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<document id="8C4394932DC01DC0D3C43D84CFCCE3E3" ID-CLB-Dataset="299687" ID-DOI="10.11646/zootaxa.5476.1.14" ID-GBIF-Dataset="83be1d12-3615-41b3-9b8f-b1490f55488e" ID-ISSN="1175-5326" ID-Zenodo-Dep="12724044" ID-ZooBank="59712C82-A1A8-43F8-9B15-7FD4B43DC0E0" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1720374679655" checkinUser="plazi" docAuthor="Hsu, Jhih-Wei &amp; Shih, Hsi-Te" docDate="2024" docId="F5111D13756DC645FF3BFD34FA404BA7" docLanguage="en" docName="zootaxa.5476.1.14.pdf" docOrigin="Zootaxa 5476 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5476.1.14" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Parahelice ngankeeae Hsu &amp; Shih, 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="144" masterDocId="0928656B756EC643FFACFFB7FFDC4D46" masterDocTitle="First records of Parahelice K. Sakai, Türkay &amp; Yang, 2006 and Pseudohelice K. Sakai, Türkay &amp; Yang, 2006 from Vanuatu, with description of a new species of Parahelice (Crustacea: Brachyura: Varunidae)" masterLastPageNumber="151" masterPageNumber="138" pageNumber="141" updateTime="1720709943798" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="9D63809538047FBF9821B81E33F42A87">First records of Parahelice K. Sakai, Türkay &amp; Yang, 2006 and Pseudohelice K. Sakai, Türkay &amp; Yang, 2006 from Vanuatu, with description of a new species of Parahelice (Crustacea: Brachyura: Varunidae)</mods:title>
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<mods:name id="F241060B3C3E82556BDA0BA5C1B5A381" type="personal">
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<mods:namePart id="7B0991FD73DEFBF3BDB79DB4D73F5B01">Hsu, Jhih-Wei</mods:namePart>
<mods:affiliation id="14AC43B3245F37DD44AB1FEDF7F4429C">National Chung Hsing University, Department of Life Science, 250 Kuo Kuang Road, Taichung 402, Taiwan.</mods:affiliation>
</mods:name>
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<mods:role id="6EB067FA0D153EAA0F330DB9AAE08B7A">
<mods:roleTerm id="E800498ADDB44B01E3998CFB61F39E36">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="171BF031C6F400B99C0D1A4CEBEAC717">Shih, Hsi-Te</mods:namePart>
<mods:affiliation id="584BAD7CE31234A4D4AF827CD0FD289A">National Chung Hsing University, Department of Life Science and Global Change Biology Research Center, 250 Kuo Kuang Road, Taichung 402, Taiwan.</mods:affiliation>
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<mods:title id="725A34AA2895E9768DB3B38749CF96B4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EA18BDA0393BB1B203E839C9E88FE22C">
<mods:date id="9FD80968017B4A72F3F171DEE21E1A76">2024</mods:date>
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<mods:number id="FEF94CF0FD7FF7D4E2C2EBA175298EBC">2024-07-04</mods:number>
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<mods:url id="5AC5336318210E57486BD0CCCB1B5788">http://dx.doi.org/10.11646/zootaxa.5476.1.14</mods:url>
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<mods:classification id="C2F944A8B5BAB5F3283389862A914745">journal article</mods:classification>
<mods:identifier id="428B90F7B621B1395930B632B7783F94" type="CLB-Dataset">299687</mods:identifier>
<mods:identifier id="5790A9897569124E7E5D40D824A377EE" type="DOI">10.11646/zootaxa.5476.1.14</mods:identifier>
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<treatment id="F5111D13756DC645FF3BFD34FA404BA7" ID-DOI="http://doi.org/10.5281/zenodo.12724048" ID-GBIF-Taxon="235037553" ID-Zenodo-Dep="12724048" LSID="urn:lsid:plazi:treatment:F5111D13756DC645FF3BFD34FA404BA7" httpUri="http://treatment.plazi.org/id/F5111D13756DC645FF3BFD34FA404BA7" lastPageId="6" lastPageNumber="144" pageId="3" pageNumber="141">
<subSubSection id="35A2FF8E756DC640FF3BFD34FDD44FD8" box="[151,520,643,670]" pageId="3" pageNumber="141" type="nomenclature">
<paragraph id="7D07AC05756DC640FF3BFD34FDD44FD8" blockId="3.[151,520,643,706]" box="[151,520,643,670]" pageId="3" pageNumber="141">
<heading id="264F1B69756DC640FF3BFD34FDD44FD8" bold="true" box="[151,520,643,670]" fontSize="11" level="1" pageId="3" pageNumber="141" reason="1">
<emphasis id="4FCC7017756DC640FF3BFD34FDD44FD8" bold="true" box="[151,520,643,670]" pageId="3" pageNumber="141">
<taxonomicName id="BAB8D786756DC640FF3BFD34FE7F4FD8" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[151,419,643,670]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="141" phylum="Arthropoda" rank="species" species="ngankeeae" status="sp. nov.">
<emphasis id="4FCC7017756DC640FF3BFD34FE7F4FD8" bold="true" box="[151,419,643,670]" italics="true" pageId="3" pageNumber="141">Parahelice ngankeeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="54FFCD6C756DC640FE07FD33FDD44FD8" box="[427,520,644,670]" pageId="3" pageNumber="141" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="35A2FF8E756DC640FF3BFD1FFECF4F84" box="[151,275,680,706]" pageId="3" pageNumber="141" type="description">
<paragraph id="7D07AC05756DC640FF3BFD1FFECF4F84" blockId="3.[151,520,643,706]" box="[151,275,680,706]" pageId="3" pageNumber="141">
(
<figureCitation id="E583B080756DC640FF33FD1FFF314F84" box="[159,237,680,706]" captionStart="FIGURE 1" captionStartId="4.[151,250,1921,1945]" captionTargetBox="[151,1435,450,1895]" captionTargetId="figure-154@4.[151,1435,448,1896]" captionTargetPageId="4" captionText="FIGURE 1. Parahelice ngankeeae sp. nov. A, B, F, H, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, D, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043); E, G, male (20.0 × 16.2 mm, ZRC 2023.0244). EH, color in life (courtesy of Heok Hui Tan); A, C, E, F, dorsal view; B, D, ventral view; G, H, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12724046" httpUri="https://zenodo.org/record/12724046/files/figure.png" pageId="3" pageNumber="141">Figs. 1</figureCitation>
,
<figureCitation id="E583B080756DC640FF57FD1FFED64F84" box="[251,266,680,706]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="3" pageNumber="141">2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="35A2FF8E756DC640FF3BFD47FDE84908" pageId="3" pageNumber="141" type="materials_examined">
<paragraph id="7D07AC05756DC640FF3BFD47FDE84908" blockId="3.[151,1437,752,2039]" pageId="3" pageNumber="141">
<emphasis id="4FCC7017756DC640FF3BFD47FE944E4C" bold="true" box="[151,328,752,778]" pageId="3" pageNumber="141">Type material.</emphasis>
<materialsCitation id="CDD0A658756DC640FEFEFD47FCCA4E14" ID-GBIF-Occurrence="4905139301" collectingDate="2006-09-10" collectionCode="ZRC" country="Vanuatu" location="Espiritu Santo Island" pageId="3" pageNumber="141" specimenCode="ZRC 2023.0238" specimenCount="1" stateProvince="Sanma" typeStatus="holotype">
<emphasis id="4FCC7017756DC640FEFEFD47FE654E4C" box="[338,441,752,778]" italics="true" pageId="3" pageNumber="141">
<typeStatus id="A20312A7756DC640FEFEFD47FE654E4C" box="[338,441,752,778]" pageId="3" pageNumber="141" type="holotype">Holotype</typeStatus>
</emphasis>
: male (24.3 × 19.1 mm),
<specimenCode id="2D1E047E756DC640FD45FD47FC7F4E4C" box="[745,931,752,778]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="141">ZRC 2023.0238</specimenCode>
, st. H4, shoreline behind
<collectingCountry id="05AFEC95756DC640FB62FD46FAF64E4D" box="[1230,1322,753,779]" name="Vanuatu" pageId="3" pageNumber="141">Vanuatu</collectingCountry>
Maritime College,
<locationDeviation id="6CB5B527756DC640FF50FCA2FE3F4E68" box="[252,483,788,815]" location="Sarakata River" pageId="3" pageNumber="141">W of Sarakata River</locationDeviation>
mouth, Luganville
<collectingRegion id="BF7C62E7756DC640FD6EFCA3FCA74E68" box="[706,891,788,814]" country="Vanuatu" name="Sanma" pageId="3" pageNumber="141">Sanma Province</collectingRegion>
,
<location id="7867FADE756DC640FC24FCA3FBAA4E68" LSID="urn:lsid:plazi:treatment:F5111D13756DC645FF3BFD34FA404BA7:7867FADE756DC640FC24FCA3FBAA4E68" box="[904,1142,788,814]" country="Vanuatu" name="Espiritu Santo Island" pageId="3" pageNumber="141" stateProvince="Sanma">Espiritu Santo Island</location>
, Vanuatu, coll.
