treatments-xml/data/F1/F6/00/F1F60027744556EF968FBDD4C67717A7.xml
2024-06-21 12:57:21 +02:00

466 lines
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XML

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1172.107112" ID-Pensoft-Pub="1313-2970-1172-117" ID-Pensoft-UUID="7CDB8E4C94ED5FC8BE3DE34846860463" ID-ZooBank="C8B5DE1C0482491DA61C4BFB55C93A13" ModsDocID="1313-2970-1172-117" checkinTime="1690372417257" checkinUser="pensoft" docAuthor="Kosyan, Armine, Zamani, Alireza &amp; Marusik, Yuri M." docDate="2023" docId="F1F60027744556EF968FBDD4C67717A7" docLanguage="en" docName="ZooKeys 1172: 117-130" docOrigin="ZooKeys 1172" docPubDate="2023-07-25" docSource="http://dx.doi.org/10.3897/zookeys.1172.107112" docTitle="Harpactea armenica Dunin 1989" docType="treatment" docVersion="1" id="7CDB8E4C94ED5FC8BE3DE34846860463" lastPageNumber="117" masterDocId="7CDB8E4C94ED5FC8BE3DE34846860463" masterDocTitle="Notes on Dysderidae (Arachnida, Araneae) of Armenia and Iran, with new species and records" masterLastPageNumber="130" masterPageNumber="117" pageNumber="117" updateTime="1690372417257" updateUser="pensoft">
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<mods:titleInfo id="AA366A05B8CBB097967FDB15E869F8E8">
<mods:title id="FE99F005196B694DD6795756E1E52E97">Notes on Dysderidae (Arachnida, Araneae) of Armenia and Iran, with new species and records</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="52517A69C222BCA864F0F00C5955F01F">Kosyan, Armine</mods:namePart>
<mods:nameIdentifier id="82BD1072715E8433D7D49B02F3026173" type="ORCID">https://orcid.org/0000-0001-5692-5369</mods:nameIdentifier>
<mods:affiliation id="ABDD906B86D77172E7052B9EEB0C99AB">Faculty of Biology, Yerevan State University, Yerevan, Armenia</mods:affiliation>
</mods:name>
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<mods:namePart id="E1ED8130BF5F56541382B91F59CC00EE">Zamani, Alireza</mods:namePart>
<mods:nameIdentifier id="7571CDBFE7EA5DB67EBB0B41446E521B" type="ORCID">https://orcid.org/0000-0002-8084-9666</mods:nameIdentifier>
<mods:affiliation id="F737FAEE98D67486396E32D9FCE9CE60">Zoological Museum, Biodiversity Unit, FI- 20014 University of Turku, Finland</mods:affiliation>
<mods:nameIdentifier id="2E4C4DF2AAD705E37846A06B09C21753" type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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<mods:namePart id="2232811A0104EB87F3868169D5C7D341">Marusik, Yuri M.</mods:namePart>
<mods:nameIdentifier id="9778D113A44EEE4B3FBDE0CD494D04C9" type="ORCID">https://orcid.org/0000-0002-4499-5148</mods:nameIdentifier>
<mods:affiliation id="3A6D02109FE7205B4AE11158662C7AEB">Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa &amp; Altai State University, Lenina Pr., 61, Barnaul, RF- 656049, Russia</mods:affiliation>
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<mods:date id="5D76D17674D051BA74553799121EC9C3">2023</mods:date>
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<mods:number id="CD5B5BFEDF58A72DCF4F935479B1496A">2023-07-25</mods:number>
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<mods:number id="1C62D151F4F0D078F131264C2A3E5ED5">1172</mods:number>
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<mods:start id="3A925CCD8DA2E501962A11D587170F33">117</mods:start>
