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<document id="26BC6A1CA31913B5595847FC7322D04F" ID-CLB-Dataset="20083" ID-DOI="10.11646/zootaxa.5120.1.4" ID-GBIF-Dataset="95654c76-8405-45f2-bbe3-358622359c7c" ID-ISSN="1175-5326" ID-Zenodo-Dep="6388863" ID-ZooBank="F3C1D9EB-74C0-4BCA-95B2-2BECD7E7DF92" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648452352969" checkinUser="plazi" docAuthor="Ruta, Rafał &amp; Yoshitomi, Hiroyuki" docDate="2022" docId="F96687F90B65FFC6FF75FF74DAC83F61" docLanguage="en" docName="zootaxa.5120.1.4.pdf" docOrigin="Zootaxa 5120 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Calvariomorphus sakaii Ruta &amp; Yoshitomi 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="78" masterDocId="055FFF810B68FFCBFFE2FFE3DA383B74" masterDocTitle="Calvariomorphus-a new genus of marsh beetles (Coleoptera: Scirtidae) with remarkable elytral excitators" masterLastPageNumber="82" masterPageNumber="65" pageNumber="78" updateTime="1699318575542" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="0327E34D2C63EB6D79D933BFD973141B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8F9B4D32A76AC4D0F2E14785E8FFBE29">
<mods:title id="C7CAFE5F1DD5BA54575F73CF40F60CB5">Calvariomorphus-a new genus of marsh beetles (Coleoptera: Scirtidae) with remarkable elytral excitators</mods:title>
</mods:titleInfo>
<mods:name id="B426EF91644D5544846BAA8AA26B581C" type="personal">
<mods:role id="77620C22F39742E81D29A22E7BD08264">
<mods:roleTerm id="291F0E197CE7C59B73BFF78086D9E1B6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2444973F98C823B52C1CDAF93B36DC66">Ruta, Rafał</mods:namePart>
<mods:nameIdentifier id="FDBE49C4C838E97A17575C0DF4D6F444" type="ORCID">0000-0001-8515-2385</mods:nameIdentifier>
<mods:affiliation id="6EB39C679FAA8E7400E1B1C7109B2E4C">Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Przybyszewskiego 65, 51 - 148 Wrocław, Poland. rafal. ruta @ uwr. edu. pl; https: // orcid. org / 0000 - 0001 - 8515 - 2385</mods:affiliation>
<mods:nameIdentifier id="1B61071A5C23F1E768DA957DEEC99AEE" type="email">rafal.ruta@uwr.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name id="4637EE9C52AC2D3B7135E18ABE88428F" type="personal">
<mods:role id="2CDB6687A2B36E9995862B523236E0F9">
<mods:roleTerm id="032A7AC75432E91A1FA2FD08C085A9D1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3E6E327C92A56B2CAD70A8E7E281C750">Yoshitomi, Hiroyuki</mods:namePart>
<mods:affiliation id="2956E7AE0626D72E86916CE8D30263DA">Ehime University Museum, Bunkyo 3, Matsuyama, 790 - 8577 Japan.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8760E06D64408D4EF2B3C321B038D62D">text</mods:typeOfResource>
<mods:relatedItem id="64CA4F30324AD12C3BCC484F843DF73E" type="host">
<mods:titleInfo id="0E3E01975B6358E2897E973FA5E3C004">
<mods:title id="91C50DDFAC4F7E574D61D51717340CC3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="18BBC0A97FE0FF158A066FB30351CA9C">
<mods:date id="E0CBE5ECE6ABD1F5D8DFBC95419CE371">2022</mods:date>
<mods:detail id="983649305D1A5D67B55FBC18D072F35E" type="pubDate">
<mods:number id="67DFF262B33F442A3E0841F48DD2325E">2022-03-24</mods:number>
</mods:detail>
<mods:detail id="03E735E621E311A84E37293210AF03D7" type="volume">
<mods:number id="B1DC27BCA8B64A5B8DDA072669DC4D26">5120</mods:number>
</mods:detail>
<mods:detail id="51AB62CEB9A2C7ECD672060FBFC60B6C" type="issue">
<mods:number id="1CF75ECCE5A7D3EE16FD2D6B2C68A022">1</mods:number>
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<mods:extent id="FE74C023B7BA276AB2B1632E0130F34A" unit="page">
<mods:start id="196326F42A8AD091309FDE6098CBD943">65</mods:start>
<mods:end id="5763031D2E39035DF4B698C72B00217F">82</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="991ADCB4835B04E9A4A88AB4AB0F52B4">journal article</mods:classification>
