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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1131.95038" ID-GBIF-Dataset="46e05d1b-511b-4d8e-8b1e-0c733d66f878" ID-PMC="PMC9836511" ID-Pensoft-Pub="1313-2970-1131-99" ID-Pensoft-UUID="1ACF6E07EE8A54DB9906143C0532304D" ID-PubMed="36761460" ID-ZooBank="EE89E8CF4B764FBC821A79BC51D28D67" ModsDocID="1313-2970-1131-99" checkinTime="1669182088444" checkinUser="pensoft" docAuthor="Xia, Mei-Hua, Tang, Liang &amp; Schillhammer, Harald" docDate="2022" docId="F57127A3116356D6B3CCF8B8FDAB0E80" docLanguage="en" docName="ZooKeys 1131: 99-134" docOrigin="ZooKeys 1131" docPubDate="2022-11-22" docSource="http://dx.doi.org/10.3897/zookeys.1131.95038" docTitle="Thoracostrongylus acerosus Yang, Zhou &amp; Schillhammer 2011" docType="treatment" docVersion="4" id="1ACF6E07EE8A54DB9906143C0532304D" lastPageNumber="99" masterDocId="1ACF6E07EE8A54DB9906143C0532304D" masterDocTitle="Review of Chinese species of the genus Thoracostrongylus Bernhauer, 1915 (Coleoptera, Staphylinidae, Staphylininae)" masterLastPageNumber="134" masterPageNumber="99" pageNumber="99" updateTime="1732489169057" updateUser="ExternalLinkService">
<mods:mods id="45EC2B028A72F079FC2E610AB3627AC5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="B1669DB45FA7E59B6797DB2EAE2BF5EF">Review of Chinese species of the genus Thoracostrongylus Bernhauer, 1915 (Coleoptera, Staphylinidae, Staphylininae)</mods:title>
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<mods:roleTerm id="0C45E6AD7B1C147F782B1F94FA0D89EB">Author</mods:roleTerm>
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<mods:namePart id="69094A858C16B3AF6F04E28BBD139944">Xia, Mei-Hua</mods:namePart>
<mods:affiliation id="878D9BBD14CCD04D0E5C16451EE75089">College of Life Sciences, Shanghai Normal University, 100 Guilin Road, 1</mods:affiliation>
</mods:name>
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<mods:namePart id="926A0380A39F407649C4EC2E2EB881E1">Tang, Liang</mods:namePart>
<mods:affiliation id="FC9691DE33333E53E6294E2F4349D10B">College of Life Sciences, Shanghai Normal University, 100 Guilin Road, 1</mods:affiliation>
<mods:nameIdentifier id="9A1C4560CD0B4533F5CFE3CBE58238DF" type="email">staphylinidae@shnu.edu.cn</mods:nameIdentifier>
</mods:name>
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</mods:role>
<mods:namePart id="C95AD8E68A79FB7D4A718F0579B78934">Schillhammer, Harald</mods:namePart>
<mods:affiliation id="0894E8C1461F79E7BE45F4B94CECD137">st</mods:affiliation>
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<mods:typeOfResource id="2AE1CAF70686CDF14D46EF45FCE7F768">text</mods:typeOfResource>
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<mods:title id="1560F3E24D6AFD851D8231E33BDBB997">ZooKeys</mods:title>
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<mods:part id="1D2AB15EDDD63EF6734ECAA7C549A10B">
<mods:date id="6C0C3E85AB2D9ED8A2A1376659F000E1">2022</mods:date>
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<mods:number id="981D72686239CE196401069D8837E887">2022-11-22</mods:number>
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<mods:number id="C93B8A0CE7FED2D14BF78AFE92F0CBB1">1131</mods:number>
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<mods:start id="D8D04C771CC21114481AB85AD4A2BC6B">99</mods:start>
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<mods:classification id="F1EFFDDE35B379192C75F7E42D5C7041">journal article</mods:classification>
<mods:identifier id="C6C6F530A696C1EABADC09F8EE28DE7A" type="DOI">http://dx.doi.org/10.3897/zookeys.1131.95038</mods:identifier>
<mods:identifier id="275C37816C2D1E0A5E828BA9B9F3977B" type="Pensoft-Pub">1313-2970-1131-99</mods:identifier>
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<treatment id="F57127A3116356D6B3CCF8B8FDAB0E80" ID-GBIF-Taxon="205119581" LSID="urn:lsid:plazi:treatment:F57127A3116356D6B3CCF8B8FDAB0E80" httpUri="http://treatment.plazi.org/id/F57127A3116356D6B3CCF8B8FDAB0E80" lastPageNumber="99" pageId="0" pageNumber="99" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Staphylininae">
<subSubSection id="01CBA0C486076ED995362F0E4A2763B1" pageId="0" pageNumber="99" type="nomenclature">
<paragraph id="37EC7BADEB43AC5CEF96E50D0C11E5DD" pageId="0" pageNumber="99">