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<mods:classification id="8C472D2E2D1EF7E259EEDA5887B7B4BF">journal article</mods:classification>
<mods:identifier id="188705B6698361C59756EFF7EE372A88" type="DOI">http://dx.doi.org/10.3897/zookeys.1172.107112</mods:identifier>
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<treatment id="F1F60027744556EF968FBDD4C67717A7" LSID="urn:lsid:plazi:treatment:F1F60027744556EF968FBDD4C67717A7" httpUri="http://treatment.plazi.org/id/F1F60027744556EF968FBDD4C67717A7" lastPageNumber="117" pageId="0" pageNumber="117">
<subSubSection id="DE35B24BEC8008B2A701D4E07FB3C2D5" pageId="0" pageNumber="117" type="nomenclature">
<paragraph id="FC74948E34E3C62B725ABFECF045E429" pageId="0" pageNumber="117">
<taxonomicName id="F15F4697EF8D4B7553DFB3BB39ABA038" ID-CoL="6LCHY" LSID="F1F60027-7445-56EF-968F-BDD4C67717A7" authority="Dunin, 1989" authorityName="Dunin" authorityYear="1989" class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea armenica" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="armenica">Harpactea armenica Dunin, 1989</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="F5175845EB16818AAAD3F3846FA1744F" pageId="0" pageNumber="117" type="description">
<paragraph id="9F25BC86073CD025C4CCAEF8CF482249" pageId="0" pageNumber="117">
<figureCitation id="DB2F6F33785CBB5C4AE1D6A1269B8627" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Habitus of Harpactea armenica, male (A) and female (B), dorsal." figureDoi="10.3897/zookeys.1172.107112.figure8" httpUri="https://binary.pensoft.net/fig/883883" pageId="0" pageNumber="117">Figs 8A, B</figureCitation>
<figureCitation id="5A5516434B480B7FB4D5BD29E4DA4F45" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Copulatory organs of Harpactea armenica A-C bulb, prolateral, retrolateral and ventral D endogyne, dorsal. Abbreviations: Ad - anterior diverticulum, Eb - base of embolus, Re - receptacle. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1172.107112.figure9" httpUri="https://binary.pensoft.net/fig/883884" pageId="0" pageNumber="117">, 9A-D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="E10082667FEB751251D171F912C3BF2D" pageId="0" pageNumber="117" type="reference_group">
<paragraph id="44A0082DE81090FD95DCF135433F7F90" pageId="0" pageNumber="117">
<taxonomicName id="F56016154CFF7B7488EED2F52F6E242B" authorityName="Dunin" authorityYear="1989" class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea armenica" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="armenica">Harpactea armenica</taxonomicName>
Dunin, 1989: 142, figs 1-3 (♂♀).
</paragraph>
<paragraph id="E5617BA86EA092B8620A2706097CD7E8" pageId="0" pageNumber="117">
<taxonomicName id="51F272D83A5DA1819AADCAE5A2915901" authorityName="Dunin" authorityYear="1989" class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea armenica" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="armenica">Harpactea armenica</taxonomicName>
:
<bibRefCitation id="EDAB3C95E3178F39C9EE46C2803CDE16" author="Dunin, PM" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="117" pagination="35 - 76" refId="B11" refString="Dunin, PM, 1992. The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta 1 (3): 35 - 76" title="The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae)." volume="1" year="1992">Dunin 1992</bibRefCitation>
: 68, fig. 13 (♂♀).
