210 lines
22 KiB
XML
210 lines
22 KiB
XML
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<mods:title id="7DF8DED58F987AF18E8E217E47C424C9">Calvariomorphus-a new genus of marsh beetles (Coleoptera: Scirtidae) with remarkable elytral excitators</mods:title>
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<mods:namePart id="22C2A2F49EF6AAA8391A8C9C91E4A825">Ruta, Rafał</mods:namePart>
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<mods:affiliation id="CE7A2C1D7E1F6B82541F7434587CF81E">Department of Biodiversity and Evolutionary Taxonomy, University of Wrocław, Przybyszewskiego 65, 51 - 148 Wrocław, Poland. rafal. ruta @ uwr. edu. pl; https: // orcid. org / 0000 - 0001 - 8515 - 2385</mods:affiliation>
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<mods:nameIdentifier id="865CF397305ACC4BCB8A122D7DAF11E0" type="email">rafal.ruta@uwr.edu.pl</mods:nameIdentifier>
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<mods:namePart id="A446CD0D79F39122953B56F97B1DA593">Yoshitomi, Hiroyuki</mods:namePart>
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<mods:affiliation id="8BD84F5D33966C9DAFCF636C2097F2A2">Ehime University Museum, Bunkyo 3, Matsuyama, 790 - 8577 Japan.</mods:affiliation>
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<mods:title id="1B4ECA3CD4AE20E3C89EF86EE1E7D91F">Zootaxa</mods:title>
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<mods:date id="2743E64C970632E1B3FA6AFC841E984C">2022</mods:date>
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<mods:number id="99A9284F802FD568872D6B065B87E095">5120</mods:number>
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<paragraph id="717036EF0B64FFC7FF75FF74D9513BC5" blockId="12.[151,873,151,213]" box="[151,873,151,178]" pageId="12" pageNumber="77">
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<heading id="2A3881830B64FFC7FF75FF74D9513BC5" bold="true" box="[151,873,151,178]" fontSize="11" level="1" pageId="12" pageNumber="77" reason="1">
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<emphasis id="43BBEAFD0B64FFC7FF75FF74D9513BC5" bold="true" box="[151,873,151,178]" pageId="12" pageNumber="77">
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<taxonomicName id="B6CF4D6C0B64FFC7FF75FF74D8ED3BC5" authority="(Klausnitzer, 1973)" authorityName="Ruta & Yoshitomi" authorityYear="2022" baseAuthorityName="Ruta & Yoshitomi" baseAuthorityYear="1973" box="[151,725,151,178]" class="Insecta" family="Scirtidae" genus="Calvariomorphus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="77" phylum="Arthropoda" rank="species" species="peterseni" status="comb. nov.">
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<emphasis id="43BBEAFD0B64FFC7FF75FF74DBE73BC5" bold="true" box="[151,479,151,178]" italics="true" pageId="12" pageNumber="77">Calvariomorphus peterseni</emphasis>
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(
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<bibRefCitation id="155E4B1E0B64FFC7FE0DFF74D8F43BC5" author="Klausnitzer, B." box="[495,716,151,177]" pageId="12" pageNumber="77" pagination="97 - 110" refId="ref7136" refString="Klausnitzer, B. (1973) Zur Kenntnis der Cyphon-Fauna der Philippinen, Sumbawas, Neuguineas und des Bismarck-Archipels (Coleoptera, Helodidae). Folia Entomologica Hungarica, 26 (Supplement), 97 - 110." type="journal article" year="1973">Klausnitzer, 1973</bibRefCitation>
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)
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</taxonomicName>
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,
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<taxonomicNameLabel id="588857860B64FFC7FD00FF74D9513BC5" box="[738,873,151,177]" pageId="12" pageNumber="77" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="39D565640B64FFC7FF75FF58DB223BA1" box="[151,282,187,213]" pageId="12" pageNumber="77" type="description">
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<paragraph id="717036EF0B64FFC7FF75FF58DB223BA1" blockId="12.[151,873,151,213]" box="[151,282,187,213]" pageId="12" pageNumber="77">
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(
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<figureCitation id="E9F42A6A0B64FFC7FF7DFF58DAD03BA1" box="[159,232,187,213]" captionStart="FIGURE 6" captionStartId="7.[151,250,771,795]" captionTargetBox="[151,1435,181,747]" captionTargetId="figure-20@7.[151,1435,181,747]" captionTargetPageId="7" captionText="FIGURE 6. Calvariomorphus peterseni (Klausnitzer), comb. nov., holotype, ventral view.A) head, ventral view, B) same, head and ventral portion of thorax, ventral view." figureDoi="http://doi.org/10.5281/zenodo.6388879" httpUri="https://zenodo.org/record/6388879/files/figure.png" pageId="12" pageNumber="77">Figs 6</figureCitation>