<taxonomicName id="57B2B16A7AB00DA3264267526D65434B" ID-CoL="56KH3" LSID="F57127A3-1163-56D6-B3CC-F8B8FDAB0E80" authority="Yang, Zhou &amp; Schillhammer, 2011" authorityName="Yang, Zhou &amp; Schillhammer" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thoracostrongylus acerosus" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="acerosus">Thoracostrongylus acerosus Yang, Zhou &amp; Schillhammer, 2011</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F2EB599044A76871DB2DE0A4A0B197F5" pageId="0" pageNumber="99" type="description">
<paragraph id="45FFC20F00B44FB97A5D56E8BCAEFE91" pageId="0" pageNumber="99">
<figureCitation id="975D64FFB59501B380CBD77FDAB8215B" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Thoracostrongylus acerosus 1, 2 habitus 3 - 6 aedeagus, ventral (3, 4) and lateral (5, 6) views. Scale bars: 1 mm (1, 2); 0.2 mm (3 - 6)." figureDoi="10.3897/zookeys.1131.95038.figures1-6" httpUri="https://binary.pensoft.net/fig/772737" pageId="0" pageNumber="99">Figs 1-6</figureCitation>
<figureCitation id="582B4F4E4A494683CDF56EB2C282AA42" captionStart="Figures 7–14" captionStartId="F2" captionText="Figures 7 - 14. Thoracostrongylus acerosus 7 - 10 aedeagus from Xiling Snow Mountain, ventral (7, 8) and lateral (9, 10) views 11 - 14 aedeagus from Jiulongdong, ventral (11, 12) and lateral (13, 14) views. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1131.95038.figures7-14" httpUri="https://binary.pensoft.net/fig/772738" pageId="0" pageNumber="99">, 7-14</figureCitation>
<figureCitation id="8CF1AFDDEE1D25E9078EDB2BDEF65308" captionStart="Figures 108–122" captionStartId="F18" captionText="Figures 108 - 122. Distribution map of Thoracostrongylus species of China 108 T. acerosus 109 T. aduncatus 110 T. birmanus 111 T. diaoluoensis 112 T. formosanus formosanus 113 T. formosanus flavipes 114 T. fujianensis 115 T. malaisei 116 T. miyakei 117 T. sarawakensis 118 T. velutinus 119 T. baishanzuensis 120 T. bicolor 121 T. brachypterus 122 T. chrysites. Triangle, localities of specimens examined in this paper; square, localities of specimens listed in previous papers. Red, doubtful localities; blue, trustede localities." figureDoi="10.3897/zookeys.1131.95038.figures108-122" httpUri="https://binary.pensoft.net/fig/772754" pageId="0" pageNumber="99">
, 108
<normalizedToken id="AE6D256A00E201A167EF6DBB2E98B49E" originalValue="刺茎钝胸隐翅虫">刺茎钝胸隐翅虫</normalizedToken>
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="37417C1570087489845FF8CED755EE90" pageId="0" pageNumber="99" type="reference_group">
<paragraph id="CE688ECC18F8FF038F6C8E08FA94C5A5" pageId="0" pageNumber="99">
<taxonomicName id="02EDBA87DE72D88E1FAA5126E74EB312" authorityName="Yang, Zhou &amp; Schillhammer" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thoracostrongylus acerosus" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="acerosus">Thoracostrongylus acerosus</taxonomicName>
Yang, Zhou &amp; Schillhammer, 2011: 410.
</paragraph>
<paragraph id="551445C6644A43F3ECE993445E6928F7" pageId="0" pageNumber="99">
<taxonomicName id="F9197D18D59CD29E2E3BBA464D4059B9" authorityName="Yang, Zhuo, Hong-Zhang Zhou &amp; Schillhammer" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thoracostrongylus baoxingensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="baoxingensis">Thoracostrongylus baoxingensis</taxonomicName>
Yang, Zhou &amp; Schillhammer, 2011: 415. syn. nov.
</paragraph>
</subSubSection>
<subSubSection id="257C22A4A0F5ACF129E2A7063996B452" pageId="0" pageNumber="99" type="materials_examined">
<paragraph id="43509BA4C20F8EF9BD483B627079C5F6" pageId="0" pageNumber="99">Material examined.</paragraph>
<paragraph id="CE4FDC3D64B16B02F285B77A47937C5F" pageId="0" pageNumber="99">
<collectingCountry id="8BDD731FCAAFDF8998FA52155645DA0D" name="China">China</collectingCountry>
-
<emphasis id="040E940A2BD95583380BC422DC239924" bold="true" pageId="0" pageNumber="99">
<collectingRegion id="8EE0E96845EDF82FC884B732AD4380B8" country="China" name="Sichuan">Sichuan Prov.</collectingRegion>
</emphasis>
•
<specimenCount id="EF927F928E6A41CA47B6F45A445FDF65" type="male">2♂♂</specimenCount>
,
<specimenCount id="B9625DA91DB771423B39C2B8FE12949B" type="female">1♀</specimenCount>
; Baoxing County, Fengtongzhai;
<geoCoordinate id="AF2A9C4DAA11FFFEB11CEE36C4B8276C" degrees="30" direction="north" minutes="32" orientation="latitude" precision="15" seconds="10" value="30.53611">30°32'10&quot;N</geoCoordinate>
,
<geoCoordinate id="1F9E11DE108A1BF153F97827332EDBD5" degrees="102" direction="east" minutes="54" orientation="longitude" precision="15" seconds="20" value="102.905556">102°54'20&quot;E</geoCoordinate>
; alt.