</paragraph>
</subSubSection>
<subSubSection id="19220E2B21D738B5023AF27C5335BCDF" pageId="0" pageNumber="117" type="materials_examined">
<paragraph id="E462526A7FAE0D75C60A774E15BD09E0" pageId="0" pageNumber="117">Materials examined.</paragraph>
<paragraph id="2DB0131E68ADEE2762B873A724D85160" pageId="0" pageNumber="117">
<materialsCitation id="CC6E3091B4C2164181CB5E8EA7809048" collectingDate="2021-05-16" collectorName="Y. M. Marusik" country="Armenia" elevation="2200" latitude="40.383335" location="Aragates Mt." longLatPrecision="1163" longitude="44.216667" specimenCount="87" specimenCount-female="34" specimenCount-juvenile="29" specimenCount-male="24" stateProvince="Aragatsotn Province">
<collectingCountry id="D03747C171167D2F33F29234AB44190C" name="Armenia">Armenia</collectingCountry>
:
<emphasis id="0265D0B2821B0E6B217CFEF9B06ABA60" italics="true" pageId="0" pageNumber="117">
<collectingRegion id="B9ED8D1C7D02D5CBF25EC606B8D1A74C" country="Armenia" name="Aragatsotn">Aragatsotn Province</collectingRegion>
</emphasis>
:
<specimenCount id="FA23C2F7AB95ADC053B54FDBFD1107AB" type="male">24♂</specimenCount>
<specimenCount id="3DF0377AB637F3D590C12AF802F1713C" type="female">34♀</specimenCount>
<specimenCount id="40C86B4B156443F4B6EB10FE3A5BCDEB" type="generic">29 juv.</specimenCount>
(ZMMU), foothills of
<location id="1E46F6AFDCB673F6CF4B104F848E4ED0" LSID="urn:lsid:plazi:treatment:F1F60027744556EF968FBDD4C67717A7:1E46F6AFDCB673F6CF4B104F848E4ED0" country="Armenia" latitude="40.383335" longLatPrecision="1163" longitude="44.216667" name="Aragates Mt." stateProvince="Aragatsotn Province">Aragates Mt.</location>
,
<geoCoordinate id="2DCF00E281E2F115CE1B6446A08B20E0" degrees="40" direction="north" minutes="23" orientation="latitude" precision="925" value="40.383335">40°23'N</geoCoordinate>
,
<geoCoordinate id="71F8825100F48E83C06CD81A722EC5AC" degrees="44" direction="east" minutes="13" orientation="longitude" precision="925" value="44.216667">44°13'E</geoCoordinate>
,
<quantity id="F39C48620E003FA5FF1A865888FF54EE" metricMagnitude="3" metricUnit="m" metricValue="2.2" unit="m" value="2200.0">
<elevation id="CABC4ADB273AC2AB2C7CE1C49BABDBA8" metricMagnitude="3" metricUnit="m" metricValue="2.2" unit="m" value="2200.0">2200 m</elevation>
</quantity>
,
<collectingDate id="44F439E27AD11608882E482AB7BC028E" value="2021-05-16">16.05.2021</collectingDate>
(
<collectorName id="DE5761EBC4858614FCEFE19F2A1C169D">Y.M. Marusik</collectorName>
)
</materialsCitation>
;
<materialsCitation id="A6536B2699E1EF5C0043AABD2279076C" collectingDate="2021-05-06" collectingDateMax="2021-05-17" collectingDateMin="2021-05-06" collectorName="Y. M. Marusik, Garni" country="Armenia" elevation="1040" latitude="40.108887" location="Azat river" longLatPrecision="19" longitude="44.732502" municipality="Urtsadzor Vill." specimenCount="10" specimenCount-female="3" specimenCount-male="7" stateProvince="Ararat Province">
<emphasis id="B221280957B52320E716ADC668F8B427" italics="true" pageId="0" pageNumber="117">
<collectingRegion id="C23020456109B79CF78D0E1D43C5B7C7" country="Armenia" name="Ararat">Ararat Province</collectingRegion>