<collectorName id="D04DC9D3756DC640FA89FCA2FA444E69" box="[1317,1432,789,815]" pageId="3" pageNumber="141">P.K.L. Ng</collectorName>
, S.H.
<collectorName id="D04DC9D3756DC640FF7EFC8EFEE34E15" box="[210,319,825,851]" pageId="3" pageNumber="141">Tan, H.H</collectorName>
. Tan and
<collectorName id="D04DC9D3756DC640FE1FFC8FFDB74E14" box="[435,619,824,850]" pageId="3" pageNumber="141">J.C.E. Mendoza</collectorName>
,
<date id="09068AC5756DC640FDD6FC8FFCCA4E14" box="[634,790,824,850]" pageId="3" pageNumber="141" value="2006-09-10">
<collectingDate id="1942732D756DC640FDD6FC8FFCCA4E14" box="[634,790,824,850]" pageId="3" pageNumber="141" value="2006-09-10">10 Sep. 2006</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="CDD0A658756DC640FC8FFC8EFD2E4E30" ID-GBIF-Occurrence="4905139302" collectingDate="2006-09-09" collectionCode="ZRC" country="Vanuatu" location="Espiritu Santo Island" pageId="3" pageNumber="141" specimenCode="ZRC 2023.0239" specimenCount="1" specimenCount-male="1" stateProvince="Sanma" typeStatus="paratype">
<emphasis id="4FCC7017756DC640FC8FFC8EFC494E14" box="[803,917,825,850]" italics="true" pageId="3" pageNumber="141">
<typeStatus id="A20312A7756DC640FC8FFC8EFC494E14" box="[803,917,825,850]" pageId="3" pageNumber="141" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="6BBE678C756DC640FC0BFC8FFC294E14" box="[935,1013,824,850]" count="1" pageId="3" pageNumber="141" type="male">1 male</specimenCount>
(12.9 × 10.2 mm),
<specimenCode id="2D1E047E756DC640FB72FC8FFA444E14" box="[1246,1432,824,850]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="141">ZRC 2023.0239</specimenCode>
, same locality and collectors as holotype,
<date id="09068AC5756DC640FDC4FCEBFD2E4E30" box="[616,754,860,886]" pageId="3" pageNumber="141" value="2006-09-09">
<collectingDate id="1942732D756DC640FDC4FCEBFD2E4E30" box="[616,754,860,886]" pageId="3" pageNumber="141" value="2006-09-09">9 Sep. 2006</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="6BBE678C756DC640FD52FCEBFC954E30" box="[766,841,860,886]" count="1" pageId="3" pageNumber="141" type="male">1 male</specimenCount>
(17.0 × 13.4 mm),
<materialsCitation id="CDD0A658756DC640FB8BFCEBFCA24EDC" ID-GBIF-Occurrence="4905139305" collectingDate="2006-09-10" collectionCode="ZRC" country="Vanuatu" location="Espiritu Santo Island" pageId="3" pageNumber="141" specimenCode="ZRC 2023.0240" specimenCount="1" specimenCount-female="1" stateProvince="Sanma" typeStatus="paratype">
<specimenCount id="6BBE678C756DC640FB8BFCEBFB544E30" box="[1063,1160,860,886]" count="1" pageId="3" pageNumber="141" type="female">1 female</specimenCount>
(22.7 × 17.6 mm),
<specimenCode id="2D1E047E756DC640FACAFCEBFECD4EDC" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="141">ZRC 2023.0240</specimenCode>
, same locality and collectors as holotype,
<date id="09068AC5756DC640FD4BFC37FCA24EDC" box="[743,894,896,922]" pageId="3" pageNumber="141" value="2006-09-10">
<collectingDate id="1942732D756DC640FD4BFC37FCA24EDC" box="[743,894,896,922]" pageId="3" pageNumber="141" value="2006-09-10">10 Sep. 2006</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="6BBE678C756DC640FC24FC37FC0F4EDC" box="[904,979,896,922]" count="1" pageId="3" pageNumber="141" type="male">1 male</specimenCount>
(20.9 × 17.1 mm),
<materialsCitation id="CDD0A658756DC640FB00FC37FBDA4EF8" ID-GBIF-Occurrence="4905139309" collectingDate="2006-09-10" collectionCode="ZRC" country="Vanuatu" location="Espiritu Santo Island" pageId="3" pageNumber="141" specimenCode="ZRC 2023.0241" specimenCount="1" specimenCount-female="1" stateProvince="Sanma" typeStatus="paratype">
<specimenCount id="6BBE678C756DC640FB00FC37FAD14EDC" box="[1196,1293,896,922]" count="1" pageId="3" pageNumber="141" type="female">1 female</specimenCount>
(23.9 × 18.7 mm),
<specimenCode id="2D1E047E756DC640FF77FC13FE4F4EF8" box="[219,403,932,958]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="141">ZRC 2023.0241</specimenCode>
, same locality and collectors as holotype,
<date id="09068AC5756DC640FCC2FC13FBDA4EF8" box="[878,1030,932,958]" pageId="3" pageNumber="141" value="2006-09-10">
<collectingDate id="1942732D756DC640FCC2FC13FBDA4EF8" box="[878,1030,932,958]" pageId="3" pageNumber="141" value="2006-09-10">10 Sep. 2006</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="6BBE678C756DC640FBBEFC13FB814EF8" box="[1042,1117,932,958]" count="1" pageId="3" pageNumber="141" type="male">1 male</specimenCount>
(23.8 × 18.6 mm),
<materialsCitation id="CDD0A658756DC640FA96FC13FB434EA4" ID-GBIF-Occurrence="4905139312" collectingDate="2006-09-11" collectionCode="ZRC" country="Vanuatu" location="Espiritu Santo Island" pageId="3" pageNumber="141" specimenCode="ZRC 2023.0242" specimenCount="1" specimenCount-female="1" stateProvince="Sanma" typeStatus="paratype">
<specimenCount id="6BBE678C756DC640FA96FC13FA404EF8" box="[1338,1436,932,958]" count="1" pageId="3" pageNumber="141" type="female">1 female</specimenCount>
(24.5 × 18.8 mm),
<specimenCode id="2D1E047E756DC640FEC2FC7FFDFB4EA4" box="[366,551,968,994]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="141">ZRC 2023.0242</specimenCode>
, same locality and collectors as holotype,
<date id="09068AC5756DC640FBABFC7FFB434EA4" box="[1031,1183,968,994]" pageId="3" pageNumber="141" value="2006-09-11">
<collectingDate id="1942732D756DC640FBABFC7FFB434EA4" box="[1031,1183,968,994]" pageId="3" pageNumber="141" value="2006-09-11">11 Sep. 2006</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="6BBE678C756DC640FB06FC7FFADE4EA4" box="[1194,1282,968,994]" count="3" pageId="3" pageNumber="141" type="male">3 males</specimenCount>
(17.6 × 14.6, 14.9 × 12.0, 13.2 × 10.8 mm),
<materialsCitation id="CDD0A658756DC640FE4BFC5BFEF3496C" ID-GBIF-Occurrence="4905139307" collectingDate="2006-09-10" collectionCode="ZRC" country="Vanuatu" location="Espiritu Santo Island" pageId="3" pageNumber="141" specimenCode="ZRC 2023.2043" specimenCount="1" specimenCount-female="1" stateProvince="Sanma" typeStatus="paratype">
<specimenCount id="6BBE678C756DC640FE4BFC5BFD9B4940" box="[487,583,1004,1030]" count="1" pageId="3" pageNumber="141" type="female">1 female</specimenCount>
(15.6 × 12.8 mm),
<specimenCode id="2D1E047E756DC640FCB7FC5BFC0E4940" box="[795,978,1004,1030]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="141">ZRC 2023.2043</specimenCode>
, same locality and collectors as
<typeStatus id="A20312A7756DC640FA9FFC5BFA444940" box="[1331,1432,1004,1030]" pageId="3" pageNumber="141" type="holotype">holotype</typeStatus>
,
<date id="09068AC5756DC640FF3BFBA7FEF3496C" box="[151,303,1040,1066]" pageId="3" pageNumber="141" value="2006-09-10">
<collectingDate id="1942732D756DC640FF3BFBA7FEF3496C" box="[151,303,1040,1066]" pageId="3" pageNumber="141" value="2006-09-10">10 Sep. 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="CDD0A658756DC640FE96FBA7FDED4908" ID-GBIF-Occurrence="4905139306" collectingDate="2006-09-23" collectionCode="ZRC" country="Vanuatu" location="Palikulo Peninsula" municipality="Espiritu Santo Island" pageId="3" pageNumber="141" specimenCode="ZRC 2023.0244" specimenCount="1" specimenCount-male="1" stateProvince="Sanma" typeStatus="paratype">