<quantity id="93A5E534417A492DDF42EA96FE062D47" metricMagnitude="3" metricUnit="m" metricValue="1.49" unit="m" value="1490.0">1490 m</quantity>
;
<date id="FC789079E5141126DFF780827D9BB772" value="2015-07-22">22 July 2015</date>
; Jiang, Peng, Tu &amp; Zhou leg.; SHNU •
<specimenCount id="6DA5A4CEA7604956670E1633ADE2E7EC" type="male">1♂</specimenCount>
,
<specimenCount id="F1DC6797F065D2231535A7392A61D43C" type="female">1♀</specimenCount>
; Baoxing County, Fengtongzhai N.R., Dengchigou;
<geoCoordinate id="2D014AE6C7464050805C171D6B62ACE4" degrees="30" direction="north" minutes="32" orientation="latitude" precision="925" value="30.533333">30°32'N</geoCoordinate>
,
<geoCoordinate id="BA371319719BBE5032D7AF76B4D20CD6" degrees="102" direction="east" minutes="56" orientation="longitude" precision="925" value="102.933334">102°56'E</geoCoordinate>
; alt.
<quantity id="3C9BB5D35D50EC912FE7C3036475656E" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" unit="m" value="1870.0">1870 m</quantity>
;
<date id="BB2F6AA2DAB3F4A566E1AA4361942717" value="2016-08-01">01 August 2016</date>
; Zhou, Jiang, Liu &amp; Gao leg.; SHNU •
<specimenCount id="1C08F0625F374520799C16499CE54E6F" type="female">1♀</specimenCount>
; Baoxing County, Fengtongzhai N.R., Dengchigou;
<geoCoordinate id="F4F7AE230857B21BA563232749C8E015" degrees="30" direction="north" minutes="29" orientation="latitude" precision="925" value="30.483334">30°29'N</geoCoordinate>
,
<geoCoordinate id="36AD3D7BFF2B4BA439084BE1CD1985D1" degrees="102" direction="east" minutes="51" orientation="longitude" precision="925" value="102.85">102°51'E</geoCoordinate>
; alt.
<quantity id="C3E5B4EFE73D632C7727D52446A23FBF" metricMagnitude="3" metricUnit="m" metricValue="1.6919999999999997" unit="m" value="1692.0">1692 m</quantity>
;
<date id="8A0D64491A792CAAF4D55067E31CED27" value="2016-08-02">02 August 2016</date>
; Zhou, Jiang, Liu &amp; Gao leg.; SHNU •
<specimenCount id="9190884B3D8ABFF79955F7ADB9C48069" type="female">2♀♀</specimenCount>
; Tianquan County, Liangluxiang Village;
<geoCoordinate id="30B03F8CE63719741B01A5F4D6DB9396" degrees="29" direction="north" minutes="56" orientation="latitude" precision="925" value="29.933332">29°56'N</geoCoordinate>
,
<geoCoordinate id="7DC23978F97EAA403B7356EE32E9DB3F" degrees="102" direction="east" minutes="23" orientation="longitude" precision="925" value="102.38333">102°23'E</geoCoordinate>
; alt.
<quantity id="A2FE1CA6B2265DCA0AB4B646D5DF2CFF" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">1500-1700 m</quantity>
;
<date id="7DC13997AFC1E36FD079834FEA158AEA" value="2012-07-10">10 July 2012</date>
; Peng, Dai &amp; Yin leg.; SHNU •
<specimenCount id="34EEAE74E76016A8AE6C240B4748266A" type="female">1♀</specimenCount>
; Tianquan County, Liangluxiang;
<geoCoordinate id="4CDFC9EAD428E94B29DEFBEC34561603" degrees="29" direction="north" minutes="56" orientation="latitude" precision="925" value="29.933332">29°56'N</geoCoordinate>
,
<geoCoordinate id="7BDFF8AF89CE50EAF1E7552111087A6D" degrees="102" direction="east" minutes="23" orientation="longitude" precision="925" value="102.38333">102°23'E</geoCoordinate>
; alt.
<quantity id="FE7E8C8A108E15310F38C274470F0851" metricMagnitude="3" metricUnit="m" metricValue="1.95" metricValueMax="2.0" metricValueMin="1.9" unit="m" value="1950.0" valueMax="2000.0" valueMin="1900.0">1900-2000 m</quantity>
; Peng, Dai &amp; Yin leg.; SHNU •
<specimenCount id="715EFD13B03EBFF1E7A3A8FD19527076" type="female">5♀♀</specimenCount>
; Tianquan County, Lianglu County; alt.
<quantity id="A2C022046AB2D1D1A17ABDD56A2E4076" metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">1400 m</quantity>
;
<date id="2ACA7B7B11DFBED7934DF4D71D857236" value="2011-08-01">01 August 2011</date>
; Hao Huang leg.; SHNU •
<specimenCount id="B5E06EFDB3BB6E4449853DFA981494A6" type="male">2♂♂</specimenCount>
,
<specimenCount id="7B3995B87AE026E1E4AB9331A0623296" type="female">1♀</specimenCount>
; Dayi County, Xiling Snow Mt.;
<geoCoordinate id="226E941CBC336DA8212275029121967D" degrees="30" direction="north" minutes="38" orientation="latitude" precision="1" seconds="6.25" value="30.635069">30°38'6.25&quot;N</geoCoordinate>
, 103°10'99.08&quot;E; alt.