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,
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<figureCitation id="E9F42A6A0B64FFC7FF16FF58DB293BA1" box="[244,273,187,213]" captionStart="FIGURE 12" captionStartId="12.[151,250,1815,1839]" captionTargetBox="[151,1435,1126,1791]" captionTargetId="figure-337@12.[151,1435,1126,1791]" captionTargetPageId="12" captionText="FIGURE 12. Calvariomorphus peterseni (Klausnitzer), comb. nov., holotype, male genitalia. A) penis, B) tegmen, C) sternite IX, D) tergite IX. Scale bar = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.6388891" httpUri="https://zenodo.org/record/6388891/files/figure.png" pageId="12" pageNumber="77">12</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="39D565640B64FFC7FF75FEE3D96C3A42" pageId="12" pageNumber="77" type="reference_group">
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<paragraph id="717036EF0B64FFC7FF75FEE3D8733A6C" blockId="12.[151,852,256,310]" box="[151,587,256,280]" pageId="12" pageNumber="77">
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<treatmentCitationGroup id="51DF11C10B64FFC7FF75FEE3D8733A6C" box="[151,587,256,280]" pageId="12" pageNumber="77">
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=
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<treatmentCitation id="F06E10FE0B64FFC7FF4EFEE3D8733A6C" author="Klausnitzer, B." box="[172,587,256,280]" page="109" pageId="12" pageNumber="77" year="1973">
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<taxonomicName id="B6CF4D6C0B64FFC7FF4EFEE3D8733A6C" authority="Klausnitzer, 1973: 109" authorityName="Klausnitzer" authorityPageNumber="109" authorityYear="1973" box="[172,587,256,280]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="77" phylum="Arthropoda" rank="species" species="peterseni">
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<emphasis id="43BBEAFD0B64FFC7FF4EFEE3DB663A63" box="[172,350,256,280]" italics="true" pageId="12" pageNumber="77">Cyphon peterseni</emphasis>
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<bibRefCitation id="155E4B1E0B64FFC7FE86FEE3D8733A6C" author="Klausnitzer, B." box="[356,587,256,280]" pageId="12" pageNumber="77" pagination="97 - 110" refId="ref7136" refString="Klausnitzer, B. (1973) Zur Kenntnis der Cyphon-Fauna der Philippinen, Sumbawas, Neuguineas und des Bismarck-Archipels (Coleoptera, Helodidae). Folia Entomologica Hungarica, 26 (Supplement), 97 - 110." type="journal article" year="1973">Klausnitzer, 1973: 109</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="717036EF0B64FFC7FF75FEFCD96C3A42" blockId="12.[151,852,256,310]" box="[151,852,286,310]" pageId="12" pageNumber="77">
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<treatmentCitationGroup id="51DF11C10B64FFC7FF75FEFCD96C3A42" box="[151,852,286,310]" pageId="12" pageNumber="77">
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=
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<taxonomicName id="B6CF4D6C0B64FFC7FF4EFEFDD85C3A42" authority="(Klausnitzer, 1973)" baseAuthorityName="Klausnitzer" baseAuthorityYear="1973" box="[172,612,286,310]" class="Insecta" family="Scirtidae" genus="Contacyphon" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="77" phylum="Arthropoda" rank="species" species="peterseni">
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<emphasis id="43BBEAFD0B64FFC7FF4EFEFDDBAE3A42" box="[172,406,286,310]" italics="true" pageId="12" pageNumber="77">Contacyphon peterseni</emphasis>
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(
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<bibRefCitation id="155E4B1E0B64FFC7FE41FEFDD8643A42" author="Klausnitzer, B." box="[419,604,286,310]" pageId="12" pageNumber="77" pagination="97 - 110" refId="ref7136" refString="Klausnitzer, B. (1973) Zur Kenntnis der Cyphon-Fauna der Philippinen, Sumbawas, Neuguineas und des Bismarck-Archipels (Coleoptera, Helodidae). Folia Entomologica Hungarica, 26 (Supplement), 97 - 110." type="journal article" year="1973">Klausnitzer, 1973</bibRefCitation>
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)
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</taxonomicName>
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:
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<treatmentCitation id="F06E10FE0B64FFC7FD8CFEFDD96C3A42" author="Zwick, P. & Klausnitzer, B. & Ruta, R." box="[622,852,286,310]" page="345" pageId="12" pageNumber="77" year="2013">
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<bibRefCitation id="155E4B1E0B64FFC7FD8CFEFDD96C3A42" author="Zwick, P. & Klausnitzer, B. & Ruta, R." box="[622,852,286,310]" pageId="12" pageNumber="77" pagination="337 - 353" refId="ref7567" refString="Zwick, P., Klausnitzer, B. & Ruta, R. (2013) Contacyphon Gozis, 1886 removed from synonymy (Coleoptera: Scirtidae) to ac- commodate species so far combined with the invalid name, Cyphon Paykull, 1799. Entomologische Blatter und Coleoptera, 109, 337 - 353." type="journal article" year="2013">