<specimenCount id="6BBE678C756DC640FE96FBA7FE5A496C" box="[314,390,1040,1066]" count="1" pageId="3" pageNumber="141" type="male">1 male</specimenCount>
(20.0 × 16.2 mm),
<specimenCode id="2D1E047E756DC640FDCEFBA7FCC6496C" box="[610,794,1040,1066]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="3" pageNumber="141">ZRC 2023.0244</specimenCode>
, st. VM34,
<location id="7867FADE756DC640FC32FBA7FBAF496C" LSID="urn:lsid:plazi:treatment:F5111D13756DC645FF3BFD34FA404BA7:7867FADE756DC640FC32FBA7FBAF496C" box="[926,1139,1040,1066]" country="Vanuatu" municipality="Espiritu Santo Island" name="Palikulo Peninsula" pageId="3" pageNumber="141" stateProvince="Sanma">Palikulo Peninsula</location>
,
<collectingRegion id="BF7C62E7756DC640FBD2FBA7FAE4496C" box="[1150,1336,1040,1066]" country="Vanuatu" name="Sanma" pageId="3" pageNumber="141">Sanma Province</collectingRegion>
,
<collectingMunicipality id="9D63367F756DC640FAE8FBA7FEF94908" pageId="3" pageNumber="141">Espiritu Santo Island</collectingMunicipality>
,
<collectingCountry id="05AFEC95756DC640FE9CFB82FE534909" box="[304,399,1077,1103]" name="Vanuatu" pageId="3" pageNumber="141">Vanuatu</collectingCountry>
,
<date id="09068AC5756DC640FE36FB83FDED4908" box="[410,561,1076,1102]" pageId="3" pageNumber="141" value="2006-09-23">
<collectingDate id="1942732D756DC640FE36FB83FDED4908" box="[410,561,1076,1102]" pageId="3" pageNumber="141" value="2006-09-23">23 Sep. 2006</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="35A2FF8E756DC640FF6BFBEFFD424B01" pageId="3" pageNumber="141" type="diagnosis">
<paragraph id="7D07AC05756DC640FF6BFBEFFD424B01" blockId="3.[151,1437,752,2039]" pageId="3" pageNumber="141">
<emphasis id="4FCC7017756DC640FF6BFBEFFE9E4934" bold="true" box="[199,322,1112,1138]" pageId="3" pageNumber="141">Diagnosis.</emphasis>
Carapace (
<figureCitation id="E583B080756DC640FE64FBEFFD444935" box="[456,664,1112,1139]" captionStart="FIGURE 1" captionStartId="4.[151,250,1921,1945]" captionTargetBox="[151,1435,450,1895]" captionTargetId="figure-154@4.[151,1435,448,1896]" captionTargetPageId="4" captionText="FIGURE 1. Parahelice ngankeeae sp. nov. A, B, F, H, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, D, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043); E, G, male (20.0 × 16.2 mm, ZRC 2023.0244). EH, color in life (courtesy of Heok Hui Tan); A, C, E, F, dorsal view; B, D, ventral view; G, H, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12724046" httpUri="https://zenodo.org/record/12724046/files/figure.png" pageId="3" pageNumber="141">Figs. 1A, C, E, F</figureCitation>
,
<figureCitation id="E583B080756DC640FD04FBEFFD174934" box="[680,715,1112,1138]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="3" pageNumber="141">2A</figureCitation>
) quadrate, 1.221.29 times as broad as long; surface convex, irregularly punctated and finely granulated. Front broad, frontal margin slightly concave in both dorsal and anterior views; postfrontal region separated into 2 lobes.Anterolateral margins with 3 distinct teeth including external orbital tooth. Suborbital crista (
<figureCitation id="E583B080756DC640FE0AFB73FDC14998" box="[422,541,1220,1246]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="3" pageNumber="141">Fig. 2B, C</figureCitation>
) sexually heteromorphic; in male (
<figureCitation id="E583B080756DC640FC08FB73FC204998" box="[932,1020,1220,1246]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="3" pageNumber="141">Fig. 2B</figureCitation>
), mesial part with 69 small rounded tubercles, followed by several (1625) larger, elongated and fused tubercles, ending with 1 elliptical tubercle, lateral part with 1 largest, very convex elliptical tubercle, followed laterally by 2 or 3 smaller tubercle; in female (
<figureCitation id="E583B080756DC640FA94FABBFA534860" box="[1336,1423,1292,1318]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="3" pageNumber="141">Fig. 2C</figureCitation>
), mesial part with 5 or 6 small rounded tubercles, followed by several (712) larger, elongated and mostly unfused tubercles, lateral part with 2 or 3 more convex, larger elliptical tubercles, ending with 1 or 2 small rounded tubercles. Chelipedal palm bulky in male, smaller in female; inner surface glabrous; outer surface with distinct patch of setae at base of fingers in male (
<figureCitation id="E583B080756DC640FE16FA2BFDCF48F0" box="[442,531,1436,1462]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="3" pageNumber="141">Fig. 2D</figureCitation>
); only line of short setae at base of occlusal margin of immovable finger in female (
<figureCitation id="E583B080756DC640FF33FA77FF26489C" box="[159,250,1472,1498]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="3" pageNumber="141">Fig. 2E</figureCitation>
). Ambulatory legs slender, anterior margins of meri, carpi, and propodi covered with short setae. Male pleon (
<figureCitation id="E583B080756DC640FF4FFA53FEEA48B8" box="[227,310,1508,1534]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="3" pageNumber="141">Fig. 2F</figureCitation>
) narrow, distal margin of telson not concave. G1 (
<figureCitation id="E583B080756DC640FCF1FA53FC0748B8" box="[861,987,1508,1534]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="3" pageNumber="141">Fig. 2HM</figureCitation>
) slender, slightly curved, tapering, distal part curved laterally and ventrally; tip chitinous, chitinous endpiece shorter, wider and thicker. Female vulvae (
<figureCitation id="E583B080756DC640FAC3F9BFFF674B00" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="3" pageNumber="141">Fig. 2N</figureCitation>
) with elongated semicircular sternal cover.
</paragraph>
</subSubSection>
<subSubSection id="35A2FF8E756DC645FF6BF9E7FB774CEB" lastPageId="6" lastPageNumber="144" pageId="3" pageNumber="141" type="description">
<paragraph id="7D07AC05756DC640FF6BF9E7FB374A21" blockId="3.[151,1437,752,2039]" pageId="3" pageNumber="141">
<emphasis id="4FCC7017756DC640FF6BF9E7FE854B2C" bold="true" box="[199,345,1616,1642]" pageId="3" pageNumber="141">Description.</emphasis>
Carapace (
<figureCitation id="E583B080756DC640FE79F9E7FDF14B2C" box="[469,557,1616,1642]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="3" pageNumber="141">Fig. 2A</figureCitation>
) quadrate, slightly broader than long, 1.221.29 times as broad as long; surface convex, irregularly punctated and finely granulated; meso- to protogastric regions convex, with longitudinal deep groove; H-shaped gastro-cardiac groove prominent. Front broad, sloping forward; frontal margin lined with small, rounded granules, slightly concave in both dorsal and anterior views; postfrontal region slightly separated into 2 convex lobes. Supraorbital margins and anterolateral margins lined with small granules. Anterolateral margins with 3 distinct teeth including external orbital tooth; first tooth (external orbital tooth) largest, broad, slightly sloping forward; second tooth slightly narrower and sharper than preceding; last tooth smallest, distinct, acute. Posterolateral margins slightly convergent posteriorly; posterolateral regions regularly furnished with short setae.