<quantity id="4A252372673680ECBF786DB01297B8AD" metricMagnitude="3" metricUnit="m" metricValue="1.25" unit="m" value="1250.0">1250 m</quantity>
;
<date id="4ED08EDF0C5C10BD4CBEC22602A166C3" value="2021-07-31">31 July 2021</date>
; Zhao &amp; Cai leg.; SHNU. -
<emphasis id="4F11553124CA61B719781D7504EA0ED2" bold="true" pageId="0" pageNumber="99">
<collectingRegion id="425B9994101B72A28645DDEE502DFB73" country="China" name="Shaanxi">Shaanxi Prov.</collectingRegion>
</emphasis>
•
<specimenCount id="42CE953C5C058FD48B1B29340C79C354" type="female">1♀</specimenCount>
; Hanzhong, Tiantaishan;
<geoCoordinate id="FC103A8DC89F58B3CDC5035224EEE315" degrees="33" direction="north" minutes="16" orientation="latitude" precision="15" seconds="20" value="33.27222">33°16'20&quot;N</geoCoordinate>
,
<geoCoordinate id="A1680D3EBAD43700AAF51CCAF1C6C0E5" degrees="107" direction="east" minutes="04" orientation="longitude" precision="15" seconds="52" value="107.08111">107°04'52&quot;E</geoCoordinate>
; alt.
<quantity id="8E489293D27EF5D9D6481CC2066254F2" metricMagnitude="3" metricUnit="m" metricValue="1.326" unit="m" value="1326.0">1326 m</quantity>
;
<date id="A185A5EBBB20263E868EE8223D3E3D96" value="2021-05-08">08 May 2021</date>
; Juan Li et al. leg.; SHNU •
<specimenCount id="168867E4BF26BCE3AD1CA1807C489480" type="female">2♀♀</specimenCount>
; Liuba, Huoshaodian;
<geoCoordinate id="592D0432C378C75BA17F0A969E2BE637" degrees="33" direction="north" minutes="30" orientation="latitude" precision="15" seconds="08" value="33.502224">33°30'08&quot;N</geoCoordinate>
,
<geoCoordinate id="667742B5C01ADA5375219C8CB22359C9" degrees="106" direction="east" minutes="56" orientation="longitude" precision="15" seconds="08" value="106.935555">106°56'08&quot;E</geoCoordinate>
; alt.
<quantity id="3407691A6472DC6A229A0D810D92E786" metricMagnitude="3" metricUnit="m" metricValue="1.041" unit="m" value="1041.0">1041 m</quantity>
;
<date id="744D68CF7B93E910E1E7B59FAFBF142F" value="2021-07-08">08 July 2021</date>
; Juan Li et al. leg.; SHNU •
<specimenCount id="34080250FCD0FAB70BE8A14F048B3885" type="female">2♀♀</specimenCount>
; Zhouzhi Coun., Houzhenzi, Qinling, west Sangongli Gou;
<geoCoordinate id="8F3CC026950A12C1BA602D651B258801" degrees="33" direction="north" minutes="50" orientation="latitude" precision="1" seconds="6.13" value="33.835033">33°50'6.13&quot;N</geoCoordinate>
,
<geoCoordinate id="38C14A1053C51588ADE99DDA6EE5DB93" degrees="107" direction="east" minutes="48" orientation="longitude" precision="1" seconds="52.4" value="107.81456">107°48'52.4&quot;E</geoCoordinate>
; alt.
<quantity id="0316F5EE0FE4B2553D322ECA853072A1" metricMagnitude="3" metricUnit="m" metricValue="1.3359999999999999" unit="m" value="1336.0">1336 m</quantity>
;
<date id="C9F678D71FCD9D2CEA8317AE28D2E470" value="2008-05-17" valueMax="2008-05-19" valueMin="2008-05-17">17-19 May 2008</date>
; Huang &amp; Xu leg.; SHNU •
<specimenCount id="B7F36C206235F6CB7F16713B4453E90F" type="female">1♀</specimenCount>
; Zhouzhi Coun., Houzhenzi, Qinling;
<geoCoordinate id="AA7878D4AA70034B3C0A354E0781D892" degrees="33" direction="north" minutes="51" orientation="latitude" precision="1" seconds="20.3" value="33.855637">33°51'20.3&quot;N</geoCoordinate>
,
<geoCoordinate id="DC3C9721D10097DAC954F1D886CFC867" degrees="107" direction="east" minutes="50" orientation="longitude" precision="1" seconds="18.3" value="107.83842">107°50'18.3&quot;E</geoCoordinate>
; alt.
<quantity id="7A18CCFCEED6D59DAE450E178C06DA71" metricMagnitude="3" metricUnit="m" metricValue="1.26" unit="m" value="1260.0">1260 m</quantity>
;
<date id="6190F2D6378A1B475E4C52A92040A35D" value="2008-05-05" valueMax="2008-05-10" valueMin="2008-05-05">05-10 May 2008</date>
; Huang &amp; Xu leg.; SHNU •
<specimenCount id="5015C7DB00F929E81FA87D9300C2142D" type="male">1♂</specimenCount>
; Baoji City, Jiulongdong;
<geoCoordinate id="2EFC89B74D1A3A74AB8D72C876EBA847" degrees="34" direction="north" minutes="19" orientation="latitude" precision="15" seconds="56" value="34.332222">34°19'56&quot;N</geoCoordinate>
,
<geoCoordinate id="71473A2D51C2E11028973208AE50D719" degrees="106" direction="east" minutes="52" orientation="longitude" precision="15" seconds="22" value="106.87278">106°52'22&quot;E</geoCoordinate>
; alt.