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Zwick
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<emphasis id="43BBEAFD0B64FFC7FD57FEFCD8D03A42" box="[693,744,286,310]" italics="true" pageId="12" pageNumber="77">et al.</emphasis>
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2013: 345
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</bibRefCitation>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="39D565640B64FFC7FF75FE86D90C3AD7" pageId="12" pageNumber="77" type="materials_examined">
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<paragraph id="717036EF0B64FFC7FF75FE86D90C3AD7" blockId="12.[151,1437,356,1067]" pageId="12" pageNumber="77">
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<emphasis id="43BBEAFD0B64FFC7FF75FE86DB7D3A0A" bold="true" box="[151,325,356,383]" pageId="12" pageNumber="77">Type material.</emphasis>
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<materialsCitation id="C1A73CB20B64FFC7FEAEFE86D90C3AD7" ID-GBIF-Occurrence="3738405307" collectingDate="1962-03-07" collectionCode="ZMUC" county="Sumuna" location="Bismarck Isl." municipality="Isl. Dyaul" pageId="12" pageNumber="77" specimenCount="2" typeStatus="holotype">
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<emphasis id="43BBEAFD0B64FFC7FEAEFE86D8403A0A" bold="true" box="[332,632,356,383]" pageId="12" pageNumber="77">
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<typeStatus id="AE74884D0B64FFC7FEAEFE86DB833A0B" box="[332,443,357,383]" pageId="12" pageNumber="77" type="holotype">Holotype</typeStatus>
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, male (
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<collectionCode id="17DEAE2A0B64FFC7FDEDFE87D8533A0A" box="[527,619,356,382]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="12" pageNumber="77">ZMUC</collectionCode>
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):
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</emphasis>
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“
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<location id="741060340B64FFC7FD6BFE87D91A3A0A" LSID="urn:lsid:plazi:treatment:F96687F90B64FFC7FF75FF74DF353F5E:741060340B64FFC7FD6BFE87D91A3A0A" box="[649,802,356,382]" county="Sumuna" municipality="Isl. Dyaul" name="Bismarck Isl." pageId="12" pageNumber="77">
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Bismarck
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<collectingMunicipality id="9114AC950B64FFC7FD1CFE87D91A3A0A" box="[766,802,356,382]" pageId="12" pageNumber="77">Isl.</collectingMunicipality>
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</location>
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Dyaul \
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<collectingCounty id="98114E630B64FFC7FC60FE86D9E63A0B" box="[898,990,357,383]" pageId="12" pageNumber="77">Sumuna</collectingCounty>
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\
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<date id="0571102F0B64FFC7FC16FE86DEA03A0A" box="[1012,1176,356,383]" pageId="12" pageNumber="77" value="1962-03-07">
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<collectingDate id="1535E9C70B64FFC7FC16FE86DEA03A0A" box="[1012,1176,356,383]" pageId="12" pageNumber="77" value="1962-03-07">7. March 1962</collectingDate>
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</date>
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\ Noona Dan. Exp. 61- 62”; “
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<typeStatus id="AE74884D0B64FFC7FF3AFE6BDB683AD6" box="[216,336,392,418]" pageId="12" pageNumber="77" type="holotype">Holotypus</typeStatus>
|
||
\
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||
<taxonomicName id="B6CF4D6C0B64FFC7FE87FE6BDB863AD6" box="[357,446,392,418]" class="Insecta" family="Scirtidae" genus="Cyphon" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="82" phylum="Arthropoda" rank="genus">Cyphon</taxonomicName>
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\ peterseni \
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<bibRefCitation id="155E4B1E0B64FFC7FDADFE6AD91E3AD7" author="Klausnitzer, B." box="[591,806,393,419]" pageId="12" pageNumber="77" pagination="97 - 110" refId="ref7136" refString="Klausnitzer, B. (1973) Zur Kenntnis der Cyphon-Fauna der Philippinen, Sumbawas, Neuguineas und des Bismarck-Archipels (Coleoptera, Helodidae). Folia Entomologica Hungarica, 26 (Supplement), 97 - 110." type="journal article" year="1973">KLAUSNITZER, 1973</bibRefCitation>
|
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”.