</paragraph>
<paragraph id="7D07AC05756DC647FF6BF8C7FBCC4DBF" blockId="3.[151,1437,752,2039]" lastBlockId="4.[151,1437,150,429]" lastPageId="4" lastPageNumber="142" pageId="3" pageNumber="141">
Suborbital crista (
<figureCitation id="E583B080756DC640FE3CF8C7FDD44ACC" box="[400,520,1904,1930]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="3" pageNumber="141">Fig. 2B, C</figureCitation>
) sexually heteromorphic; in male, mesial part with several (69) less interspaced small rounded tubercles, followed by several (1625) larger, elongated, less convex and fused tubercles, ending with 1 significantly larger and convex elliptical tubercle, lateral part with 1 significantly largest, very convex and elliptical tubercle, followed laterally by 1 large convex elongated tubercle and 1 or 2 small, convex, rounded tubercle (
<figureCitation id="E583B080756AC647FF33FF20FF2B4DF7" box="[159,247,151,177]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="4" pageNumber="142">Fig. 2B</figureCitation>
); in female, mesial part with several (5 or 6) less interspaced small rounded tubercles, followed by several (712) larger, less convex elongated and mostly unfused tubercles, lateral part with well-spaced 2 or 3 more convex, larger elliptical tubercles, ending with 1 or 2 small rounded tubercles (
<figureCitation id="E583B080756AC647FC06FF68FBDF4DBF" box="[938,1027,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="4" pageNumber="142">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="7D07AC05756AC647FF6BFEB4FBD04CEB" blockId="4.[151,1437,150,429]" pageId="4" pageNumber="142">
Chelipeds (
<figureCitation id="E583B080756AC647FEEBFEB4FE5B4C5B" box="[327,391,259,285]" captionStart="FIGURE 1" captionStartId="4.[151,250,1921,1945]" captionTargetBox="[151,1435,450,1895]" captionTargetId="figure-154@4.[151,1435,448,1896]" captionTargetPageId="4" captionText="FIGURE 1. Parahelice ngankeeae sp. nov. A, B, F, H, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, D, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043); E, G, male (20.0 × 16.2 mm, ZRC 2023.0244). EH, color in life (courtesy of Heok Hui Tan); A, C, E, F, dorsal view; B, D, ventral view; G, H, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12724046" httpUri="https://zenodo.org/record/12724046/files/figure.png" pageId="4" pageNumber="142">Fig. 1</figureCitation>
) usually unequal in adult male and equal in female. Merus without spines, dorsal and ventral margins glabrous. Surface of carpus glabrous, margins of carpus with dense short setae, inner distal angle with 13 tiny teeth. Palm bulky; inner surface almost smooth, glabrous; outer surface almost smooth; in male (
<figureCitation id="E583B080756AC647FA9EFEFCFA534C23" box="[1330,1423,331,357]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="4" pageNumber="142">Fig. 2D</figureCitation>
), distinct patch of setae at base of fingers, mostly expanding onto fixed finger, in both chelae; in female (
<figureCitation id="E583B080756AC647FA9BFED8FA534CCF" box="[1335,1423,367,393]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="4" pageNumber="142">Fig. 2E</figureCitation>
), only line of short setae present at base of occlusal margin of immovable finger.
</paragraph>
<caption id="29C7FC8D756AC647FF3BF836FC904AA4" ID-DOI="http://doi.org/10.5281/zenodo.12724046" ID-Zenodo-Dep="12724046" httpUri="https://zenodo.org/record/12724046/files/figure.png" pageId="4" pageNumber="142" startId="4.[151,250,1921,1945]" targetBox="[151,1435,450,1895]" targetPageId="4" targetType="figure">
<paragraph id="7D07AC05756AC647FF3BF836FC904AA4" blockId="4.[151,1436,1921,2018]" pageId="4" pageNumber="142">
<emphasis id="4FCC7017756AC647FF3BF836FECF4ADF" bold="true" box="[151,275,1921,1945]" pageId="4" pageNumber="142">FIGURE 1.</emphasis>
<taxonomicName id="BAB8D786756AC647FEB4F835FE254ADC" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[280,505,1922,1946]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="142" phylum="Arthropoda" rank="species" species="ngankeeae" status="sp. nov.">
<emphasis id="4FCC7017756AC647FEB4F835FE254ADC" box="[280,505,1922,1946]" italics="true" pageId="4" pageNumber="142">Parahelice ngankeeae</emphasis>
</taxonomicName>
<emphasis id="4FCC7017756AC647FE53F835FD934ADC" bold="true" box="[511,591,1922,1946]" pageId="4" pageNumber="142">
<taxonomicNameLabel id="54FFCD6C756AC647FE53F835FD934ADC" box="[511,591,1922,1946]" pageId="4" pageNumber="142" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, B, F, H, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, D, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043); E, G, male (20.0 × 16.2 mm, ZRC 2023.0244). EH, color in life (courtesy of Heok Hui Tan); A, C, E, F, dorsal view; B, D, ventral view; G, H, frontal view.
</paragraph>
</caption>
<caption id="29C7FC8D756BC646FF3BF8C2FC6C4ABF" ID-DOI="http://doi.org/10.5281/zenodo.12724052" ID-Zenodo-Dep="12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="5" pageNumber="143" startId="5.[151,250,1909,1933]" targetBox="[205,1381,189,1883]" targetPageId="5" targetType="figure">
<paragraph id="7D07AC05756BC646FF3BF8C2FC6C4ABF" blockId="5.[151,1436,1909,2041]" pageId="5" pageNumber="143">
<emphasis id="4FCC7017756BC646FF3BF8C2FEC84ACB" bold="true" box="[151,276,1909,1933]" pageId="5" pageNumber="143">FIGURE 2.</emphasis>
<taxonomicName id="BAB8D786756BC646FEB7F8C2FE214ACB" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[283,509,1909,1933]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="143" phylum="Arthropoda" rank="species" species="ngankeeae" status="sp. nov.">
<emphasis id="4FCC7017756BC646FEB7F8C2FE214ACB" box="[283,509,1909,1933]" italics="true" pageId="5" pageNumber="143">Parahelice ngankeeae</emphasis>
</taxonomicName>
<emphasis id="4FCC7017756BC646FDA9F8C2FD8A4ACB" bold="true" box="[517,598,1909,1933]" pageId="5" pageNumber="143">
<taxonomicNameLabel id="54FFCD6C756BC646FDA9F8C2FD8A4ACB" box="[517,598,1909,1933]" pageId="5" pageNumber="143" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva.
</paragraph>
</caption>
<paragraph id="7D07AC057568C645FF6BFF20FDBB4D93" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="144">
Ambulatory legs (
<figureCitation id="E583B0807568C645FE3AFF20FE044DF7" box="[406,472,151,177]" captionStart="FIGURE 1" captionStartId="4.[151,250,1921,1945]" captionTargetBox="[151,1435,450,1895]" captionTargetId="figure-154@4.[151,1435,448,1896]" captionTargetPageId="4" captionText="FIGURE 1. Parahelice ngankeeae sp. nov. A, B, F, H, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, D, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043); E, G, male (20.0 × 16.2 mm, ZRC 2023.0244). EH, color in life (courtesy of Heok Hui Tan); A, C, E, F, dorsal view; B, D, ventral view; G, H, frontal view." figureDoi="http://doi.org/10.5281/zenodo.12724046" httpUri="https://zenodo.org/record/12724046/files/figure.png" pageId="6" pageNumber="144">Fig. 1</figureCitation>
) slender, anterior margins of merus, carpus, and propodus covered with short setae, posterior margins with sparse short setae.
</paragraph>
<paragraph id="7D07AC057568C645FF6BFF68FC394C5B" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="144">
Male pleon (
<figureCitation id="E583B0807568C645FEFFFF68FE794DBF" box="[339,421,223,249]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="6" pageNumber="144">Fig. 2F</figureCitation>
) narrow, surface smooth, without any granules, lateral margins lined with short setae; telson tongue-shaped, short than sixth somite, distal margin of telson not concave.
</paragraph>
<paragraph id="7D07AC057568C645FF6BFE90FF2B4CCF" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="144">
G1 (
<figureCitation id="E583B0807568C645FF57FE90FE594C07" box="[251,389,295,321]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="6" pageNumber="144">Fig. 2 HM</figureCitation>
) slender, slightly curved towards lateral, tapering, distal part curved laterally and ventrally; tip chitinous, chitinous endpiece shorter, wider and thicker, genital pore widely open distally towards laterally and dorsally.
</paragraph>
<paragraph id="7D07AC057568C645FF6BFE24FB774CEB" blockId="6.[151,1437,151,1761]" box="[199,1195,403,429]" pageId="6" pageNumber="144">
Female vulvae (
<figureCitation id="E583B0807568C645FED7FE24FE0F4CEB" box="[379,467,403,429]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="6" pageNumber="144">Fig. 2N</figureCitation>
) with elongated semicircular sternal cover; sunken on inner part.
</paragraph>
</subSubSection>
<subSubSection id="35A2FF8E7568C645FF6BFE00FDEF4F7B" pageId="6" pageNumber="144" type="etymology">
<paragraph id="7D07AC057568C645FF6BFE00FC5C4CB3" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="144">
<emphasis id="4FCC70177568C645FF6BFE00FE924C97" bold="true" box="[199,334,439,465]" pageId="6" pageNumber="144">Etymology.</emphasis>
The species is named after the late Dr. Ngan Kee Ng for her contribution to the taxonomy of brachyurans, especially the family
<taxonomicName id="BAB8D7867568C645FDB2FE6CFCA04CB3" authority="(Lee &amp; Jaafar 2022)" baseAuthorityName="Lee &amp; Jaafar" baseAuthorityYear="2022" box="[542,892,475,501]" class="Malacostraca" family="Varunidae" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="family">
Varunidae (
<bibRefCitation id="1929D1F47568C645FD32FE6CFCA84CB3" author="Lee, B. Y. &amp; Jaafar, Z." box="[670,884,475,501]" pageId="6" pageNumber="144" pagination="1255 - 1264" refId="ref9545" refString="Lee, B. Y. &amp; Jaafar, Z. (2022) In memorian Ng Ngan Kee (1966 - 2022). Crustaceana, 95, 1255 - 1264. https: // doi. org / 10.1163 / 15685403 - bja 10229" type="journal article" year="2022">Lee &amp; Jaafar 2022</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="7D07AC057568C645FF6BFE49FDEF4F7B" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="144">
<emphasis id="4FCC70177568C645FF6BFE49FE4D4F5F" bold="true" box="[199,401,510,537]" pageId="6" pageNumber="144">Ecological notes.</emphasis>
This species is common in mangrove and intertidal areas with sandy sediments (J. C. E. Mendoza, personal communication).