<mods:identifier id="7480518AC8C7B4CBCDB3211A9551604E" type="CLB-Dataset">20083</mods:identifier>
<mods:identifier id="41C4C191FA17B2BB5A586B64B4F89420" type="DOI">10.11646/zootaxa.5120.1.4</mods:identifier>
<mods:identifier id="9617453DC4DCC24B6974F3DF57347EB5" type="GBIF-Dataset">95654c76-8405-45f2-bbe3-358622359c7c</mods:identifier>
<mods:identifier id="31D012460AE9D9882249128E151BFFB2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D5BEE44CA0343AB83375CFC25A1F5F28" type="Zenodo-Dep">6388863</mods:identifier>
<mods:identifier id="699E5855D32D149BC4D418DA10EAFFCF" type="ZooBank">F3C1D9EB-74C0-4BCA-95B2-2BECD7E7DF92</mods:identifier>
</mods:mods>
<treatment id="F96687F90B65FFC6FF75FF74DAC83F61" ID-DOI="http://doi.org/10.5281/zenodo.6393856" ID-GBIF-Taxon="194286790" ID-Zenodo-Dep="6393856" LSID="urn:lsid:plazi:treatment:F96687F90B65FFC6FF75FF74DAC83F61" httpUri="http://treatment.plazi.org/id/F96687F90B65FFC6FF75FF74DAC83F61" lastPageNumber="78" pageId="13" pageNumber="78">
<subSubSection id="39D565640B65FFC6FF75FF74D8273BC5" box="[151,543,151,178]" pageId="13" pageNumber="78" type="nomenclature">
<paragraph id="717036EF0B65FFC6FF75FF74D8273BC5" blockId="13.[151,543,151,212]" box="[151,543,151,178]" pageId="13" pageNumber="78">
<heading id="2A3881830B65FFC6FF75FF74D8273BC5" bold="true" box="[151,543,151,178]" fontSize="11" level="1" pageId="13" pageNumber="78" reason="1">
<emphasis id="43BBEAFD0B65FFC6FF75FF74D8273BC5" bold="true" box="[151,543,151,178]" pageId="13" pageNumber="78">
<taxonomicName id="B6CF4D6C0B65FFC6FF75FF74DB823BC6" authority="Ruta &amp; Yoshitomi, 2022" authorityName="Ruta &amp; Yoshitomi" authorityYear="2022" box="[151,442,151,178]" class="Insecta" family="Scirtidae" genus="Calvariomorphus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="78" phylum="Arthropoda" rank="species" species="sakaii" status="sp. nov.">
<emphasis id="43BBEAFD0B65FFC6FF75FF74DB823BC6" bold="true" box="[151,442,151,178]" italics="true" pageId="13" pageNumber="78">Calvariomorphus sakaii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="588857860B65FFC6FE23FF74D8273BC5" box="[449,543,151,177]" pageId="13" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="39D565640B65FFC6FF75FF59DBA73BA0" box="[151,415,186,212]" pageId="13" pageNumber="78" type="description">
<paragraph id="717036EF0B65FFC6FF75FF59DBA73BA0" blockId="13.[151,543,151,212]" box="[151,415,186,212]" pageId="13" pageNumber="78">
(
<figureCitation id="E9F42A6A0B65FFC6FF7DFF59DB2D3BA0" box="[159,277,186,212]" captionStart="FIGURE 1" captionStartId="2.[151,250,1894,1918]" captionTargetBox="[151,1435,179,1868]" captionTargetId="figure-28@2.[151,1435,179,1868]" captionTargetPageId="2" captionText="FIGURE 1. Calvariomorphus gen. nov., dorsal view (with the exception of H). A) C. malayanus sp. nov., male, B) same, fe- male, C) C. palauensis sp. nov., male, D) same, female, E) C. sakaii sp. nov., male, F) same, female, G) C. sp. (Nicobar Islands), H) same, semilateral view. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6388865" httpUri="https://zenodo.org/record/6388865/files/figure.png" pageId="13" pageNumber="78">Figs 1E–F</figureCitation>
,
<figureCitation id="E9F42A6A0B65FFC6FEC2FF59DB7B3BA0" box="[288,323,186,212]" captionStart="FIGURE 7" captionStartId="7.[151,250,1785,1809]" captionTargetBox="[151,1435,855,1762]" captionTargetId="figure-47@7.[151,1435,855,1762]" captionTargetPageId="7" captionText="FIGURE 7. Calvariomorphus gen. nov., abdomen. A) C. malayanus sp. nov., male, B) C. sakaii sp. nov., female, C) same, close-up of ventrite 4." figureDoi="http://doi.org/10.5281/zenodo.6388881" httpUri="https://zenodo.org/record/6388881/files/figure.png" pageId="13" pageNumber="78">7B</figureCitation>
,