|
||
</materialsCitation>
|
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="39D565640B64FFC7FF25FE4FD9583A9E" pageId="12" pageNumber="77" type="diagnosis">
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<paragraph id="717036EF0B64FFC7FF25FE4FD9583A9E" blockId="12.[151,1437,356,1067]" pageId="12" pageNumber="77">
|
||
<emphasis id="43BBEAFD0B64FFC7FF25FE4FDB7A3AB2" bold="true" box="[199,322,428,454]" pageId="12" pageNumber="77">Diagnosis.</emphasis>
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Small species (TL
|
||
<quantity id="B6379B0A0B64FFC7FDF8FE4ED84C3AB3" box="[538,628,429,455]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="77" unit="mm" value="1.8">1.8 mm</quantity>
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), body widest in the middle of elytra, trigonium of penis slightly shorter than parameroids, parameroids with denticles on outer margins.
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="39D565640B64FFC7FF25FE17DBA638CA" pageId="12" pageNumber="77" type="description">
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<paragraph id="717036EF0B64FFC7FF25FE17DB6239B6" blockId="12.[151,1437,356,1067]" pageId="12" pageNumber="77">
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<emphasis id="43BBEAFD0B64FFC7FF25FE17DB4D397A" bold="true" box="[199,373,500,526]" pageId="12" pageNumber="77">Redescription.</emphasis>
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||
Body very small, TL
|
||
<quantity id="B6379B0A0B64FFC7FD91FE16D8F5397B" box="[627,717,501,527]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="77" unit="mm" value="1.8">1.8 mm</quantity>
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, TL/EW ~ 1.6, oval, light brown, legs and antennae yellowish, covered with yellowish semierect setae. Head strongly transverse, punctation sparse, punctures separated by ca. 2 diameters. Pronotum transverse, anterolateral and posterolateral angles right angled, lateral carinae almost straight, punctation sparse, similar to that on head. Angle between pronotum and elytra not marked in dorsal view. Scutellar shield subtriangular, impunctate. Elytra elongate, punctation stronger than on pronotum, punctures separated by ca. 0.7–1.0 diameter.
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</paragraph>
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<paragraph id="717036EF0B64FFC7FF25FD2FD97638EE" blockId="12.[151,1437,356,1067]" pageId="12" pageNumber="77">
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Penis (L
|
||
<quantity id="B6379B0A0B64FFC7FEC8FD2EDBB73993" box="[298,399,717,743]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="12" pageNumber="77" unit="mm" value="0.49">0.49 mm</quantity>
|
||
,
|
||
<geoCoordinate id="14FB50280B64FFC7FE7BFD2EDBD23993" box="[409,490,717,743]" degrees="0.12" direction="west" orientation="longitude" pageId="12" pageNumber="77" precision="555" value="-0.12">
|
||
W
|
||
<quantity id="B6379B0A0B64FFC7FE5BFD2EDBD23993" box="[441,490,717,743]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="12" pageNumber="77" unit="mm" value="0.12">0.12</quantity>
|
||
</geoCoordinate>
|
||
mm) symmetrical, widest in the middle of its length, pala and parameroids of similar length, trigonium shorter than parameroids, pointed at apex, parameroids with row of denticles on outer margins; tegmen (L
|
||
<quantity id="B6379B0A0B64FFC7FEF4FCF7DB46385B" box="[278,382,788,815]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="12" pageNumber="77" unit="mm" value="0.45">0.45 mm</quantity>
|
||
,
|
||
<geoCoordinate id="14FB50280B64FFC7FE69FCF6DBE7385B" box="[395,479,789,815]" degrees="0.28" direction="west" orientation="longitude" pageId="12" pageNumber="77" precision="555" value="-0.28">