<quantity id="03FA9ABD1599EE3D2323D435FCD331B6" metricMagnitude="2" metricUnit="m" metricValue="9.86" unit="m" value="986.0">986 m</quantity>
;
<date id="77FBE882DF71C22056D604AE28160A99" value="2021-05-26">26 May 2021</date>
; Juan Li et al. leg.; SHNU •
<specimenCount id="3590916155C09A3FCA15BC88CF142FD1" type="female">1♀</specimenCount>
; Baoji City, Jiulongdong;
<geoCoordinate id="9B4CE4AE3460E84C3EEB31EDD611F2F7" degrees="34" direction="north" minutes="19" orientation="latitude" precision="15" seconds="59" value="34.333054">34°19'59&quot;N</geoCoordinate>
,
<geoCoordinate id="C03CEDF26CC4E8BA71C89E51F10171B5" degrees="106" direction="east" minutes="52" orientation="longitude" precision="15" seconds="21" value="106.872505">106°52'21&quot;E</geoCoordinate>
; alt.
<quantity id="5844A167341CAAA2A7FA381CE28F472A" metricMagnitude="2" metricUnit="m" metricValue="9.75" unit="m" value="975.0">975 m</quantity>
;
<date id="66C0F9FD2AC881698BB3470B136FC409" value="2021-08-05">05 August 2021</date>
; Juan Li et al. leg.; SHNU •
<specimenCount id="F2770FBA7CA9E1590EBAC8778544FA84" type="female">1♀</specimenCount>
; Baoji City, Jiulongdong;
<geoCoordinate id="CD2E93E4DA1621A9513E21140ADFC314" degrees="34" direction="north" minutes="20" orientation="latitude" precision="15" seconds="10" value="34.33611">34°20'10&quot;N</geoCoordinate>
,
<geoCoordinate id="CFE1E0B492F4489D7CB2D35F46DD368B" degrees="106" direction="east" minutes="51" orientation="longitude" precision="15" seconds="51" value="106.86417">106°51'51&quot;E</geoCoordinate>
; alt.
<quantity id="822DAB08A93A6DBAFDB16CC95393F519" metricMagnitude="2" metricUnit="m" metricValue="9.69" unit="m" value="969.0">969 m</quantity>
;
<date id="2E3792CD3D685A7B7EDAA23EEA21ECC2" value="2021-08-05">05 August 2021</date>
; Juan Li et al. leg.; SHNU •
<specimenCount id="C070142D9F24E18856872DF721E5838E" type="male">1♂</specimenCount>
; Liuba, Zhangliang Temple;
<geoCoordinate id="AA00C54C5B1F6C146C80C6375D0071D2" degrees="33" direction="north" minutes="41" orientation="latitude" precision="15" seconds="51" value="33.697502">33°41'51&quot;N</geoCoordinate>
,
<geoCoordinate id="47CD2564F5701F5765D71654364A009F" degrees="106" direction="east" minutes="47" orientation="longitude" precision="15" seconds="15" value="106.7875">106°47'15&quot;E</geoCoordinate>
; alt.
<quantity id="E4A8A8FCAFDBDCA562161025A8A8B7DD" metricMagnitude="3" metricUnit="m" metricValue="1.476" unit="m" value="1476.0">1476 m</quantity>
;
<date id="4E61D6B1B1D99564711417FDB2E2A124" value="2021-07-11">11 July 2021</date>
; Juan Li et al. leg.; SHNU •
<specimenCount id="A04AA5D882D7BD7A37E106869BF1AFEB" type="female">1♀</specimenCount>
; Lueyang, Wulongdong;
<geoCoordinate id="AB5AECAFED0D546399FC6030BFA26930" degrees="33" direction="north" minutes="31" orientation="latitude" precision="15" seconds="16" value="33.52111">33°31'16&quot;N</geoCoordinate>
,
<geoCoordinate id="5F9EF9D81D022035988B32D23BDD3C9A" degrees="106" direction="east" minutes="16" orientation="longitude" precision="15" seconds="22" value="106.27278">106°16'22&quot;E</geoCoordinate>
; alt.
<quantity id="73BB0E809A34314CE6EA0A5ED5713B1C" metricMagnitude="3" metricUnit="m" metricValue="1.107" unit="m" value="1107.0">1107 m</quantity>
;
<date id="1C6766E589356A286F83212E824ABF58" value="2021-07-20">20 July 2021</date>
; Juan Li et al. leg.; SHNU •
<specimenCount id="A4992654A58C7419B5FD79E393B874C2" type="female">1♀</specimenCount>
; Lueyang, Wulongdong;
<geoCoordinate id="3126C95131C02506EA5ED0CB557DCD01" degrees="33" direction="north" minutes="30" orientation="latitude" precision="15" seconds="51" value="33.514168">33°30'51&quot;N</geoCoordinate>
,
<geoCoordinate id="5C224F1F9C24853BE13B2855F0E38F24" degrees="106" direction="east" minutes="15" orientation="longitude" precision="15" seconds="04" value="106.251114">106°15'04&quot;E</geoCoordinate>
; alt.
<quantity id="45CD315BA38AD676C28D1E4E3F93E0F8" metricMagnitude="3" metricUnit="m" metricValue="1.237" unit="m" value="1237.0">1237 m</quantity>
;
<date id="7A30EF7B93FBFE8A1950A5DEA102F497" value="2021-07-20">20 July 2021</date>
; Juan Li et al. leg.; SHNU •
<specimenCount id="4BCFD1091E4AF9F76B3129852E2CAB3F" type="male">2♂♂</specimenCount>
; Ankang City, Ningshan, Guanghuojie Town; 33°45'81&quot;N,
<geoCoordinate id="DB5940DA8481D868D8D523E029B111BD" degrees="108" direction="east" minutes="46" orientation="longitude" precision="15" seconds="48" value="108.78001">108°46'48&quot;E</geoCoordinate>
; alt.
<quantity id="91CBA0D2BD291914DB3EA4BDD29BC1A0" metricMagnitude="3" metricUnit="m" metricValue="1.176" unit="m" value="1176.0">1176 m</quantity>
;
<date id="062E293316D1F128851AD515FF19AF90" value="2011-05-07" valueMax="2011-05-08" valueMin="2011-05-07">07-08 May 2011</date>
; Bao-Xiang Zhan leg.; SHNU. -
<emphasis id="E945643F2C40420E94214FC7FF17D373" bold="true" pageId="0" pageNumber="99">
<collectingRegion id="7008E41D543E04CE178F1ED58E5ADBDE" country="China" name="Gansu">Gansu Prov.</collectingRegion>
</emphasis>
•
<specimenCount id="305D343B901AC8C36D0ADAAC7C253507" type="male">1♂</specimenCount>
; Hui County, Gaoqiaolinchang;
<geoCoordinate id="636A1BB7084C68F9D956CDFC4B8FB938" degrees="34" direction="north" minutes="05" orientation="latitude" precision="15" seconds="44" value="34.095554">34°05'44&quot;N</geoCoordinate>
,
<geoCoordinate id="706768B88906C9213A5282FE84B8AAFE" degrees="105" direction="east" minutes="57" orientation="longitude" precision="15" seconds="42" value="105.96166">105°57'42&quot;E</geoCoordinate>
; alt.
<quantity id="29437063DF8DA9D82C498DA8369144E2" metricMagnitude="3" metricUnit="m" metricValue="1.305" unit="m" value="1305.0">1305 m</quantity>
;
<date id="67AFC928A58A9ECA8403400D6DE43E2D" value="2021-05-18">18 May 2021</date>
; Juan Li et al. leg.; SHNU •
<specimenCount id="C7D36805D725A4AD0F17ECFEAF71239F" type="female">1♀</specimenCount>
; Hui County, Yanpinglinchang;
<geoCoordinate id="5156F246843AADEB25C7070F66F860EE" degrees="33" direction="north" minutes="40" orientation="latitude" precision="15" seconds="36" value="33.676666">33°40'36&quot;N</geoCoordinate>
,
<geoCoordinate id="505A3AFEF56682DDF368DD21235A475A" degrees="106" direction="east" minutes="16" orientation="longitude" precision="15" seconds="51" value="106.28084">106°16'51&quot;E</geoCoordinate>
; alt.
<quantity id="7FD5E90DD4DF41618A8D4BD9FFA042B0" metricMagnitude="3" metricUnit="m" metricValue="1.483" unit="m" value="1483.0">1483 m</quantity>
;
<date id="D103267DE4B973F11DFD48DF5785FC96" value="2021-07-18">18 July 2021</date>
; Juan Li et al. leg.; SHNU. -
<emphasis id="27D9E973BB365CD4CB36F5C5B02D4FA1" bold="true" pageId="0" pageNumber="99">
<collectingRegion id="8C5DAA006CE3144916203F9EBB05B5D7" country="China" name="Henan">Henan Prov.</collectingRegion>
</emphasis>
•
<specimenCount id="641454BBBD13C6A487BD4CC207BDB87B" type="male">1♂</specimenCount>
; Funiu Shan, Baotianman; alt.
<quantity id="3C8D2E2AA7C8EE1EE34860D5D63D6523" metricMagnitude="3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" unit="m" value="1600.0" valueMax="1700.0" valueMin="1500.0">1500-1700 m</quantity>
;
<geoCoordinate id="E079D402507099031674D1ABDBEC266E" degrees="33" direction="north" minutes="31" orientation="latitude" precision="925" value="33.516666">33°31'N</geoCoordinate>
,
<geoCoordinate id="10E8D926E43639FC015D50448583F74A" degrees="111" direction="east" minutes="56" orientation="longitude" precision="925" value="111.933334">111°56'E</geoCoordinate>
;
<date id="A380D0998214606475E3E05A6745557C" value="2009-06-15">15 June 2009</date>
; J. Turna leg.; NMW.