<figureCitation id="E9F42A6A0B65FFC6FEADFF59DB553BA0" box="[335,365,186,212]" captionStart="FIGURE 13" captionStartId="13.[151,250,1953,1977]" captionTargetBox="[151,1435,1069,1929]" captionTargetId="figure-396@13.[151,1435,1069,1929]" captionTargetPageId="13" captionText="FIGURE 13. Calvariomorphus sakaii sp. nov., male genitalia. A) penis, B) tegmen, C) sternite IX and tergite IX, D) tergite VIII. Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6388893" httpUri="https://zenodo.org/record/6388893/files/figure.png" pageId="13" pageNumber="78">13</figureCitation>
,
<figureCitation id="E9F42A6A0B65FFC6FE9BFF59DBAE3BA0" box="[377,406,186,212]" captionStart="FIGURE 14" captionStartId="14.[151,250,1056,1080]" captionTargetBox="[151,1435,181,1032]" captionTargetId="figure-28@14.[151,1435,181,1032]" captionTargetPageId="14" captionText="FIGURE 14. Calvariomorphus sakaii sp. nov., female genitalia. A) genital tract, B) coxites, C) bursal sclerite. Scale bar = 0.5 mm (A, C), 0.25 mm (B)." figureDoi="http://doi.org/10.5281/zenodo.6388895" httpUri="https://zenodo.org/record/6388895/files/figure.png" pageId="13" pageNumber="78">14</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="39D565640B65FFC6FF75FF1CD99F3AF5" pageId="13" pageNumber="78" type="materials_examined">
<paragraph id="717036EF0B65FFC6FF75FF1CD99F3AF5" blockId="13.[151,1437,254,1045]" pageId="13" pageNumber="78">
<emphasis id="43BBEAFD0B65FFC6FF75FF1CDB7E3A6D" bold="true" box="[151,326,255,281]" pageId="13" pageNumber="78">Type material.</emphasis>
<materialsCitation id="C1A73CB20B65FFC6FEACFF1CDBCF3A48" ID-GBIF-Occurrence="3738405305" collectingDate="1985-08-04" collectionCode="EUM" collectorName="M. Sakai" county="Eagle" elevation="1100" location="Mt. Apo" municipality="Baracatan" pageId="13" pageNumber="78" specimenCount="1" stateProvince="Centre" typeStatus="holotype">
<emphasis id="43BBEAFD0B65FFC6FEACFF1CD8503A6D" bold="true" box="[334,616,255,281]" pageId="13" pageNumber="78">
<typeStatus id="AE74884D0B65FFC6FEACFF1CDB853A6D" box="[334,445,255,281]" pageId="13" pageNumber="78" type="holotype">Holotype</typeStatus>
, male (
<collectionCode id="17DEAE2A0B65FFC6FDF1FF1CD8633A6D" box="[531,603,255,281]" pageId="13" pageNumber="78">EUM</collectionCode>
):
</emphasis>
“[MINDANAO] \
<collectingCounty id="98114E630B65FFC6FCA0FF1CD9BA3A6D" box="[834,898,255,281]" pageId="13" pageNumber="78">Eagle</collectingCounty>
<collectingRegion id="B30BF80D0B65FFC6FC68FF1CD9E03A6D" box="[906,984,255,281]" country="France" name="Centre" pageId="13" pageNumber="78">Centre</collectingRegion>
,
<quantity id="B6379B0A0B65FFC6FC01FF1CDE083A6D" box="[995,1072,255,281]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="78" unit="m" value="1100.0">
<elevation id="FAE2D1DC0B65FFC6FC01FF1CDE083A6D" box="[995,1072,255,281]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="78" unit="m" value="1100.0">1100m</elevation>
</quantity>
\
<collectingMunicipality id="9114AC950B65FFC6FBAAFEE3DE803A6E" box="[1096,1208,256,282]" pageId="13" pageNumber="78">Baracatan</collectingMunicipality>
north \ slope of
<location id="741060340B65FFC6FA97FEE3DAFF3A48" LSID="urn:lsid:plazi:treatment:F96687F90B65FFC6FF75FF74DAC83F61:741060340B65FFC6FA97FEE3DAFF3A48" county="Eagle" municipality="Baracatan" name="Mt. Apo" pageId="13" pageNumber="78" stateProvince="Centre">Mt. Apo</location>
\
<date id="0571102F0B65FFC6FF03FEC1DB4F3A49" box="[225,375,290,317]" pageId="13" pageNumber="78" value="1985-08-04">
<collectingDate id="1535E9C70B65FFC6FF03FEC1DB4F3A49" box="[225,375,290,317]" pageId="13" pageNumber="78" value="1985-08-04">4. VIII. 1985</collectingDate>
</date>
<collectorName id="DC3A53390B65FFC6FE63FEC1DBD63A48" box="[385,494,290,316]" pageId="13" pageNumber="78">M. Sakai</collectorName>
”
</materialsCitation>
.
<materialsCitation id="C1A73CB20B65FFC6FDE6FEC1DEE23A48" ID-GBIF-Occurrence="3738405312" box="[516,1242,289,316]" collectingDate="1985-08-04" collectionCode="EUM" collectorName="M. Sakai" county="Eagle" elevation="1100" location="Mt. Apo" municipality="Baracatan" pageId="13" pageNumber="78" specimenCount="8" specimenCount-female="2" specimenCount-male="6" stateProvince="Centre" typeStatus="paratype">
<emphasis id="43BBEAFD0B65FFC6FDE6FEC1D9F73A4F" bold="true" box="[516,975,289,316]" pageId="13" pageNumber="78">
<typeStatus id="AE74884D0B65FFC6FDE6FEC1D8453A48" box="[516,637,290,316]" pageId="13" pageNumber="78" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="67C9FD660B65FFC6FD6EFEC1D8D13A48" box="[652,745,290,316]" pageId="13" pageNumber="78" type="male">6 males</specimenCount>
,
<specimenCount id="67C9FD660B65FFC6FD1AFEC1D9513A4F" box="[760,873,289,316]" pageId="13" pageNumber="78" type="female">2 females</specimenCount>
(
<collectionCode id="17DEAE2A0B65FFC6FC98FEC2D9FA3A4F" box="[890,962,289,315]" pageId="13" pageNumber="78">EUM</collectionCode>
):
</emphasis>
same data as holotype
</materialsCitation>
.
<materialsCitation id="C1A73CB20B65FFC6FB04FEC1D91D3A2B" ID-GBIF-Occurrence="3738405303" collectingDate="1985-08-05" collectionCode="EUM" collectorName="M. Sakai" county="Eagle" elevation="1100" location="Mt. Apo" municipality="Baracatan" pageId="13" pageNumber="78" specimenCount="1" stateProvince="Centre" typeStatus="paratype">
<emphasis id="43BBEAFD0B65FFC6FB04FEC1DACC3A2A" bold="true" pageId="13" pageNumber="78">
<typeStatus id="AE74884D0B65FFC6FB04FEC1DF6D3A48" box="[1254,1365,290,316]" pageId="13" pageNumber="78" type="paratype">Paratype</typeStatus>
, male (
<collectionCode id="17DEAE2A0B65FFC6FF7DFEA7DADF3A2A" box="[159,231,324,350]" pageId="13" pageNumber="78">EUM</collectionCode>
):
</emphasis>
same data as holotype, but collected
<date id="0571102F0B65FFC6FD7CFEA7D91D3A2B" box="[670,805,324,351]" pageId="13" pageNumber="78" value="1985-08-05">
<collectingDate id="1535E9C70B65FFC6FD7CFEA7D91D3A2B" box="[670,805,324,351]" pageId="13" pageNumber="78" value="1985-08-05">5.VIII.1985</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="C1A73CB20B65FFC6FCD3FEA6D99B3AF5" ID-GBIF-Occurrence="3738405313" collectingDate="1959-10-26" collectionCode="EUM" collectorName="L. W. Quate" elevation="1250" location="Mt. Katanglad" pageId="13" pageNumber="78" specimenCount="1" stateProvince="Centre" typeStatus="paratype">
<emphasis id="43BBEAFD0B65FFC6FCD3FEA6DE743A2A" bold="true" box="[817,1100,324,351]" pageId="13" pageNumber="78">
<typeStatus id="AE74884D0B65FFC6FCD3FEA6D9983A2B" box="[817,928,325,351]" pageId="13" pageNumber="78" type="paratype">Paratype</typeStatus>
, male (
<collectionCode id="17DEAE2A0B65FFC6FC15FEA7DE073A2A" box="[1015,1087,324,350]" pageId="13" pageNumber="78">EUM</collectionCode>
):
</emphasis>