|
||
W
|
||
<quantity id="B6379B0A0B64FFC7FE4CFCF6DBE7385B" box="[430,479,789,815]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="12" pageNumber="77" unit="mm" value="0.28">0.28</quantity>
|
||
</geoCoordinate>
|
||
mm) u-shaped, with very narrow parameres; sternite IX (L
|
||
<quantity id="B6379B0A0B64FFC7FB76FCF7DEC5385B" box="[1172,1277,788,815]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="12" pageNumber="77" unit="mm" value="0.45">0.45 mm</quantity>
|
||
,
|
||
<geoCoordinate id="14FB50280B64FFC7FAE8FCF6DF66385A" box="[1290,1374,788,815]" degrees="0.26" direction="west" orientation="longitude" pageId="12" pageNumber="77" precision="555" value="-0.26">
|
||
W
|
||
<quantity id="B6379B0A0B64FFC7FACFFCF7DF66385A" box="[1325,1374,788,814]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="12" pageNumber="77" unit="mm" value="0.26">0.26</quantity>
|
||
</geoCoordinate>
|
||
mm) consisting of two elongate and relatively wide hemisternites, apices subtriangular; tergite VIII missing; tergite IX (L
|
||
<quantity id="B6379B0A0B64FFC7FF75FCBEDAC53803" box="[151,253,861,887]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="12" pageNumber="77" unit="mm" value="0.48">0.48 mm</quantity>
|
||
,
|
||
<geoCoordinate id="14FB50280B64FFC7FEE5FCBEDB613803" box="[263,345,861,887]" degrees="0.32" direction="west" orientation="longitude" pageId="12" pageNumber="77" precision="555" value="-0.32">
|
||
W
|
||
<quantity id="B6379B0A0B64FFC7FECAFCBEDB613803" box="[296,345,861,887]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="12" pageNumber="77" unit="mm" value="0.32">0.32</quantity>
|
||
</geoCoordinate>
|
||
mm) with short apical plate and much longer apodemes, anterior margin of apical plate with wide emargination and subtriangular outgrowths in lateral portions.
|
||
</paragraph>
|
||
<paragraph id="717036EF0B64FFC7FF25FC47DBA638CA" blockId="12.[151,1437,356,1067]" box="[199,414,932,958]" pageId="12" pageNumber="77">Female. Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="39D565640B64FFC7FF25FC2BDE843897" box="[199,1212,968,995]" pageId="12" pageNumber="77" type="distribution">
|
||
<paragraph id="717036EF0B64FFC7FF25FC2BDE843897" blockId="12.[151,1437,356,1067]" box="[199,1212,968,995]" pageId="12" pageNumber="77">
|
||
<emphasis id="43BBEAFD0B64FFC7FF25FC2BDB593896" bold="true" box="[199,353,968,994]" pageId="12" pageNumber="77">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="AE74884D0B64FFC7FD82FC2AD8A83897" box="[608,656,969,995]" pageId="12" pageNumber="77">type</typeStatus>
|
||
locality in Dyaul Island (
|
||
<collectingCountry id="09D8767F0B64FFC7FC4CFC2BDE883897" box="[942,1200,968,995]" name="Papua New Guinea" pageId="12" pageNumber="77">Bismarck Archipelago</collectingCountry>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="39D565640B64FFC7FF25FC0EDF353F5E" pageId="12" pageNumber="77" type="discussion">
|
||
<paragraph id="717036EF0B64FFC7FF25FC0EDF353F5E" blockId="12.[151,1437,356,1067]" pageId="12" pageNumber="77">
|
||
<emphasis id="43BBEAFD0B64FFC7FF25FC0EDB033F73" bold="true" box="[199,315,1005,1031]" pageId="12" pageNumber="77">Remarks.</emphasis>
|
||
Figure
|
||
<quantity id="B6379B0A0B64FFC7FE71FC0EDBF43F72" box="[403,460,1004,1031]" metricMagnitude="-1" metricUnit="m" metricValue="8.382" pageId="12" pageNumber="77" unit="in" value="33.0">33 in</quantity>
|
||
the original description shows the tegmen, not sternite IX, and figure 34 is sternite IX, not the tegmen. The
|
||
<typeStatus id="AE74884D0B64FFC7FE9CFBF2DB963F5F" box="[382,430,1041,1067]" pageId="12" pageNumber="77">type</typeStatus>
|
||
specimen is in poor condition, it is disarticulated, with right elytron missing.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |