treatments-xml/data/F4/A5/69/F4A56930A8FB03130B0D3D7A8C7EBD9F.xml
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<document id="F821EB525571AEF33271AE3DA8B89B37" ID-DOI="http://dx.doi.org/10.3897/AfrInvertebr.58.12782" ID-GBIF-Dataset="0c3635ae-8b7b-4559-ab86-20908d81041f" ID-Pensoft-Pub="2305-2562-2-21" ID-ZooBank="2B0E4FE5663944F6A5EF4DCB034BAA0A" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="2305-2562-2-21" ModsDocOrigin="African Invertebrates 58 (2)" ModsDocTitle="Review of the ant genus Nesomyrmex Wheeler, 1910 (Hymenoptera, Formicidae, Myrmicinae) from the Arabian Peninsula" checkinTime="1552155284496" checkinUser="pensoft" docAuthor="Sharaf, Mostafa R., Akbar, Shahid Ali, Aldawood, Abdulrahman S. &amp; Garcia, Francisco Hita" docDate="2017" docId="F4A56930A8FB03130B0D3D7A8C7EBD9F" docLanguage="en" docName="AfrInvert 58(2): 21-37" docOrigin="African Invertebrates 58 (2)" docSource="http://dx.doi.org/10.3897/AfrInvertebr.58.12782" docTitle="Nesomyrmex humerosus Emery 1896" docType="treatment" docVersion="6" lastPageNumber="28" masterDocId="124AFFD7FFAF7E75447BFE62FFDCFF9A" masterDocTitle="Review of the ant genus Nesomyrmex Wheeler, 1910 (Hymenoptera, Formicidae, Myrmicinae) from the Arabian Peninsula" masterLastPageNumber="37" masterPageNumber="21" pageNumber="26" updateTime="1701316593355" updateUser="plazi">
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<mods:title id="D9B04B459955818C104213DA207605ED">Review of the ant genus Nesomyrmex Wheeler, 1910 (Hymenoptera, Formicidae, Myrmicinae) from the Arabian Peninsula</mods:title>
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<mods:namePart id="7B1B24C5758C4684E4437C4242CFD4F5">Sharaf, Mostafa R.</mods:namePart>
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<mods:namePart id="3E58502434CAF30A6F59FC52221B419E">Akbar, Shahid Ali</mods:namePart>
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<mods:namePart id="C750E3FC1DC6CE61C683DB19FEA8D754">Aldawood, Abdulrahman S.</mods:namePart>
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<mods:namePart id="11A300AF29A6437E12C17DADD759354B">Garcia, Francisco Hita</mods:namePart>
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<mods:title id="305D5B2A9803B1E159E79494FBC2D2F8">African Invertebrates</mods:title>
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<subSubSection id="5E3E41E98F4D6E138192F432B5E3C153" pageId="5" pageNumber="26" type="nomenclature">
<paragraph id="57CFB4C2ED5D6760902498F36152E245" pageId="5" pageNumber="26">
<taxonomicName id="A7F267BCD1BA85DD4BCCDE6C18FC514F" authority="Emery, 1896" authorityName="Emery" authorityYear="1896" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex" order="Hymenoptera" pageId="5" pageNumber="26" phylum="Arthropoda" rank="species" species="humerosus">Nesomyrmex humerosus (Emery, 1896)</taxonomicName>
Figs 2A, B, 5
</paragraph>
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<subSubSection id="BEB236503697A22E354B6668764F3E6F" pageId="6" pageNumber="27" type="reference_group">
<paragraph id="C98208299766FC9A8F2AE2C565B03FC7" pageId="6" pageNumber="27">
<taxonomicName id="6A5701CB10BFE474564A484AE4E6D22A" class="Insecta" family="Formicidae" genus="Leptothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leptothorax (Goniothorax) humerosus" order="Hymenoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="humerosus" subGenus="Goniothorax">
<pageBreakToken id="72643901CAC2563628D431FE1B17BEA7" pageId="6" pageNumber="27" start="start">Leptothorax</pageBreakToken>
(Goniothorax) humerosus
</taxonomicName>
Emery, 1896: 62 (w.) EAST AFRICA (no locality given, very likely KENYA). [Combination in
<taxonomicName id="F681EEF78EC37411615BF35C2D94C75C" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex" order="Hymenoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="genus">Nesomyrmex</taxonomicName>
:
<bibRefCitation id="7A0AA5261C9079DD362FEBBF78994544" author="Bolton, B" journalOrPublisher="Memoirs of the American Entomological Institute" pageId="9" pageNumber="30" pagination="1 - 370" title="Synopsis and classification of Formicidae." volume="71" year="2003">Bolton 2003</bibRefCitation>
: 272. See also:
<bibRefCitation id="C1FC4A784F10F955B060A715E41055C6" author="Bolton, B" journalOrPublisher="Bulletin of the British Museum (Natural History) Entomology" pageId="9" pageNumber="30" pagination="245 - 307" title="A revision of six minor genera of Myrmicinae (Hymenoptera: Formicidae) in the Ethiopian zoogeographical region." volume="43" year="1982">Bolton 1982</bibRefCitation>
: 329;
<bibRefCitation id="12AFAFE6AC109D9A423923A4995966D9" author="Hita, Garcia F" journalOrPublisher="European Journal of Taxonomy" pageId="10" pageNumber="31" pagination="1 - 31" title="Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique." url="http://dx.doi.org/10.5852/ejt.2017.258" volume="258" year="2017">Hita Garcia et al. 2017</bibRefCitation>
: 8].
</paragraph>
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<subSubSection id="1DBDB27CC8F21C1A873E67714B36A7AF" pageId="6" pageNumber="27" type="materials_examined">
<paragraph id="2818847251952F72A27DDED7003ADF90" pageId="6" pageNumber="27">Material examined.</paragraph>
<paragraph id="609F15317F8781C823306A92AEAB05F3" pageId="6" pageNumber="27">KENYA: Coast Province, Malindi, Arabuko Sokoke Forest, -3.321, 39.929, 50 m, coastal dry forest, VI.2009 (F. Hita Garcia &amp; G. Fischer) (FHGC: 3w, 1q).</paragraph>
</subSubSection>
<subSubSection id="B5300D91BE3D79F1CC2E5129B91D1113" pageId="6" pageNumber="27" type="diagnosis">
<paragraph id="0888780EA575770770F8EF964400696F" pageId="6" pageNumber="27">Diagnosis.</paragraph>
<paragraph id="D2229A2CF255E04E27412250985D432D" pageId="6" pageNumber="27">
The following character combination distinguishes
<taxonomicName id="E0D92AC7561F48E66F7840AD5FABC756" lsidName="N. humerosus" pageId="6" pageNumber="27" rank="species" species="humerosus">N. humerosus</taxonomicName>
from the other members of the group in the Arabian Peninsula: anterior clypeal lobe short, flat-margined, and never convex, with small median triangular projection; clypeus without median longitudinal carina; pronotum anterodorsally sharply marginate, with sharp, dentate corners; in profile mesosomal outline conspicuously concave; petiole barrel-shaped with very weakly developed, short and triangular petiolar node.
</paragraph>
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<subSubSection id="3F43C62EE921B0ECD346CE2A9C414E14" pageId="6" pageNumber="27" type="description">
<paragraph id="98695BF30C152D28085D2AC4E62126BB" pageId="6" pageNumber="27">Worker measurements</paragraph>
<paragraph id="06F53912A97701661052F0CA9D9D368C" pageId="6" pageNumber="27">(n=3). HL 0.74-0.75; HW 0.70-0.71; SL 0.58-0.59; EL 0.27-0.28; PH 0.34-0.35; PW 0.52-0.53; WL 0.89-0.92; PSL 0.19-0.20; PTL 0.12-0.13; PTH 0.24-0.25; PTW 0.21-0.22; PPL 0.15-0.16; PPH 0.21-0.22; PPW 0.33-0.34; OI 39; SI 83; CI 95; DMI 58; LMI 38; PSLI 26-27; LPeI 50-52; DPeI 169-175; LPpI 71-73; DPpI 213-220; PPI 155-157.</paragraph>
</subSubSection>
<subSubSection id="90C26B40D9110FB98B730C70741A28E4" pageId="6" pageNumber="27" type="distribution">
<paragraph id="7E43BC7F21E5E31B225B39A7EF2E7A67" pageId="6" pageNumber="27">Distribution.</paragraph>
<paragraph id="425E57FAD2E140F360528798127EF8B9" pageId="6" pageNumber="27">
<taxonomicName id="4E9AA55D392B644A308DED1185B6EE70" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex humerosus" order="Hymenoptera" pageId="6" pageNumber="27" phylum="Arthropoda" rank="species" species="humerosus">Nesomyrmex humerosus</taxonomicName>
is currently only known to occur in Kenya, Tanzania, and Yemen. Very little is known about the biology of the species. Rarely collected species seems to live on vegetation but has also been sampled from the ground (
<bibRefCitation id="93ED37A9591A4198464A5CE82B38FCB6" author="Hita, Garcia F" journalOrPublisher="European Journal of Taxonomy" pageId="10" pageNumber="31" pagination="1 - 31" title="Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique." url="http://dx.doi.org/10.5852/ejt.2017.258" volume="258" year="2017">Hita Garcia et al. 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CD6E9D159D4B16324EE301A1543B8E77" pageId="7" pageNumber="28" type="comments">
<paragraph id="19959C5266B512058B204629F3F20D31" pageId="7" pageNumber="28">
<pageBreakToken id="81EDE53ACA6B82EF4FB0277E71B60D73" pageId="7" pageNumber="28" start="start">Comments</pageBreakToken>
.