“P. I., MINDANAO \ Bukindon,
<quantity id="B6379B0A0B65FFC6FF2DFE84DB193AF6" box="[207,289,359,386]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="13" pageNumber="78" unit="m" value="1250.0">
<elevation id="FAE2D1DC0B65FFC6FF2DFE84DB193AF6" box="[207,289,359,386]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="13" pageNumber="78" unit="m" value="1250.0">1250m</elevation>
</quantity>
. \
<location id="741060340B65FFC6FED8FE8BDBE33AF5" LSID="urn:lsid:plazi:treatment:F96687F90B65FFC6FF75FF74DAC83F61:741060340B65FFC6FED8FE8BDBE33AF5" box="[314,475,359,386]" name="Mt. Katanglad" pageId="13" pageNumber="78" stateProvince="Centre">Mt. Katanglad</location>
\
<date id="0571102F0B65FFC6FE13FE84D8533AF6" box="[497,619,359,386]" pageId="13" pageNumber="78" value="1959-10-26">
<collectingDate id="1535E9C70B65FFC6FE13FE84D8533AF6" box="[497,619,359,386]" pageId="13" pageNumber="78" value="1959-10-26">26-x-1959</collectingDate>
</date>
”; “
<collectorName id="DC3A53390B65FFC6FD6FFE84D92D3AF5" box="[653,789,359,386]" pageId="13" pageNumber="78">L. W. Quate</collectorName>
\ Collector”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="39D565640B65FFC6FF25FE6ADF0C3AB2" pageId="13" pageNumber="78" type="diagnosis">
<paragraph id="717036EF0B65FFC6FF25FE6ADF0C3AB2" blockId="13.[151,1437,254,1045]" pageId="13" pageNumber="78">
<emphasis id="43BBEAFD0B65FFC6FF25FE6ADB7A3AD7" bold="true" box="[199,322,393,419]" pageId="13" pageNumber="78">Diagnosis.</emphasis>
Small species, on average larger than other members of the genus (TL ~ 2.0–
<quantity id="B6379B0A0B65FFC6FB45FE69DF393AD0" box="[1191,1281,394,420]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="13" pageNumber="78" unit="mm" value="2.2">2.2 mm</quantity>
), body widest in the basal half of elytra, trigonium of penis as long as parameroids, elytra of females without excitators.
</paragraph>
</subSubSection>
<subSubSection id="39D565640B65FFC6FF25FE2CDF1538D8" pageId="13" pageNumber="78" type="description">
<paragraph id="717036EF0B65FFC6FF25FE2CDE3439E3" blockId="13.[151,1437,254,1045]" pageId="13" pageNumber="78">
<emphasis id="43BBEAFD0B65FFC6FF25FE2CDB613A9D" bold="true" box="[199,345,463,489]" pageId="13" pageNumber="78">Description.</emphasis>
Body small, TL 2.03–2.20 (mean 2.11, n=8) mm, TL/EW ~ 1.4–1.5 (1.4), oval, brownish black, legs and antennae lighter, yellowish, covered with dense, yellowish, erect setae. Head strongly transverse, punctation sparse, punctures separated by ca. 2 diameters. Pronotum transverse, anterolateral and posterolateral angles right angled, lateral carinae almost straight, punctation sparse, similar to that on head. Angle between pronotum and elytra not marked in dorsal view. Scutellar shield subtriangular, impunctate. Elytra elongate, widest in anterior half, punctation stronger than on pronotum, punctures separated by ca. 1.0 diameter.