</paragraph>
<caption id="AD1E19A2D09446A8F51D1C9C5AB2488E" doi="10.3897/zookeys.1131.95038.figures1-6" httpUri="https://binary.pensoft.net/fig/772737" pageId="0" pageNumber="99" start="Figures 1–6" startId="F1">
<paragraph id="E5545157748A56690D901AE09B2364FF" pageId="0" pageNumber="99">
<emphasis id="287C57FA6D92C1A0630CE9B0F8959B6D" bold="true" pageId="0" pageNumber="99">Figures 1-6.</emphasis>
<taxonomicName id="9F675585954AA15FCF8C4CFBD611122C" authorityName="Yang, Zhou &amp; Schillhammer" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thoracostrongylus acerosus" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="acerosus">
<emphasis id="534A19BCE924B4F29B14C5745DB7EF35" italics="true" pageId="0" pageNumber="99">Thoracostrongylus acerosus</emphasis>
</taxonomicName>
<emphasis id="30B1396CCDD46AC99CB214BB0F429889" bold="true" pageId="0" pageNumber="99">1, 2</emphasis>
habitus
<emphasis id="E704F935497BAAD967810025E091956F" bold="true" pageId="0" pageNumber="99">3-6</emphasis>
aedeagus, ventral (
<emphasis id="56341EDF3D665B36AFBB9762CABEDCD1" bold="true" pageId="0" pageNumber="99">3, 4</emphasis>
) and lateral (
<emphasis id="8DF39B2C8C05A12F3E805CAB9DEE0FF2" bold="true" pageId="0" pageNumber="99">5, 6</emphasis>
) views. Scale bars:
<quantity id="5BA6F1042645F881BC2CC077655B8ABB" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
(
<emphasis id="BEEEA59EE392DC663C13670049D79B74" bold="true" pageId="0" pageNumber="99">1, 2</emphasis>
);
<quantity id="5CADBF27D1B3CD68CD5F9558906FCE95" metricMagnitude="-4" metricUnit="m" metricValue="2.0" unit="mm" value="0.2">0.2 mm</quantity>
(
<emphasis id="DAA2468AAAA63D9BE7622FF1BA5FABC0" bold="true" pageId="0" pageNumber="99">3-6</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="B89A79D9180F490A0E8C6985A49708D2" pageId="0" pageNumber="99" type="description">
<paragraph id="C7CA498B823CFCD0D723F0BC65AF5096" pageId="0" pageNumber="99">Measurements.</paragraph>
<paragraph id="C4D9641B4F0C1B3DBB2C2A9BF5B90245" pageId="0" pageNumber="99">
<emphasis id="6883405790E188067E1A3AD8983DF067" bold="true" pageId="0" pageNumber="99">Male</emphasis>
: BL: 8.2-9.7 mm, FL: 4.5-5.2 mm. HL: 1.28-1.45 mm, HW: 1.78-1.95 mm, CL: 0.89-0.95 mm, PO: 0.22-0.28 mm, PL: 1.61-1.78 mm, PW: 1.45-1.50 mm, EL: 1.95-2.11 mm, EW: 1.95-2.11 mm. HL/HW: 0.70-0.78, CL/PO: 3.20-4.00, PL/PW: 1.12-1.23, EL/EW: 0.95-1.00, HW/EW: 0.87-0.94, PW/EW: 0.68-0.74, HW/PW: 1.23-1.31.
<emphasis id="837ED5524A35BEA9435A8C796502FA5E" bold="true" pageId="0" pageNumber="99">Female</emphasis>
: BL: 9.2-10.4 mm, FL: 4.8-5.3 mm. HL: 1.39-1.50 mm, HW: 2.00-2.17 mm, CL: 0.89-1.06 mm, PO: 0.22-0.28 mm, PL: 1.72-1.95 mm, PW: 1.50-1.72 mm, EL: 2.00-2.50 mm, EW: 2.11-2.50 mm. HL/HW: 0.68-0.72, CL/PO: 3.60-4.00, PL/PW: 1.00-1.19, EL/EW: 0.95-1.00, HW/EW: 0.84-0.95, PW/EW: 0.69-0.71, HW/PW: 1.23-1.33.
</paragraph>
</subSubSection>
<subSubSection id="87684A7685172798DE121600FB9361B7" pageId="0" pageNumber="99" type="distribution">
<paragraph id="0EC7C8EEE84C78DDE8359C11B456AA92" pageId="0" pageNumber="99">Distribution.</paragraph>
<paragraph id="901AC4DA37FE5E5C2D16F68C6A44B9E1" pageId="0" pageNumber="99">China (Hubei, Sichuan, Shaanxi, Gansu, Henan). New to Shaanxi, Gansu, and Henan.</paragraph>
</subSubSection>
<subSubSection id="0A71AA133C385CE27A0A05918899CF5D" pageId="0" pageNumber="99" type="diagnosis">
<paragraph id="242AD576DB8B42A8F5D794FC600B0C57" pageId="0" pageNumber="99">Diagnosis.</paragraph>
<paragraph id="E6B228E0D4A57EF7B5EFE630E75573DD" pageId="0" pageNumber="99">
In general appearance,
<taxonomicName id="18192CC45D1DDD8C8287F859A21991D9" class="Insecta" kingdom="Animalia" lsidName="T. acerosus" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="acerosus">
<emphasis id="FC69557365C16A1066BEF955F333B3CF" italics="true" pageId="0" pageNumber="99">T. acerosus</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="ABCD97DE3DFE2F13868856267AF4422B" class="Insecta" kingdom="Animalia" lsidName="T. aduncatus" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="aduncatus">
<emphasis id="77664364910DC60D15FE986AF0C4B5DD" italics="true" pageId="0" pageNumber="99">T. aduncatus</emphasis>
</taxonomicName>
Yang, Zhou &amp; Schillhammer, 2011,
<taxonomicName id="303DFAAC4E7D1CFDC84E62BAE4E73E26" class="Insecta" kingdom="Animalia" lsidName="T. fujianensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="fujianensis">
<emphasis id="B65DC1858A1B88008EC51B4DD4FC57AC" italics="true" pageId="0" pageNumber="99">T. fujianensis</emphasis>
</taxonomicName>
Yang, Zhou &amp; Schillhammer, 2011, and
<taxonomicName id="AB0D0D3FBA3A0579EA698DD6BC87853D" class="Insecta" kingdom="Animalia" lsidName="T. diaoluoensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="diaoluoensis">
<emphasis id="FF505B0023A9B6A3E786A4239081C1BC" italics="true" pageId="0" pageNumber="99">T. diaoluoensis</emphasis>
</taxonomicName>
Yang, Zhou &amp; Schillhammer, 2011, but it can be recognized by the sharply pointed tip of aedeagal median lobe.
</paragraph>
</subSubSection>
<subSubSection id="275B4F0D5D2C2D063BBC4D0EEDE63B5B" pageId="0" pageNumber="99" type="remarks">
<paragraph id="657C896CE73C1BE5407921D79CD859AF" pageId="0" pageNumber="99">Remarks.</paragraph>
<paragraph id="E5C171920AC4AD1BA458868462F199D0" pageId="0" pageNumber="99">
The apical portion of the median lobe and the paramere are subject to some variability (Figs
<figureCitation id="C98BFCE2E513E8AE248012C7CF332956" captionStart="Figures 1–6" captionStartId="F1" captionText="Figures 1 - 6. Thoracostrongylus acerosus 1, 2 habitus 3 - 6 aedeagus, ventral (3, 4) and lateral (5, 6) views. Scale bars: 1 mm (1, 2); 0.2 mm (3 - 6)." figureDoi="10.3897/zookeys.1131.95038.figures1-6" httpUri="https://binary.pensoft.net/fig/772737" pageId="0" pageNumber="99">3</figureCitation>
-
<figureCitation id="6A1451A88E04EADA576EFA89FE00F585" captionStart="Figures 7–14" captionStartId="F2" captionText="Figures 7 - 14. Thoracostrongylus acerosus 7 - 10 aedeagus from Xiling Snow Mountain, ventral (7, 8) and lateral (9, 10) views 11 - 14 aedeagus from Jiulongdong, ventral (11, 12) and lateral (13, 14) views. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1131.95038.figures7-14" httpUri="https://binary.pensoft.net/fig/772738" pageId="0" pageNumber="99">14</figureCitation>
). This may be observed not only in populations from different localities but also within one population. A closer inspection of the types of
<taxonomicName id="235C9A5F0A723C22E210559E20592387" class="Insecta" kingdom="Animalia" lsidName="T. baoxingensis" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="baoxingensis">
<emphasis id="8E0688C7156BB85EA8EE4752C3DABEB3" italics="true" pageId="0" pageNumber="99">T. baoxingensis</emphasis>
</taxonomicName>
and
<taxonomicName id="2CDCCD501ACE0D306E5E877626BC7DF6" class="Insecta" kingdom="Animalia" lsidName="T. acerosus" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="acerosus">
<emphasis id="530267979F0034406FCFDDABA81F539F" italics="true" pageId="0" pageNumber="99">T. acerosus</emphasis>
</taxonomicName>
revealed that this is the case here as well and that both species are conspecific.
</paragraph>
<caption id="9F672B50FD8AD51AB1077D79AB0E696D" doi="10.3897/zookeys.1131.95038.figures7-14" httpUri="https://binary.pensoft.net/fig/772738" pageId="0" pageNumber="99" start="Figures 7–14" startId="F2">
<paragraph id="F64AACD759F5D3BCBA76A8B0BF5D0C14" pageId="0" pageNumber="99">
<emphasis id="F7C3D9428226CD8EFC8BFA8EC460680D" bold="true" pageId="0" pageNumber="99">Figures 7-14.</emphasis>
<taxonomicName id="59D6772A9A793C58FC575333F5FB4526" authorityName="Yang, Zhou &amp; Schillhammer" authorityYear="2011" class="Insecta" family="Staphylinidae" genus="Thoracostrongylus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Thoracostrongylus acerosus" order="Coleoptera" pageId="0" pageNumber="99" phylum="Arthropoda" rank="species" species="acerosus">
<emphasis id="8E7E6B99241075583549F399E38793F5" italics="true" pageId="0" pageNumber="99">Thoracostrongylus acerosus</emphasis>
</taxonomicName>
<emphasis id="7BDA41429A10FC7C9C97C9396B339248" bold="true" pageId="0" pageNumber="99">7-10</emphasis>
aedeagus from Xiling Snow Mountain, ventral (
<emphasis id="8565EEDB565FF87EF4BEC6A457A9B789" bold="true" pageId="0" pageNumber="99">7, 8</emphasis>
) and lateral (
<emphasis id="3E1A58D76B7C17B276F13056E658AB7E" bold="true" pageId="0" pageNumber="99">9, 10</emphasis>
) views
<emphasis id="264A8A2E339836994FD5BBD440F4CA40" bold="true" pageId="0" pageNumber="99">11-14</emphasis>
aedeagus from Jiulongdong, ventral (
<emphasis id="7774787910CAB707484A353A8CD7E3A6" bold="true" pageId="0" pageNumber="99">11, 12</emphasis>
) and lateral (
<emphasis id="A5193CD1E9968804B59557AC68EFC6B6" bold="true" pageId="0" pageNumber="99">13, 14</emphasis>
) views. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
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