</emphasis>
:
<specimenCount id="2847050F8E2C466ADE2D3A6324E48B25" type="male">1♂</specimenCount>
(ZMMU), env. of
<collectingMunicipality id="D09C8D68233241DDC5DF43AE3AF1BB58">Urtsadzor Vill.</collectingMunicipality>
,
<geoCoordinate id="A6D4FAD086BFA927C7A507E83ECBDAD1" degrees="39" direction="north" minutes="55" orientation="latitude" precision="15" seconds="26" value="39.92389">39°55'26&quot;N</geoCoordinate>
,
<geoCoordinate id="3DF253B13D41BAD2A6828698E74671D3" degrees="44" direction="east" minutes="48" orientation="longitude" precision="15" seconds="53" value="44.81472">44°48'53&quot;E</geoCoordinate>
,
<quantity id="7174F9EBB599967DD11E5F8E0D87CD02" metricMagnitude="3" metricUnit="m" metricValue="1.04" unit="m" value="1040.0">
<elevation id="301DC7C6514535B4913E4B15228C460B" metricMagnitude="3" metricUnit="m" metricValue="1.04" unit="m" value="1040.0">1040 m</elevation>
</quantity>
,
<collectingDate id="CA3CA7ECCAD87779F206AC79DE647C39" value="2021-05-06">6.05.2021</collectingDate>
(
<collectorName id="501FE090F4CAD26E86A67E4DED786413">Y.M. Marusik</collectorName>
);
<specimenCount id="3C0992AC27A1CFB133E704EA2688D9C9" type="male">1♂</specimenCount>
<specimenCount id="326FCEF09E0753DE9E4EB2FE566B0283" type="female">2♀</specimenCount>
(ZMMU), env. of Urtsalanj Vill.,
<geoCoordinate id="B346D7B7480FC48B82634D35188D6550" degrees="39" direction="north" minutes="49" orientation="latitude" precision="925" value="39.816666">39°49'N</geoCoordinate>
,
<geoCoordinate id="B80E59228D61EC47F75D87B535804677" degrees="44" direction="east" minutes="59" orientation="longitude" precision="925" value="44.983334">44°59'E</geoCoordinate>
,
<quantity id="D09819977977738CEBC36F8B1665F60C" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">
<elevation id="6D31F55F2AE961E4ED9136F93D76B84B" metricMagnitude="3" metricUnit="m" metricValue="1.8" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<collectingDate id="D9A58C97AB0F0BFA23956F93189E617E" value="2021-05-11">11.05.2021</collectingDate>
(
<collectorName id="B397AD744DD1B0E9B00DD7223C34B964">Y.M. Marusik</collectorName>
);
<specimenCount id="2E39FE69F1026E5B41189B3F361859DE" type="male">5♂</specimenCount>
<specimenCount id="4F18049ECD9542528D29288E30893C97" type="female">1♀</specimenCount>
(ZMMU),
<collectorName id="8D7B1170BD138C615A848391B8E4316B">Garni</collectorName>
gorge,
<location id="6855E676E11A9A4F41159CF78F1F89A2" LSID="urn:lsid:plazi:treatment:F1F60027744556EF968FBDD4C67717A7:6855E676E11A9A4F41159CF78F1F89A2" country="Armenia" latitude="40.108887" longLatPrecision="19" longitude="44.732502" municipality="Urtsadzor Vill." name="Azat river" stateProvince="Ararat Province">Azat river</location>
,
<geoCoordinate id="E2F782A01C282212B8EB70C9D841E06A" degrees="40" direction="north" minutes="06" orientation="latitude" precision="15" seconds="32" value="40.108887">40°06'32&quot;N</geoCoordinate>
,
<geoCoordinate id="E5B2BA543DD3C7286711C2BCFA40500B" degrees="44" direction="east" minutes="43" orientation="longitude" precision="15" seconds="57" value="44.732502">44°43'57&quot;E</geoCoordinate>
,
<quantity id="933108387E31245D6D80E8D237741259" metricMagnitude="3" metricUnit="m" metricValue="1.24" unit="m" value="1240.0">
<elevation id="EB188C06E472ED4FA6B1602D9E078285" metricMagnitude="3" metricUnit="m" metricValue="1.24" unit="m" value="1240.0">1240 m</elevation>
</quantity>
,
<collectingDate id="690C65B05812787C2C142806364C52FF" value="2021-05-17">17.05.2021</collectingDate>
(
<collectorName id="862815EE464412E52CA51BFA5FE3AC71">Y.M. Marusik</collectorName>
)
</materialsCitation>
;
<materialsCitation id="BC82E86D58FDBBE87DB32FFDD8E2A620" collectingDate="2021-05-07" collectingDateMax="2021-05-15" collectingDateMin="2021-05-07" collectorName="Y. M. Marusik" country="Armenia" elevation="1040" latitude="40.498333" location="Geghadir" longLatPrecision="19" longitude="44.59" municipality="Urtsadzor Vill." specimenCount="29" specimenCount-female="14" specimenCount-juvenile="4" specimenCount-male="11" stateProvince="Kotayk Province">
<emphasis id="B79173CAA23F46CB8BF32E8856C0FF86" italics="true" pageId="0" pageNumber="117">
<collectingRegion id="86C28901285A996CF1B9EB5030B1E229" country="Armenia" name="Kotayk'">Kotayk Province</collectingRegion>
</emphasis>
:
<specimenCount id="208C22E43FBEE27542301EF5E1874AE5" type="male">3♂</specimenCount>
<specimenCount id="83B56A6EEAEA60E9EE74F631ED286FC3" type="female">2♀</specimenCount>
(ZMMU), env. of
<location id="4B3E2B885B8151B268FE9C85AD8370F2" LSID="urn:lsid:plazi:treatment:F1F60027744556EF968FBDD4C67717A7:4B3E2B885B8151B268FE9C85AD8370F2" country="Armenia" latitude="40.498333" longLatPrecision="19" longitude="44.59" name="Geghadir" stateProvince="Kotayk Province">Geghadir</location>
,
<geoCoordinate id="71430E1EADA77B87BCE317FAB1913CF4" degrees="40" direction="north" minutes="09" orientation="latitude" precision="925" value="40.15">40°09'N</geoCoordinate>
,
<geoCoordinate id="03AD78DD02DC68EF22A8B1D1F948D69E" degrees="44" direction="east" minutes="38" orientation="longitude" precision="925" value="44.633335">44°38'E</geoCoordinate>
,
<collectingDate id="27919387C456A51EBD5BDB030777EBD5" value="2021-05-15">15.05.2021</collectingDate>
(
<collectorName id="4F4E2B26FA7048871FE1737A2F437461">Y.M. Marusik</collectorName>
);
<specimenCount id="E9D7CB66039CA2A00648BEC09E751F7D" type="male">2♂</specimenCount>
<specimenCount id="D6CE531D68059358A1E3F2AB9B829EF9" type="female">4♀</specimenCount>
(ZMMU), env. of Solak Vill.,
<geoCoordinate id="D2CA705715E89C5EFB13807429FAD6FA" degrees="40" direction="north" minutes="28" orientation="latitude" precision="15" seconds="24" value="40.473335">40°28'24&quot;N</geoCoordinate>
,
<geoCoordinate id="E97913FE0C984186B858F36B9E92F1FB" degrees="44" direction="east" minutes="42" orientation="longitude" precision="15" seconds="57" value="44.715836">44°42'57&quot;E</geoCoordinate>
,
<collectingDate id="4377DC8E460C04A5CC42D9E9F58F3B6A" value="2021-05-14">14.05.2021</collectingDate>
(
<collectorName id="577A32A13F44BC3080CACAEFF0FD7295">Y.M. Marusik</collectorName>
);
<specimenCount id="ABA89BB28066E1FF4D92C783BC87138B" type="male">1♂</specimenCount>