</paragraph>
<paragraph id="717036EF0B65FFC6FF25FD7CDF0F3830" blockId="13.[151,1437,254,1045]" pageId="13" pageNumber="78">
Penis (L
<quantity id="B6379B0A0B65FFC6FECFFD7CDBAD39CE" box="[301,405,671,698]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="13" pageNumber="78" unit="mm" value="0.58">0.58 mm</quantity>
,
<geoCoordinate id="14FB50280B65FFC6FE43FD43DBCC39CD" box="[417,500,671,698]" degrees="0.12" direction="west" orientation="longitude" pageId="13" pageNumber="78" precision="555" value="-0.12">
W
<quantity id="B6379B0A0B65FFC6FE21FD7CDBCC39CD" box="[451,500,671,697]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="78" unit="mm" value="0.12">0.12</quantity>
</geoCoordinate>
mm) elongate, with short, oval pala and long parameroids, parameroids and trigonium of similar lengths, apex of trigonium subtriangular; tegmen (L
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,
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W
<quantity id="B6379B0A0B65FFC6FBE1FD21DE0B39A8" box="[1027,1075,706,732]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="13" pageNumber="78" unit="mm" value="0.19">0.19</quantity>
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mm) u-shaped, with very narrow parameres and pointed apices; sternite IX (L
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,
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W
<quantity id="B6379B0A0B65FFC6FCEAFD06D90F398B" box="[776,823,741,767]" metricMagnitude="-4" metricUnit="m" metricValue="1.1" pageId="13" pageNumber="78" unit="mm" value="0.11">0.11</quantity>
</geoCoordinate>
mm) u-shaped, with very narrow lateral portions; tergite VIII (L
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,
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W
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mm) with apodemes as long as apical plate, margin of apical plate with sparse, long setae and dense, much shorter ones; tergite IX (L
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,
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W
<quantity id="B6379B0A0B65FFC6FD3AFCC9D9303830" box="[728,776,810,836]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="13" pageNumber="78" unit="mm" value="0.32">0.32</quantity>
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mm) with membranous, subtriangular apical plate.
</paragraph>
<paragraph id="717036EF0B65FFC6FF25FCAFDF1538D8" blockId="13.[151,1437,254,1045]" pageId="13" pageNumber="78">
Female. Size similar to that in males, TL 2.11–2.23 (2.17) mm, TL/EW ~ 1.4–1.6 (1.5). Elytra without excitators. Ventrite 4 with small, oval microreticulate area in central portion (
<figureCitation id="E9F42A6A0B65FFC6FC08FC8CDE7C38FD" box="[1002,1092,879,905]" captionStart="FIGURE 7" captionStartId="7.[151,250,1785,1809]" captionTargetBox="[151,1435,855,1762]" captionTargetId="figure-47@7.[151,1435,855,1762]" captionTargetPageId="7" captionText="FIGURE 7. Calvariomorphus gen. nov., abdomen. A) C. malayanus sp. nov., male, B) C. sakaii sp. nov., female, C) same, close-up of ventrite 4." figureDoi="http://doi.org/10.5281/zenodo.6388881" httpUri="https://zenodo.org/record/6388881/files/figure.png" pageId="13" pageNumber="78">Fig. 7C</figureCitation>
). Ovipositor long (L
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), bursal sclerite relatively big (L
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), subrectangular, plate-like, covered with subtle microsculpture.
</paragraph>
</subSubSection>
<subSubSection id="39D565640B65FFC6FF25FC57D96A38BB" box="[199,850,948,975]" pageId="13" pageNumber="78" type="distribution">
<paragraph id="717036EF0B65FFC6FF25FC57D96A38BB" blockId="13.[151,1437,254,1045]" box="[199,850,948,975]" pageId="13" pageNumber="78">
<emphasis id="43BBEAFD0B65FFC6FF25FC57DB5938BA" bold="true" box="[199,353,948,974]" pageId="13" pageNumber="78">Distribution.</emphasis>
Known from two localities on Mindanao Is.
</paragraph>
</subSubSection>
<subSubSection id="39D565640B65FFC6FF25FC34DAC83F61" pageId="13" pageNumber="78" type="etymology">
<paragraph id="717036EF0B65FFC6FF25FC34DAC83F61" blockId="13.[151,1437,254,1045]" pageId="13" pageNumber="78">
<emphasis id="43BBEAFD0B65FFC6FF25FC34DB763885" bold="true" box="[199,334,983,1009]" pageId="13" pageNumber="78">Etymology.</emphasis>
Named after Dr Masahiro Sakai, who is a coleopterist and was the collector of the
<typeStatus id="AE74884D0B65FFC6FB0FFC34DF773885" box="[1261,1359,983,1009]" pageId="13" pageNumber="78" type="holotype">holotype</typeStatus>
of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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