typeStatus box="[264,350,344,363]" pageId="3" pageNumber="192">holotype</typeStatus>
:
<specimenCount box="[358,409,343,363]" pageId="3" pageNumber="192" type="male">2 ♂♂</specimenCount>
,
<specimenCount box="[419,470,343,363]" pageId="3" pageNumber="192" type="female">9 ♀♀</specimenCount>
(in
<collectionCode box="[505,552,344,363]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737972" collectingDate="2000-09-22" collectingDateMax="2000-09-27" collectingDateMin="2000-09-22" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" county="Danum Valley Field Centre" latitude="5.0" location="Valley Field Centre" longLatPrecision="1305" longitude="117.833336" municipality="Danum Valley" pageId="3" pageNumber="192" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sabah" typeStatus="paratype">
same locality as
<typeStatus box="[718,804,344,363]" pageId="3" pageNumber="192">holotype</typeStatus>
,
<date box="[812,934,344,363]" pageId="3" pageNumber="192" value="2000-09-22" valueMax="2000-09-27" valueMin="2000-09-22">
<collectingDate box="[812,934,344,363]" pageId="3" pageNumber="192" value="2000-09-22" valueMax="2000-09-27" valueMin="2000-09-22">22-27.ix.2000</collectingDate>
</date>
,
<collectingMethod box="[942,998,344,363]" pageId="3" pageNumber="192">pitfall</collectingMethod>
with human dung, leg.
<collectorName box="[252,403,372,391]" pageId="3" pageNumber="192">M. Schilthuizen</collectorName>
:
<specimenCount box="[411,463,371,391]" pageId="3" pageNumber="192" type="male">2 ♂♂</specimenCount>
<specimenCount box="[469,503,371,391]" pageId="3" pageNumber="192" type="female">1 ♀</specimenCount>
(
<collectionCode box="[514,562,372,391]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737987" collectingDate="1997-02-27" collectionCode="BOR" country="Malaysia" location="Kiansom" pageId="3" pageNumber="192" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Sabah" typeStatus="paratype">
<collectingCountry box="[578,666,372,391]" name="Malaysia" pageId="3" pageNumber="192">Malaysia</collectingCountry>
:
<collectingRegion box="[674,733,372,391]" country="Malaysia" name="Sabah" pageId="3" pageNumber="192">Sabah</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD:740C602F7403E66365AAFED1FCE8FE62" box="[742,826,372,391]" country="Malaysia" name="Kiansom" pageId="3" pageNumber="192" stateProvince="Sabah">Kiansom</location>
(
<locationDeviation box="[838,1109,372,391]" location="Kota Kinabalu" pageId="3" pageNumber="192">
<quantity box="[838,897,372,391]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="192" unit="km" value="20.0">20 km</quantity>
ESE of Kota Kinabalu
</locationDeviation>
),
<date box="[151,240,400,419]" pageId="3" pageNumber="192" value="1997-02-27">
<collectingDate box="[151,240,400,419]" pageId="3" pageNumber="192" value="1997-02-27">27.ii.1997</collectingDate>
</date>
,
<collectingMethod box="[249,465,400,419]" pageId="3" pageNumber="192">flight interception trap</collectingMethod>
, leg. UMS students:
<specimenCount box="[658,711,399,419]" pageId="3" pageNumber="192" type="male">2 ♂♂</specimenCount>
,
<specimenCount box="[721,756,399,419]" pageId="3" pageNumber="192" type="female">1 ♀</specimenCount>
(
<collectionCode box="[767,815,400,419]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737946" collectingDate="2000-04-11" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" location="Kiansom" pageId="3" pageNumber="192" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
same locality,
<date box="[962,1053,400,419]" pageId="3" pageNumber="192" value="2000-04-11">
<collectingDate box="[962,1053,400,419]" pageId="3" pageNumber="192" value="2000-04-11">11.iv.2000</collectingDate>
</date>
,
<collectingMethod box="[1063,1119,400,419]" pageId="3" pageNumber="192">pitfall</collectingMethod>
with chicken, leg.