</paragraph>
</subSubSection>
<subSubSection id="35A2FF8E7568C645FF6BFDF1FA854FC3" pageId="6" pageNumber="144" type="distribution">
<paragraph id="7D07AC057568C645FF6BFDF1FD464F27" blockId="6.[151,1437,151,1761]" box="[199,666,582,609]" pageId="6" pageNumber="144">
<emphasis id="4FCC70177568C645FF6BFDF1FEBD4F26" bold="true" box="[199,353,582,608]" pageId="6" pageNumber="144">Distribution.</emphasis>
Only known from
<collectingCountry id="05AFEC957568C645FD9BFDF0FD4B4F27" box="[567,663,583,609]" name="Vanuatu" pageId="6" pageNumber="144">Vanuatu</collectingCountry>
.
</paragraph>
<paragraph id="7D07AC057568C645FF6BFDDDFA854FC3" blockId="6.[151,1437,151,1761]" box="[199,1369,618,645]" pageId="6" pageNumber="144">
<emphasis id="4FCC70177568C645FF6BFDDDFF224FC2" bold="true" box="[199,254,618,644]" pageId="6" pageNumber="144">Size.</emphasis>
Largest male 24.3 ×
<quantity id="BA4001E07568C645FE40FDDCFD8C4FC3" box="[492,592,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="1.91" pageId="6" pageNumber="144" unit="mm" value="19.1">19.1 mm</quantity>
(ZRC 2023.0238); largest female 24.5 ×
<quantity id="BA4001E07568C645FB8CFDDCFB584FC3" box="[1056,1156,619,645]" metricMagnitude="-2" metricUnit="m" metricValue="1.88" pageId="6" pageNumber="144" unit="mm" value="18.8">18.8 mm</quantity>
(ZRC 2023.0242).
</paragraph>
</subSubSection>
<subSubSection id="35A2FF8E7568C645FF6BFD38FA404BA7" pageId="6" pageNumber="144" type="discussion">
<paragraph id="7D07AC057568C645FF6BFD38FCE34E1B" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="144">
<emphasis id="4FCC70177568C645FF6BFD38FEE74FEF" bold="true" box="[199,315,655,681]" pageId="6" pageNumber="144">Remarks.</emphasis>
Among the males of species in
<taxonomicName id="BAB8D7867568C645FD07FD38FCFA4FEF" authorityName="K. Sakai, Turkay &amp; Yang" authorityYear="2006" box="[683,806,655,681]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="4FCC70177568C645FD07FD38FCFA4FEF" box="[683,806,655,681]" italics="true" pageId="6" pageNumber="144">Parahelice</emphasis>
</taxonomicName>
and
<taxonomicName id="BAB8D7867568C645FCF3FD38FC284FEF" authorityName="K. Sakai, Turkay &amp; Yang" authorityYear="2006" box="[863,1012,655,681]" class="Malacostraca" family="Varunidae" genus="Pseudohelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="4FCC70177568C645FCF3FD38FC284FEF" box="[863,1012,655,681]" italics="true" pageId="6" pageNumber="144">Pseudohelice</emphasis>
</taxonomicName>
,
<taxonomicName id="BAB8D7867568C645FBAFFD38FB534FEE" baseAuthorityName="Clark" baseAuthorityYear="1987" box="[1027,1167,655,680]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="georgei">
<emphasis id="4FCC70177568C645FBAFFD38FB534FEE" box="[1027,1167,655,680]" italics="true" pageId="6" pageNumber="144">Par. georgei</emphasis>
</taxonomicName>
and all three species of
<taxonomicName id="BAB8D7867568C645FF3BFD04FEF04F8B" authorityName="K. Sakai, Turkay &amp; Yang" authorityYear="2006" box="[151,300,691,717]" class="Malacostraca" family="Varunidae" genus="Pseudohelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="genus">
<emphasis id="4FCC70177568C645FF3BFD04FEF04F8B" box="[151,300,691,717]" italics="true" pageId="6" pageNumber="144">Pseudohelice</emphasis>
</taxonomicName>
(
<taxonomicName id="BAB8D7867568C645FE93FD04FE2C4F8B" authorityName="Prema, Hsu, Shih &amp; Ravichandran" authorityYear="2022" box="[319,496,691,717]" class="Malacostraca" family="Varunidae" genus="Pseudohelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="annamalai">
<emphasis id="4FCC70177568C645FE93FD04FE2C4F8B" box="[319,496,691,717]" italics="true" pageId="6" pageNumber="144">Pse. annamalai</emphasis>
</taxonomicName>
,
<taxonomicName id="BAB8D7867568C645FDACFD04FD4F4F8B" baseAuthorityName="H. Milne Edwards" baseAuthorityYear="1837" box="[512,659,691,717]" class="Malacostraca" family="Varunidae" genus="Pseudohelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="latreillii">
<emphasis id="4FCC70177568C645FDACFD04FD4F4F8B" box="[512,659,691,717]" italics="true" pageId="6" pageNumber="144">Pse. latreillii</emphasis>
</taxonomicName>
, and
<taxonomicName id="BAB8D7867568C645FD79FD04FC404F8B" baseAuthorityName="Dana" baseAuthorityYear="1851" box="[725,924,691,717]" class="Malacostraca" family="Varunidae" genus="Pseudohelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="subquadrata">
<emphasis id="4FCC70177568C645FD79FD04FC404F8B" box="[725,924,691,717]" italics="true" pageId="6" pageNumber="144">Pse. subquadrata</emphasis>
</taxonomicName>
) can be easily distinguished from remaining four
<taxonomicName id="BAB8D7867568C645FF7DFD60FE7B4FB7" box="[209,423,727,753]" pageId="6" pageNumber="144">
<emphasis id="4FCC70177568C645FF7DFD60FE904FB7" box="[209,332,727,753]" italics="true" pageId="6" pageNumber="144">Parahelice</emphasis>
species
</taxonomicName>
by the absence of a patch of setae in
<taxonomicName id="BAB8D7867568C645FCC3FD60FC214FB6" baseAuthorityName="Clark" baseAuthorityYear="1987" box="[879,1021,727,752]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="georgei">
<emphasis id="4FCC70177568C645FCC3FD60FC214FB6" box="[879,1021,727,752]" italics="true" pageId="6" pageNumber="144">Par. georgei</emphasis>
</taxonomicName>
and
<taxonomicName id="BAB8D7867568C645FB97FD60FAF04FB7" box="[1083,1324,727,753]" pageId="6" pageNumber="144">
<emphasis id="4FCC70177568C645FB97FD60FB0C4FB7" box="[1083,1232,727,753]" italics="true" pageId="6" pageNumber="144">Pseudohelice</emphasis>
species
</taxonomicName>
[vs. with patch of setae in other
<taxonomicName id="BAB8D7867568C645FE3AFD4CFDB04E53" box="[406,620,763,789]" pageId="6" pageNumber="144">
<emphasis id="4FCC70177568C645FE3AFD4CFDCD4E53" box="[406,529,763,789]" italics="true" pageId="6" pageNumber="144">Parahelice</emphasis>
species
</taxonomicName>
] (K.
<bibRefCitation id="1929D1F47568C645FD0AFD4CFCBA4E53" author="Sakai, K. &amp; Turkay, M. &amp; Yang, S. - L." box="[678,870,763,789]" pageId="6" pageNumber="144" pagination="1 - 76" refId="ref10458" refString="Sakai, K., Turkay, M. &amp; Yang, S. - L. (2006) Revision of the Helice / Chasmagnathus complex (Crustacea: Decapoda: Brachyura). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 565, 1 - 76." type="journal article" year="2006">
Sakai
<emphasis id="4FCC70177568C645FD47FD4CFCC04E53" box="[747,796,763,789]" italics="true" pageId="6" pageNumber="144">et al</emphasis>
. 2006
</bibRefCitation>
;
<bibRefCitation id="1929D1F47568C645FCDDFD4CFBFD4E53" author="Hsu, J. - W. &amp; Shih, H. - T. &amp; Innocenti, G." box="[881,1057,763,789]" pageId="6" pageNumber="144" pagination="94 - 107" refId="ref9196" refString="Hsu, J. - W., Shih, H. - T. &amp; Innocenti, G. (2022) A review of the mud crab genus Pseudohelice Sakai, Turkay &amp; Yang, 2006 (Crustacea: Brachyura: Varunidae), with redescription of Cyclograpsus latreillii H. Milne Edwards, 1837, from the western Indian Ocean. Raffles Bulletin of Zoology, 70, 94 - 107. https: // doi. org / 10.26107 / RBZ- 2022 - 0007" type="journal article" year="2022">
Hsu
<emphasis id="4FCC70177568C645FC0AFD4CFC024E53" box="[934,990,763,789]" italics="true" pageId="6" pageNumber="144">et al.</emphasis>
2022
</bibRefCitation>
;
<bibRefCitation id="1929D1F47568C645FB80FD4CFB2A4E53" author="Prema, M. &amp; Hsu, J. - W. &amp; Shih, H. - T. &amp; Ravichandran, S." box="[1068,1270,763,789]" pageId="6" pageNumber="144" refId="ref10142" refString="Prema, M., Hsu, J. - W., Shih, H. - T. &amp; Ravichandran, S. (2022) First record of the genus Pseudohelice Sakai, Turkay &amp; Yang, 2006 from India and description of a new pseudocryptic species (Crustacea: Brachyura: Varunidae). Zoological Studies, 61, 56. https: // doi. org / 10.6620 / ZS. 2022.61 - 56" type="journal volume" year="2022">
Prema
<emphasis id="4FCC70177568C645FBD7FD4CFB6F4E53" box="[1147,1203,763,789]" italics="true" pageId="6" pageNumber="144">et al.</emphasis>
2022
</bibRefCitation>
; this study). In this regard,
<taxonomicName id="BAB8D7867568C645FE8CFCA8FE124E7F" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[288,462,799,825]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="ngankeeae" status="sp. nov.">
<emphasis id="4FCC70177568C645FE8CFCA8FE124E7F" box="[288,462,799,825]" italics="true" pageId="6" pageNumber="144">Par. ngankeeae</emphasis>
</taxonomicName>
<emphasis id="4FCC70177568C645FE7BFCA8FDEE4E7F" bold="true" box="[471,562,799,825]" pageId="6" pageNumber="144">
<taxonomicNameLabel id="54FFCD6C7568C645FE7BFCA8FDEE4E7F" box="[471,562,799,825]" pageId="6" pageNumber="144" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="BAB8D7867568C645FD18FCA8FCF74E7F" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[692,811,799,825]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="4FCC70177568C645FD18FCA8FCF74E7F" box="[692,811,799,825]" italics="true" pageId="6" pageNumber="144">Par. balssi</emphasis>
</taxonomicName>
,
<taxonomicName id="BAB8D7867568C645FC90FCA8FC6A4E7F" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[828,950,799,825]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="4FCC70177568C645FC90FCA8FC6A4E7F" box="[828,950,799,825]" italics="true" pageId="6" pageNumber="144">Par. daviei</emphasis>
</taxonomicName>
,
<taxonomicName id="BAB8D7867568C645FC6AFCA8FBBD4E7F" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[966,1121,799,825]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="pilimana">
<emphasis id="4FCC70177568C645FC6AFCA8FBBD4E7F" box="[966,1121,799,825]" italics="true" pageId="6" pageNumber="144">Par. pilimana</emphasis>
</taxonomicName>
and
<taxonomicName id="BAB8D7867568C645FB37FCA8FAC94E7F" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[1179,1301,799,825]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="4FCC70177568C645FB37FCA8FAC94E7F" box="[1179,1301,799,825]" italics="true" pageId="6" pageNumber="144">Par. pilosa</emphasis>
</taxonomicName>
, but can be separated by the characters of the suborbital cristae and G1s.
</paragraph>
<paragraph id="7D07AC057568C645FF6BFCD0FEFE491F" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="144">
<taxonomicName id="BAB8D7867568C645FF6BFCD0FE1E4EC7" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[199,450,871,897]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="4FCC70177568C645FF6BFCD0FE1E4EC7" box="[199,450,871,897]" italics="true" pageId="6" pageNumber="144">Parahelice ngankeeae</emphasis>
</taxonomicName>
can be discerned from
<taxonomicName id="BAB8D7867568C645FD62FCD0FC994EC7" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[718,837,871,897]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="4FCC70177568C645FD62FCD0FC994EC7" box="[718,837,871,897]" italics="true" pageId="6" pageNumber="144">Par. balssi</emphasis>
</taxonomicName>
by the male suborbital cristae and G1s. In male
<taxonomicName id="BAB8D7867568C645FAC2FCD0FED34EE3" authorityName="Hsu &amp; Shih" authorityYear="2024" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="4FCC70177568C645FAC2FCD0FED34EE3" italics="true" pageId="6" pageNumber="144">Par. ngankeeae</emphasis>
</taxonomicName>
, the lateral part of the suborbital crista consists of 1 significantly largest elliptical tubercle and 23 elongated or rounded tubercles, which are significantly smaller than the largest elliptical tubercle (
<figureCitation id="E583B0807568C645FAA4FC18FABD4E8F" box="[1288,1377,943,969]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="6" pageNumber="144">Fig. 2B</figureCitation>
) [vs. elongated crest ending with inflated part, followed by 24 elongated tubercles, which are larger or equal in size to the inflated part in
<taxonomicName id="BAB8D7867568C645FEDEFC40FE354957" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[370,489,1015,1041]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="4FCC70177568C645FEDEFC40FE354957" box="[370,489,1015,1041]" italics="true" pageId="6" pageNumber="144">Par. balssi</emphasis>
</taxonomicName>
(K.
<bibRefCitation id="1929D1F47568C645FDB3FC40FD384957" author="Sakai, K. &amp; Turkay, M. &amp; Yang, S. - L." box="[543,740,1015,1041]" pageId="6" pageNumber="144" pagination="1 - 76" refId="ref10458" refString="Sakai, K., Turkay, M. &amp; Yang, S. - L. (2006) Revision of the Helice / Chasmagnathus complex (Crustacea: Decapoda: Brachyura). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 565, 1 - 76." type="journal article" year="2006">
Sakai
<emphasis id="4FCC70177568C645FDCAFC40FD7C4957" box="[614,672,1015,1041]" italics="true" pageId="6" pageNumber="144">et al.</emphasis>
2006
</bibRefCitation>
: fig. 60;
<bibRefCitation id="1929D1F47568C645FCE7FC40FBE24957" author="Bouchard, J. M. &amp; Poupin, J. &amp; Cleva, R. &amp; Dumas, J. &amp; Dinhut, V." box="[843,1086,1015,1041]" pageId="6" pageNumber="144" pagination="1 - 60" refId="ref8648" refString="Bouchard, J. M., Poupin, J., Cleva, R., Dumas, J. &amp; Dinhut, V. (2013) Land, mangrove and freshwater decapod crustaceans of Mayotte region (Crustacea Decapoda). Atoll Research Bulletin, 592, 1 - 60. https: // doi. org / 10.5479 / si. 00775630.592" type="journal article" year="2013">
Bouchard
<emphasis id="4FCC70177568C645FC6CFC40FC2F4957" box="[960,1011,1015,1041]" italics="true" pageId="6" pageNumber="144">et al</emphasis>
. 2013
</bibRefCitation>
: fig. 24D)]; and its male G1 is slender and obviously tapering (
<figureCitation id="E583B0807568C645FE50FBACFDA74973" box="[508,635,1051,1077]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="6" pageNumber="144">Fig. 2HM</figureCitation>
) [vs. stout and only slightly tapering in
<taxonomicName id="BAB8D7867568C645FB98FBACFB754973" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[1076,1193,1051,1077]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="4FCC70177568C645FB98FBACFB754973" box="[1076,1193,1051,1077]" italics="true" pageId="6" pageNumber="144">Par. balssi</emphasis>
</taxonomicName>
(K.
<bibRefCitation id="1929D1F47568C645FB76FBACFA444973" author="Sakai, K. &amp; Turkay, M. &amp; Yang, S. - L." box="[1242,1432,1051,1077]" pageId="6" pageNumber="144" pagination="1 - 76" refId="ref10458" refString="Sakai, K., Turkay, M. &amp; Yang, S. - L. (2006) Revision of the Helice / Chasmagnathus complex (Crustacea: Decapoda: Brachyura). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 565, 1 - 76." type="journal article" year="2006">
Sakai
<emphasis id="4FCC70177568C645FAB3FBACFA934973" box="[1311,1359,1051,1077]" italics="true" pageId="6" pageNumber="144">et al</emphasis>
. 2006
</bibRefCitation>
: fig. 70a, b)].
</paragraph>
<paragraph id="7D07AC057568C645FF6BFBD4FB70483F" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="144">
<taxonomicName id="BAB8D7867568C645FF6BFBD4FE1E493B" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[199,450,1123,1149]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="4FCC70177568C645FF6BFBD4FE1E493B" box="[199,450,1123,1149]" italics="true" pageId="6" pageNumber="144">Parahelice ngankeeae</emphasis>
</taxonomicName>
can be differentiated from
<taxonomicName id="BAB8D7867568C645FD54FBD4FCAE493B" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[760,882,1123,1149]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="4FCC70177568C645FD54FBD4FCAE493B" box="[760,882,1123,1149]" italics="true" pageId="6" pageNumber="144">Par. daviei</emphasis>
</taxonomicName>
and
<taxonomicName id="BAB8D7867568C645FC07FBD4FB99493B" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[939,1093,1123,1149]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="pilimana">
<emphasis id="4FCC70177568C645FC07FBD4FB99493B" box="[939,1093,1123,1149]" italics="true" pageId="6" pageNumber="144">Par. pilimana</emphasis>
</taxonomicName>
by the lateral part of the male suborbital cristae and G1s. The lateral part of the suborbital crista comprises 1 significantly largest elliptical tubercle (without Y-shaped or particularly vertically inflated crest) and 23 smaller tubercles in
<taxonomicName id="BAB8D7867568C645FBD0FB1CFAF74983" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[1148,1323,1195,1221]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="4FCC70177568C645FBD0FB1CFAF74983" box="[1148,1323,1195,1221]" italics="true" pageId="6" pageNumber="144">Par. ngankeeae</emphasis>
</taxonomicName>
(
<figureCitation id="E583B0807568C645FA90FB1CFA484983" box="[1340,1428,1195,1221]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="6" pageNumber="144">Fig. 2B</figureCitation>
) [vs. elongated Y-shaped crest and 12 smaller tubercles in
<taxonomicName id="BAB8D7867568C645FC81FB78FC7A49AF" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[813,934,1231,1257]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="4FCC70177568C645FC81FB78FC7A49AF" box="[813,934,1231,1257]" italics="true" pageId="6" pageNumber="144">Par. daviei</emphasis>
</taxonomicName>
(K.
<bibRefCitation id="1929D1F47568C645FC76FB78FB4049AF" author="Sakai, K. &amp; Turkay, M. &amp; Yang, S. - L." box="[986,1180,1231,1257]" pageId="6" pageNumber="144" pagination="1 - 76" refId="ref10458" refString="Sakai, K., Turkay, M. &amp; Yang, S. - L. (2006) Revision of the Helice / Chasmagnathus complex (Crustacea: Decapoda: Brachyura). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 565, 1 - 76." type="journal article" year="2006">
Sakai
<emphasis id="4FCC70177568C645FB8CFB78FB8449AF" box="[1056,1112,1231,1257]" italics="true" pageId="6" pageNumber="144">et al.</emphasis>
2006
</bibRefCitation>
: fig. 66a, b;
<bibRefCitation id="1929D1F47568C645FA85FB78FF0F484B" author="Shih, H. - T. &amp; Hsu, J. - W. &amp; Li, J. - J. &amp; Ng, N. K. &amp; Lee, J. - H." pageId="6" pageNumber="144" pagination="249 - 265" refId="ref10696" refString="Shih, H. - T., Hsu, J. - W., Li, J. - J., Ng, N. K. &amp; Lee, J. - H. (2020) The identities of three species of Parahelice Sakai, Turkay &amp; Yang, 2006 (Crustacea: Brachyura: Varunidae) from the western Pacific, based on morphological and molecular evidence. Zootaxa, 4728 (2), 249 - 265. https: // doi. org / 10.11646 / zootaxa. 4728.2.6" type="journal article" year="2020">
Shih
<emphasis id="4FCC70177568C645FAC8FB78FA4149AF" box="[1380,1437,1231,1257]" italics="true" pageId="6" pageNumber="144">et al.</emphasis>
2020
</bibRefCitation>
: fig. 2C); and elongated, laterally, particularly vertically inflated crest and 2 smaller tubercles in
<taxonomicName id="BAB8D7867568C645FAA8FB44FA41484B" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1284,1437,1267,1293]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="pilimana">
<emphasis id="4FCC70177568C645FAA8FB44FA41484B" box="[1284,1437,1267,1293]" italics="true" pageId="6" pageNumber="144">Par. pilimana</emphasis>
</taxonomicName>
(K.
<bibRefCitation id="1929D1F47568C645FF6FFAA0FE584877" author="Sakai, K. &amp; Turkay, M. &amp; Yang, S. - L." box="[195,388,1303,1329]" pageId="6" pageNumber="144" pagination="1 - 76" refId="ref10458" refString="Sakai, K., Turkay, M. &amp; Yang, S. - L. (2006) Revision of the Helice / Chasmagnathus complex (Crustacea: Decapoda: Brachyura). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 565, 1 - 76." type="journal article" year="2006">
Sakai
<emphasis id="4FCC70177568C645FEA4FAA0FE9D4877" box="[264,321,1303,1329]" italics="true" pageId="6" pageNumber="144">et al.</emphasis>
2006
</bibRefCitation>
: fig. 65;
<bibRefCitation id="1929D1F47568C645FE4BFAA0FD424877" author="Shih, H. - T. &amp; Hsu, J. - W. &amp; Li, J. - J. &amp; Ng, N. K. &amp; Lee, J. - H." box="[487,670,1303,1329]" pageId="6" pageNumber="144" pagination="249 - 265" refId="ref10696" refString="Shih, H. - T., Hsu, J. - W., Li, J. - J., Ng, N. K. &amp; Lee, J. - H. (2020) The identities of three species of Parahelice Sakai, Turkay &amp; Yang, 2006 (Crustacea: Brachyura: Varunidae) from the western Pacific, based on morphological and molecular evidence. Zootaxa, 4728 (2), 249 - 265. https: // doi. org / 10.11646 / zootaxa. 4728.2.6" type="journal article" year="2020">
Shih
<emphasis id="4FCC70177568C645FD8EFAA0FD8F4877" box="[546,595,1303,1329]" italics="true" pageId="6" pageNumber="144">et al</emphasis>
. 2020
</bibRefCitation>
: fig. 4C)]; and the part near the tip of G1 is relatively stout and only slightly tapering in
<taxonomicName id="BAB8D7867568C645FEDDFA8CFDC24813" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[369,542,1339,1365]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="4FCC70177568C645FEDDFA8CFDC24813" box="[369,542,1339,1365]" italics="true" pageId="6" pageNumber="144">Par. ngankeeae</emphasis>
</taxonomicName>
(
<figureCitation id="E583B0807568C645FD81FA8CFD704813" box="[557,684,1339,1365]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="6" pageNumber="144">Fig. 2HM</figureCitation>
) [vs. relatively slender and significantly tapering in
<taxonomicName id="BAB8D7867568C645FB58FA8CFAB14813" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[1268,1389,1339,1365]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="4FCC70177568C645FB58FA8CFAB14813" box="[1268,1389,1339,1365]" italics="true" pageId="6" pageNumber="144">Par. daviei</emphasis>
</taxonomicName>
and
<taxonomicName id="BAB8D7867568C645FF3BFAE8FEEC483F" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[151,304,1375,1401]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="pilimana">
<emphasis id="4FCC70177568C645FF3BFAE8FEEC483F" box="[151,304,1375,1401]" italics="true" pageId="6" pageNumber="144">Par. pilimana</emphasis>
</taxonomicName>
(K.
<bibRefCitation id="1929D1F47568C645FECEFAE8FDFE483F" author="Sakai, K. &amp; Turkay, M. &amp; Yang, S. - L." box="[354,546,1375,1401]" pageId="6" pageNumber="144" pagination="1 - 76" refId="ref10458" refString="Sakai, K., Turkay, M. &amp; Yang, S. - L. (2006) Revision of the Helice / Chasmagnathus complex (Crustacea: Decapoda: Brachyura). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 565, 1 - 76." type="journal article" year="2006">
Sakai
<emphasis id="4FCC70177568C645FE0BFAE8FE05483F" box="[423,473,1375,1401]" italics="true" pageId="6" pageNumber="144">et al</emphasis>
. 2006
</bibRefCitation>
: figs. 68ac, 71ac;
<bibRefCitation id="1929D1F47568C645FCA4FAE8FC61483F" author="Shih, H. - T. &amp; Hsu, J. - W. &amp; Li, J. - J. &amp; Ng, N. K. &amp; Lee, J. - H." box="[776,957,1375,1401]" pageId="6" pageNumber="144" pagination="249 - 265" refId="ref10696" refString="Shih, H. - T., Hsu, J. - W., Li, J. - J., Ng, N. K. &amp; Lee, J. - H. (2020) The identities of three species of Parahelice Sakai, Turkay &amp; Yang, 2006 (Crustacea: Brachyura: Varunidae) from the western Pacific, based on morphological and molecular evidence. Zootaxa, 4728 (2), 249 - 265. https: // doi. org / 10.11646 / zootaxa. 4728.2.6" type="journal article" year="2020">
Shih
<emphasis id="4FCC70177568C645FCEEFAE8FCAF483F" box="[834,883,1375,1401]" italics="true" pageId="6" pageNumber="144">et al</emphasis>
. 2020
</bibRefCitation>
: figs. 2EH, 4CH)].