</paragraph>
<paragraph id="391C7C2157FCBEB190EE17EEB8D38E0D" pageId="7" pageNumber="28">
<taxonomicName id="65ADFCC010609FBE8AAF4F279316F9A8" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex humerosus" order="Hymenoptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="humerosus">Nesomyrmex humerosus</taxonomicName>
is the only member of the
<taxonomicName id="2802B417DAAAD35D99E29A9F751F4544" lsidName="N. humerosus" pageId="7" pageNumber="28" rank="species" species="humerosus">N. humerosus</taxonomicName>
group and possesses an unusual character combination for an Afrotropical species due to its flat anterior clypeal margin, dentate pronotum, and barrel-shaped petiole. For more details see
<bibRefCitation id="D8C2585F85E6A0700BD988EB2CFE6840" author="Hita, Garcia F" journalOrPublisher="European Journal of Taxonomy" pageId="10" pageNumber="31" pagination="1 - 31" title="Taxonomy of the ant genus Nesomyrmex Wheeler (Formicidae, Myrmicinae) in the Afrotropical region, with a review of current species groups and description of a new species of the N. angulatus group from Mozambique." url="http://dx.doi.org/10.5852/ejt.2017.258" volume="258" year="2017">Hita Garcia et al. (2017)</bibRefCitation>
. This species has been infrequently collected and is apparently known from four separate collections. More intensive sampling in East Africa and Yemen should yield more material. It is noteworthy that we were not able to examine any material of this species from Yemen and all our knowledge of the species is based on literature (
<bibRefCitation id="34086112C56B1F83F90388EB7A7BD953" author="Collingwood, CA" journalOrPublisher="Fauna of Saudi Arabia" pageId="10" pageNumber="31" pagination="300 - 385" title="Formicidae (Insecta: Hymenoptera) of Saudi Arabia (part 2)." volume="15" year="1996">Collingwood and Agosti 1996</bibRefCitation>
,
<bibRefCitation id="030717C4D14E90A1E640B735A441F45D" author="Borowiec, L" journalOrPublisher="Genus (Wroclaw)" pageId="9" pageNumber="30" pagination="1 - 340" title="Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae)." volume="25" year="2014">Borowiec 2014</bibRefCitation>
).
</paragraph>
<caption id="F2F83B6D8A01EEB261A28E643704B5C9" pageId="7" pageNumber="28">
<paragraph id="5B1C16DEC55CF3764EC037659421F8BB" pageId="7" pageNumber="28">
Figure 5.
<taxonomicName id="E1261CAD9CEA01147FA8D5FEB31240DE" class="Insecta" family="Formicidae" genus="Nesomyrmex" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nesomyrmex humerosus" order="Hymenoptera" pageId="7" pageNumber="28" phylum="Arthropoda" rank="species" species="humerosus">Nesomyrmex humerosus</taxonomicName>
(CASENT0906196 - Antweb, Estella Ortega 2013). A Body in profile B Body in dorsal view C Head in full-face view.
</paragraph>
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