<collectorName box="[328,480,428,447]" pageId="3" pageNumber="192">M. Schilthuizen</collectorName>
,
<specimenCount box="[490,545,427,447]" pageId="3" pageNumber="192" type="male">1 ♂♂</specimenCount>
,
<specimenCount box="[557,593,427,447]" pageId="3" pageNumber="192" type="female">1 ♀</specimenCount>
(
<collectionCode box="[606,654,428,447]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737996" collectingDate="2000-04-10" collectingDateMax="2000-04-12" collectingDateMin="2000-04-10" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" location="Kota Kinabalu" municipality="Tun Fuad Stephen Municipal Park" pageId="3" pageNumber="192" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Sabah" typeStatus="paratype">
<collectingCountry box="[671,759,428,447]" name="Malaysia" pageId="3" pageNumber="192">Malaysia</collectingCountry>
:
<collectingRegion box="[769,828,428,447]" country="Malaysia" name="Sabah" pageId="3" pageNumber="192">Sabah</collectingRegion>
:
<collectingMunicipality pageId="3" pageNumber="192">Tun Fuad Stephen Municipal Park</collectingMunicipality>
(outskirts of
<location LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD:740C602F7403E6636605FE6DFE05FE3E" box="[329,471,456,475]" country="Malaysia" municipality="Tun Fuad Stephen Municipal Park" name="Kota Kinabalu" pageId="3" pageNumber="192" stateProvince="Sabah">Kota Kinabalu</location>
),
<date box="[489,609,456,475]" pageId="3" pageNumber="192" value="2000-04-10" valueMax="2000-04-12" valueMin="2000-04-10">
<collectingDate box="[489,609,456,475]" pageId="3" pageNumber="192" value="2000-04-10" valueMax="2000-04-12" valueMin="2000-04-10">10-12.iv.2000</collectingDate>
</date>
,
<collectingMethod box="[621,677,456,475]" pageId="3" pageNumber="192">pitfall</collectingMethod>
with lamb, leg.
<collectorName box="[840,993,456,475]" pageId="3" pageNumber="192">M. Schilthuizen</collectorName>
,
<specimenCount box="[1005,1059,455,475]" pageId="3" pageNumber="192" type="male">2 ♂♂</specimenCount>
,
<specimenCount box="[1073,1119,455,475]" pageId="3" pageNumber="192" type="female">5♀♀</specimenCount>
(
<collectionCode box="[157,205,484,503]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
,
<materialsCitation ID-GBIF-Occurrence="2512737969" box="[219,325,483,503]" collectingDate="2000-04-10" collectingDateMax="2000-04-12" collectingDateMin="2000-04-10" collectionCode="FRCS" collectorName="M. Schilthuizen" country="Malaysia" isTailEnumeration="true" location="Kota Kinabalu" municipality="Tun Fuad Stephen Municipal Park" pageId="3" pageNumber="192" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<specimenCount box="[219,252,483,503]" pageId="3" pageNumber="192" type="male">1 ♂</specimenCount>
(
<collectionCode box="[263,319,484,503]" country="Malaysia" httpUri="http://grbio.org/cool/sxta-ddia" name="Forest Research Centre" pageId="3" pageNumber="192">FRCS</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737993" collectingDate="2000-11-24" collectingDateMax="2000-11-27" collectingDateMin="2000-11-24" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" isTailEnumeration="true" location="Kota Kinabalu" municipality="Tun Fuad Stephen Municipal Park" pageId="3" pageNumber="192" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
same locality,
<date box="[463,586,484,503]" pageId="3" pageNumber="192" value="2000-11-24" valueMax="2000-11-27" valueMin="2000-11-24">
<collectingDate box="[463,586,484,503]" pageId="3" pageNumber="192" value="2000-11-24" valueMax="2000-11-27" valueMin="2000-11-24">24-27.xi.2000</collectingDate>
</date>
,
<collectingMethod box="[593,649,484,503]" pageId="3" pageNumber="192">pitfall</collectingMethod>
with dung or beef, leg.
<collectorName box="[868,1017,484,503]" pageId="3" pageNumber="192">M. Schilthuizen</collectorName>
,
<specimenCount box="[1025,1058,483,503]" pageId="3" pageNumber="192" type="male">1 ♂</specimenCount>
,
<specimenCount box="[1068,1119,483,503]" pageId="3" pageNumber="192" type="female">3 ♀♀</specimenCount>
(
<collectionCode box="[157,205,512,531]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737948" collectionCode="BOR" collectorName="M. Schilthuizen" country="Malaysia" elevation="200" location="Sugud Forest Reserve" pageId="3" pageNumber="192" specimenCount="1" specimenCount-female="1" stateProvince="Sabah" typeStatus="paratype">
<location LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD:740C602F7403E6636790FDA5FE79FDF6" box="[220,427,512,531]" country="Malaysia" name="Sugud Forest Reserve" pageId="3" pageNumber="192" stateProvince="Sabah">Sugud Forest Reserve</location>
,
<locationDeviation box="[436,676,512,531]" location="Kota Kinabalu" pageId="3" pageNumber="192">
<quantity box="[436,494,512,531]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="192" unit="km" value="15.0">15 km</quantity>
S of Kota Kinabalu
</locationDeviation>
(
<collectingRegion box="[688,748,512,531]" country="Malaysia" name="Sabah" pageId="3" pageNumber="192">Sabah</collectingRegion>
),
<quantity box="[764,820,512,531]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="192" unit="m" value="200.0">
<elevation box="[764,820,512,531]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="192" unit="m" value="200.0">200 m</elevation>
</quantity>
alt.,
<collectingMethod box="[866,922,512,531]" pageId="3" pageNumber="192">pitfall</collectingMethod>
with carrion, leg.