</paragraph>
<paragraph id="7D07AC057568C645FF6BFA34FC854B6B" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="144">
<taxonomicName id="BAB8D7867568C645FF6BFA34FE1948DB" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[199,453,1411,1437]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="4FCC70177568C645FF6BFA34FE1948DB" box="[199,453,1411,1437]" italics="true" pageId="6" pageNumber="144">Parahelice ngankeeae</emphasis>
</taxonomicName>
and
<taxonomicName id="BAB8D7867568C645FDA8FA34FD5D48DB" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[516,641,1411,1437]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="4FCC70177568C645FDA8FA34FD5D48DB" box="[516,641,1411,1437]" italics="true" pageId="6" pageNumber="144">Par. pilosa</emphasis>
</taxonomicName>
can be separated by the male suborbital cristae. The lateral part of the suborbital cristae in
<taxonomicName id="BAB8D7867568C645FE00FA10FD874887" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[428,603,1447,1473]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="4FCC70177568C645FE00FA10FD874887" box="[428,603,1447,1473]" italics="true" pageId="6" pageNumber="144">Par. ngankeeae</emphasis>
</taxonomicName>
is composed of 1 largest elliptical tubercle and 23 tubercles, which are significantly smaller and less convex than the largest elliptical tubercle [vs. crest ending with a slightly inflated part, followed by 2 tubercles, which are only slightly smaller and less convex than (or equal to) the inflated part in
<taxonomicName id="BAB8D7867568C645FAC2FA58FF074B6B" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="4FCC70177568C645FAC2FA58FF074B6B" italics="true" pageId="6" pageNumber="144">Par. pilosa</emphasis>
</taxonomicName>
(K.
<bibRefCitation id="1929D1F47568C645FEA2F9A4FE124B6B" author="Sakai, K. &amp; Turkay, M. &amp; Yang, S. - L." box="[270,462,1555,1581]" pageId="6" pageNumber="144" pagination="1 - 76" refId="ref10458" refString="Sakai, K., Turkay, M. &amp; Yang, S. - L. (2006) Revision of the Helice / Chasmagnathus complex (Crustacea: Decapoda: Brachyura). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 565, 1 - 76." type="journal article" year="2006">
Sakai
<emphasis id="4FCC70177568C645FEFFF9A4FE594B6B" box="[339,389,1555,1581]" italics="true" pageId="6" pageNumber="144">et al</emphasis>
. 2006
</bibRefCitation>
: fig. 61;
<bibRefCitation id="1929D1F47568C645FD9DF9A4FD3A4B6B" author="Shih, H. - T. &amp; Hsu, J. - W. &amp; Li, J. - J. &amp; Ng, N. K. &amp; Lee, J. - H." box="[561,742,1555,1581]" pageId="6" pageNumber="144" pagination="249 - 265" refId="ref10696" refString="Shih, H. - T., Hsu, J. - W., Li, J. - J., Ng, N. K. &amp; Lee, J. - H. (2020) The identities of three species of Parahelice Sakai, Turkay &amp; Yang, 2006 (Crustacea: Brachyura: Varunidae) from the western Pacific, based on morphological and molecular evidence. Zootaxa, 4728 (2), 249 - 265. https: // doi. org / 10.11646 / zootaxa. 4728.2.6" type="journal article" year="2020">
Shih
<emphasis id="4FCC70177568C645FDC7F9A4FD414B6B" box="[619,669,1555,1581]" italics="true" pageId="6" pageNumber="144">et al</emphasis>
. 2020
</bibRefCitation>
: fig. 6C)].
</paragraph>
<paragraph id="7D07AC057568C645FF6BF980FA404BA7" blockId="6.[151,1437,151,1761]" pageId="6" pageNumber="144">
The female
<taxonomicName id="BAB8D7867568C645FEFDF980FDDC4B17" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[337,512,1591,1617]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="4FCC70177568C645FEFDF980FDDC4B17" box="[337,512,1591,1617]" italics="true" pageId="6" pageNumber="144">Par. ngankeeae</emphasis>
</taxonomicName>
is morphologically remarkably similar to
<taxonomicName id="BAB8D7867568C645FC4AF980FB824B17" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[998,1118,1591,1617]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="balssi">
<emphasis id="4FCC70177568C645FC4AF980FB824B17" box="[998,1118,1591,1617]" italics="true" pageId="6" pageNumber="144">Par. balssi</emphasis>
</taxonomicName>
,
<taxonomicName id="BAB8D7867568C645FBC3F980FB364B17" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[1135,1258,1591,1617]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="daviei">
<emphasis id="4FCC70177568C645FBC3F980FB364B17" box="[1135,1258,1591,1617]" italics="true" pageId="6" pageNumber="144">Par. daviei</emphasis>
</taxonomicName>
,
<taxonomicName id="BAB8D7867568C645FB57F980FA4A4B17" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[1275,1430,1591,1617]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="pilimana">
<emphasis id="4FCC70177568C645FB57F980FA4A4B17" box="[1275,1430,1591,1617]" italics="true" pageId="6" pageNumber="144">Par. pilimana</emphasis>
</taxonomicName>
, and
<taxonomicName id="BAB8D7867568C645FF65F9ECFE994B33" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[201,325,1627,1653]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="4FCC70177568C645FF65F9ECFE994B33" box="[201,325,1627,1653]" italics="true" pageId="6" pageNumber="144">Par. pilosa</emphasis>
</taxonomicName>
, but can be distinguished by the suborbital cristae. In female
<taxonomicName id="BAB8D7867568C645FBB8F9ECFB1F4B33" authorityName="Hsu &amp; Shih" authorityYear="2024" box="[1044,1219,1627,1653]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="ngankeeae">
<emphasis id="4FCC70177568C645FBB8F9ECFB1F4B33" box="[1044,1219,1627,1653]" italics="true" pageId="6" pageNumber="144">Par. ngankeeae</emphasis>
</taxonomicName>
, the lateral part of the suborbital crista comprises 2 or 3 larger and vertically inflated (significantly larger than the tubercles in the mesial part) and 1 or 2 small, rounded tubercles (
<figureCitation id="E583B0807568C645FD18F914FCD54BFB" box="[692,777,1699,1725]" captionStart="FIGURE 2" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[205,1381,189,1883]" captionTargetId="figure-35@5.[197,1389,181,1885]" captionTargetPageId="5" captionText="FIGURE 2. Parahelice ngankeeae sp. nov. A, B, D, F, HM, male (24.3 × 19.1 mm, holotype, ZRC 2023.0238); C, E, G, N, female (15.6 × 12.8 mm, paratype, ZRC 2023.2043). A, carapace; B, left male suborbital crista; C, left female suborbital crista; D, outer view of male left cheliped; E, outer view of female left cheliped; F, male pleon; G, female pleon; HM, male right G1 (H, I, dorsal view; J, K, dorsolateral view; L, M, ventral view); N, right vulva." figureDoi="http://doi.org/10.5281/zenodo.12724052" httpUri="https://zenodo.org/record/12724052/files/figure.png" pageId="6" pageNumber="144">Fig. 2C</figureCitation>
) [vs. several isomorphically interspaced tubercles without a significantly larger one in
<emphasis id="4FCC70177568C645FE14F970FC884BA7" box="[440,852,1735,1761]" italics="true" pageId="6" pageNumber="144">
<taxonomicName id="BAB8D7867568C645FE14F970FDF34BA7" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[440,559,1735,1761]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="balssi">Par. balssi</taxonomicName>
,
<taxonomicName id="BAB8D7867568C645FD96F970FD684BA7" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[570,692,1735,1761]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="daviei">Par. daviei</taxonomicName>
,
<taxonomicName id="BAB8D7867568C645FD12F970FC884BA7" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" box="[702,852,1735,1761]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="pilimana">Par. pilimana</taxonomicName>
</emphasis>
and
<taxonomicName id="BAB8D7867568C645FC25F970FBDC4BA7" baseAuthorityName="K. Sakai, Turkay &amp; Yang" baseAuthorityYear="2006" box="[905,1024,1735,1761]" class="Malacostraca" family="Varunidae" genus="Parahelice" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="144" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="4FCC70177568C645FC25F970FBDC4BA7" box="[905,1024,1735,1761]" italics="true" pageId="6" pageNumber="144">Par. pilosa</emphasis>
</taxonomicName>
(
<bibRefCitation id="1929D1F47568C645FBA2F970FB1E4BA7" author="Shih, H. - T. &amp; Hsu, J. - W. &amp; Li, J. - J. &amp; Ng, N. K. &amp; Lee, J. - H." box="[1038,1218,1735,1761]" pageId="6" pageNumber="144" pagination="249 - 265" refId="ref10696" refString="Shih, H. - T., Hsu, J. - W., Li, J. - J., Ng, N. K. &amp; Lee, J. - H. (2020) The identities of three species of Parahelice Sakai, Turkay &amp; Yang, 2006 (Crustacea: Brachyura: Varunidae) from the western Pacific, based on morphological and molecular evidence. Zootaxa, 4728 (2), 249 - 265. https: // doi. org / 10.11646 / zootaxa. 4728.2.6" type="journal article" year="2020">
Shih
<emphasis id="4FCC70177568C645FBE5F970FBA54BA7" box="[1097,1145,1735,1761]" italics="true" pageId="6" pageNumber="144">et al</emphasis>
. 2020
</bibRefCitation>
: figs. 2D, 4D, 6D)].
</paragraph>
</subSubSection>
</treatment>
</document>