<collectorName pageId="3" pageNumber="192">M. Schilthuizen</collectorName>
,
<specimenCount box="[278,312,539,559]" pageId="3" pageNumber="192" type="female">1 ♀</specimenCount>
(
<collectionCode box="[322,370,540,559]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:13160" name="Guermonprez Museum" pageId="3" pageNumber="192" type="Herbarium">BOR</collectionCode>
)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2512737956" collectingDate="1996-06-24" collectingDateMax="1996-07-01" collectingDateMin="1996-06-24" collectionCode="CJRZ, CMPR" country="Malaysia" location="Batu Punggul Resort" pageId="3" pageNumber="192" specimenCount="20" specimenCount-female="7" specimenCount-male="13" stateProvince="Sabah" typeStatus="paratype">
<collectingCountry box="[384,472,540,559]" name="Malaysia" pageId="3" pageNumber="192">Malaysia</collectingCountry>
:
<collectingRegion box="[480,539,540,559]" country="Malaysia" name="Sabah" pageId="3" pageNumber="192">Sabah</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD:740C602F7403E663656FFDB9FD35FDCA" box="[547,743,540,559]" country="Malaysia" name="Batu Punggul Resort" pageId="3" pageNumber="192" stateProvince="Sabah">Batu Punggul Resort</location>
,
<date box="[751,893,540,559]" pageId="3" pageNumber="192" value="1996-06-24" valueMax="1996-07-01" valueMin="1996-06-24">
<collectingDate box="[751,893,540,559]" pageId="3" pageNumber="192" value="1996-06-24" valueMax="1996-07-01" valueMin="1996-06-24">24.vi-1.vii.1996</collectingDate>
</date>
,
<collectingMethod box="[901,1116,540,559]" pageId="3" pageNumber="192">flight interception trap</collectingMethod>
,
<specimenCount box="[151,214,567,587]" pageId="3" pageNumber="192" type="male">13 ♂♂</specimenCount>
,
<specimenCount box="[224,276,567,587]" pageId="3" pageNumber="192" type="female">7 ♀♀</specimenCount>
(
<collectionCode box="[287,341,568,587]" pageId="3" pageNumber="192">CJRZ</collectionCode>
;
<collectionCode box="[349,415,568,587]" country="India" name="Centre for Medicinal Plants Research" pageId="3" pageNumber="192" type="Herbarium">CMPR</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="3.[151,1120,287,727]" pageId="3" pageNumber="192">
<materialsCitation ID-GBIF-Occurrence="2512737963" collectingDate="1997-02-27" collectingDateMax="2000-12-29" collectingDateMin="1997-02-27" location="Valley Field" municipality="Danum Valley" pageId="3" pageNumber="192" specimenCount="25" specimenCount-female="9" specimenCount-male="4" stateProvince="Centre">
<typeStatus box="[151,336,596,615]" isOtherTypeStatus="true" pageId="3" pageNumber="192">Additional material</typeStatus>
(not included in the type series).— Tun Fuad Stephen Municipal Park,
<date box="[993,1115,596,615]" pageId="3" pageNumber="192" value="2000-09-24" valueMax="2000-09-27" valueMin="2000-09-24">
<collectingDate box="[993,1115,596,615]" pageId="3" pageNumber="192" value="2000-09-24" valueMax="2000-09-27" valueMin="2000-09-24">24-27.ix.2000</collectingDate>
</date>
,
<specimenCount box="[151,204,623,643]" pageId="3" pageNumber="192" type="male">4 ♂♂</specimenCount>
,
<specimenCount box="[216,269,623,643]" pageId="3" pageNumber="192" type="female">9 ♀♀</specimenCount>
in 70% ethanol; Kiansom,
<date box="[526,654,624,643]" pageId="3" pageNumber="192" value="2000-12-25" valueMax="2000-12-29" valueMin="2000-12-25">
<collectingDate box="[526,654,624,643]" pageId="3" pageNumber="192" value="2000-12-25" valueMax="2000-12-29" valueMin="2000-12-25">25-29.xii.2000</collectingDate>
</date>
,
<specimenCount box="[664,779,624,643]" pageId="3" pageNumber="192" type="generic">8 specimens</specimenCount>
of unidentified sex in 70% ethanol; Kiansom,
<date box="[250,339,652,671]" pageId="3" pageNumber="192" value="1997-02-27">
<collectingDate box="[250,339,652,671]" pageId="3" pageNumber="192" value="1997-02-27">27.ii.1997</collectingDate>
</date>
,
<specimenCount box="[352,470,652,671]" pageId="3" pageNumber="192" type="generic">2 specimens</specimenCount>
of unidentified sex in 100% ethanol;
<collectingMunicipality box="[845,985,652,671]" pageId="3" pageNumber="192">
Danum
<location LSID="urn:lsid:plazi:treatment:F97A87E27403E66466D1FF65FC98FEDD:740C602F7403E66364D2FD29FC0BFD7A" box="[926,985,652,671]" municipality="Danum Valley" name="Valley Field" pageId="3" pageNumber="192" stateProvince="Centre">Valley</location>
</collectingMunicipality>
Field
<collectingRegion box="[1050,1115,652,671]" country="France" name="Centre" pageId="3" pageNumber="192">Centre</collectingRegion>
,
<date box="[151,238,680,699]" pageId="3" pageNumber="192" value="2000-05-15">
<collectingDate box="[151,238,680,699]" pageId="3" pageNumber="192" value="2000-05-15">15.v.2000</collectingDate>
</date>
,
<specimenCount box="[246,360,680,699]" pageId="3" pageNumber="192" type="generic">2 specimens</specimenCount>
of unidentified sex in 100% ethanol. (For full collection details, see Type material above.)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="192" type="diagnosis">
<paragraph blockId="3.[151,1119,768,1657]" pageId="3" pageNumber="192">
Diagnosis.— Habitus broad, rectangular, flat. Thorax on average 1.8 times as wide as long. Aedeagus (
<figureCitation box="[368,476,800,824]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="3" pageNumber="192">figs 20-21</figureCitation>
) with two apical lateral wings and a long terminal processus. Male forelegs with long hairs on the ventral side of the profemur and protibia.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="192" type="description">
<paragraph blockId="3.[151,1119,768,1657]" pageId="3" pageNumber="192">
Description.— Habitus broad and rectangular, relatively flat. Moderately pigment- ed: mostly chestnut brown; only the tarsi and the edges of thorax and elytra lighter brown. Length
<quantity box="[323,447,928,952]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.9" metricValueMin="2.1" pageId="3" pageNumber="192" unit="mm" value="2.5" valueMax="2.9" valueMin="2.1">2.1-2.9 mm</quantity>
(n = 26). Antennae slender and relatively long, 1.1-1.2 times as long as the width of the head (n = 5). Articles 6, 9, and 10 square. Male protarsi moderately dilated: the protibia at its largest width (excluding the lateral spines) is 1.5 times broader than the first article of the protarsus (n = 1). Female protarsi undilated. In the male, the ventral side of profemur and protibia carry long hairs, some of which are as long as the tarsus. Thorax 1.76-1.84 times as wide as long (n = 5), relatively flat, with the caudal corners distinctly drawn out. The mesosternal carina is strongly developed, its edge somewhat thickened. Elytra short, as long (measured from the caudal tip of the scutellum) as their combined width. Elytral apices distinctly truncate, identical in both sexes. The 3
<superScript attach="none" box="[285,299,1215,1229]" fontSize="6" pageId="3" pageNumber="192">rd-</superScript>
5
<superScript attach="left" box="[321,334,1215,1229]" fontSize="6" pageId="3" pageNumber="192">th</superScript>
visible abdominal sternite in the male each have a broad and very indistinct central notch and are slightly depressed in the vicinity of these notches. Aedeagus (
<figureCitation box="[204,310,1280,1304]" captionStart="Figs 16-25. 16" captionStartId="20.[151,190,1437,1456]" captionTargetBox="[151,1119,192,1408]" captionTargetPageId="20" captionText="Figs 16-25. 16, Ptomaphaginus anas spec. nov. holotype; aedeagus, dorsal view. 17, Ptomaphaginus anas spec. nov., holotype; aedeagus, lateral view. 18, Ptomaphaginus similipes spec. nov., holotype; aedeagus, dorsal view. 19, Ptomaphaginus similipes spec. nov., holotype; aedeagus, lateral view. 20, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, dorsal view. 21, Ptomaphaginus bryantioides spec. nov., paratype (from Kiansom); aedeagus, lateral view (basal part partly obscured by the genital segment). 22, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, ventral view. 23, Ptomaphaginus latimanus spec. nov., holotype; aedeagus, lateral view. 24, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, dorsal view. 25, Ptomaphaginus kinabaluensis spec. nov., holotype; aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3576295" httpUri="https://zenodo.org/record/3576295/files/figure.png" pageId="3" pageNumber="192">figs 20-21</figureCitation>
) in dorsal view abruptly narrowing towards the apex, and there adorned with two lateral wings. The tip is composed of a long, fragile and slightly asymmetric processus. In lateral view, the aedeagus is moderately curved, the terminal processus lies in the plane of the rest of the aedeagus, whereas the two lateral wings point ventrad. Male genital segment relatively broad, at the apex slightly triangular, 2.5 times as long as wide.
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="192" type="distribution">
<paragraph blockId="3.[151,1119,768,1657]" pageId="3" pageNumber="192">
Distribution.— This species is widely distributed in the lowlands of
<collectingRegion box="[970,1037,1472,1496]" country="Malaysia" name="Sabah" pageId="3" pageNumber="192">Sabah</collectingRegion>
(all localities lie below
<quantity box="[339,404,1504,1528]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="192" unit="m" value="300.0">300 m</quantity>
altitude), and apparently has a broad tolerance for habitat
<typeStatus box="[1052,1114,1505,1529]" pageId="3" pageNumber="192">types</typeStatus>
, having been found both in primary (Danum Valley) and in secondary forests (Tun Fuad Stephen Park, Kota Kinabalu).
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="193" pageId="3" pageNumber="192" type="discussion">
<paragraph blockId="3.[151,1119,768,1657]" lastBlockId="4.[151,1119,192,312]" lastPageId="4" lastPageNumber="193" pageId="3" pageNumber="192">
Remarks.— Based on aedeagal shape,
<taxonomicName authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[627,779,1600,1624]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="bryantioides">
<emphasis box="[627,779,1600,1624]" italics="true" pageId="3" pageNumber="192">P. bryantioides</emphasis>
</taxonomicName>
is closely related to
<taxonomicName authorityName="Jeannel" authorityYear="1936" box="[1011,1112,1600,1624]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis box="[1011,1112,1600,1624]" italics="true" pageId="3" pageNumber="192">P. bryanti</emphasis>
</taxonomicName>
, from which it differs in the habitus, which is very broad and stocky in
<taxonomicName authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[960,1113,1632,1656]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="192" phylum="Arthropoda" rank="species" species="bryantioides">
<emphasis box="[960,1113,1632,1656]" italics="true" pageId="3" pageNumber="192">P. bryantioides</emphasis>
</taxonomicName>
. For example, the thoracal index for the new species is 1.76-1.84, whereas for the
<typeStatus pageId="4" pageNumber="193">holotype</typeStatus>
of
<taxonomicName authorityName="Jeannel" authorityYear="1936" box="[236,336,224,248]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis box="[236,336,224,248]" italics="true" pageId="4" pageNumber="193">P. bryanti</emphasis>
</taxonomicName>
it is only 1.56.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="193" type="etymology">
<paragraph blockId="4.[151,1119,192,312]" pageId="4" pageNumber="193">
Etymology.—
<taxonomicName authorityName="Schilthuizen &amp; Perreau" authorityYear="2008" box="[351,477,256,280]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="bryantioides">
<emphasis box="[351,477,256,280]" italics="true" pageId="4" pageNumber="193">bryantioides</emphasis>
</taxonomicName>
, a Latin adjective, meaning like
<taxonomicName authorityName="Jeannel" authorityYear="1936" box="[844,919,256,280]" class="Insecta" family="Leiodidae" genus="Ptomaphaginus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="193" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis box="[844,919,256,280]" italics="true" pageId="4" pageNumber="193">bryanti</emphasis>
</taxonomicName>
. The name refers to the similarity in aedeagal shape between these two species.
</paragraph>
</subSubSection>
</treatment>
</document>