<specimenCount id="AB1FA7AA0366FF581D8413377ABA914C" type="female">1♀</specimenCount>
(ZMMU), env. of Aghveran Vill.,
<geoCoordinate id="556E84C6C3F2DDBFE3030DD88121C7D4" degrees="40" direction="north" minutes="29" orientation="latitude" precision="15" seconds="54" value="40.498333">40°29'54&quot;N</geoCoordinate>
,
<geoCoordinate id="F546E6AD81FCFA70C1D77659E259A5EE" degrees="44" direction="east" minutes="35" orientation="longitude" precision="15" seconds="24" value="44.59">44°35'24&quot;E</geoCoordinate>
, 7-
<collectingDate id="3DA632935D5F5AE8C94441DD0AEC882A" value="2021-05-08">8.05.2021</collectingDate>
(
<collectorName id="1732FE3AA97C482073609A5D8B157136">Y.M. Marusik</collectorName>
);
<specimenCount id="02B0646455DDB2F663A299C37AFD5C3D" type="male">5♂</specimenCount>
<specimenCount id="6489D66F00953D00FB33A5DE6FA916AD" type="female">7♀</specimenCount>
<specimenCount id="6783830CA21CA5561B48231FEBF8A5B6" type="generic">4 juv.</specimenCount>
(ZMMU), same locality,
<collectingDate id="7DB96BF0709D96D873CE408936BE71F5" value="2021-05-07">7.05.2021</collectingDate>
(
<collectorName id="FE70A0669D8E9357DF124246F1BC5BDD">Y.M. Marusik</collectorName>
)
</materialsCitation>
;
<materialsCitation id="F95314C51D4D5CE60DAEFE7D923606DF" collectingDate="2021-05-09" collectorName="Marusik, Y. M. Marusik" country="Armenia" latitude="39.833332" location="Shatin" longLatPrecision="1166" longitude="45.316666" municipality="Shatin" specimenCount="40" specimenCount-female="17" specimenCount-male="23" stateProvince="Vayots Dzor Province">
<emphasis id="44A2AFA4DC133AFC4DFE80C41B90FF36" italics="true" pageId="0" pageNumber="117">
<collectingRegion id="8A6EED0E35F4B1838000DFE9564B00AA" country="Armenia" name="Vayots' Dzor">Vayots Dzor Province</collectingRegion>
</emphasis>
:
<specimenCount id="73E30286D5A08AFC061E0C4FCAE52AC1" type="male">21♂</specimenCount>
<specimenCount id="7CA9C712C36D80ECB901E4CB7A7A8F5E" type="female">14♀</specimenCount>
(ZMMU),
<collectingMunicipality id="6E896AB49C501E4309BDF00C504A85CA">Shatin</collectingMunicipality>
,
<geoCoordinate id="0FB44CC834D609033F36BE5F0DD220D8" degrees="39" direction="north" minutes="50" orientation="latitude" precision="925" value="39.833332">39°50'N</geoCoordinate>
,
<geoCoordinate id="5DE558DF07D4ACCCFDB8E9468456D7AC" degrees="45" direction="east" minutes="19" orientation="longitude" precision="925" value="45.316666">45°19'E</geoCoordinate>
,
<collectingDate id="F426A3F8179E13E69A727BA3F10C1C93" value="2021-05-09">9.05.2021</collectingDate>
(Y.M.
<collectorName id="6AE1CCF653E8E4EDE5F677062F71A0F6">Marusik</collectorName>
);
<specimenCount id="268D0C6B728EC7B1F55E203ED3F0C6EA" type="male">2♂</specimenCount>
<specimenCount id="4706945D7E98FD7098FDD00F281017D0" type="female">3♀</specimenCount>
(ZMMU),
<location id="B0CDC72A79B5F12200C412552D233E25" LSID="urn:lsid:plazi:treatment:F1F60027744556EF968FBDD4C67717A7:B0CDC72A79B5F12200C412552D233E25" country="Armenia" latitude="39.833332" longLatPrecision="1166" longitude="45.316666" municipality="Shatin" name="Shatin" stateProvince="Vayots Dzor Province">Shatin</location>
,
<geoCoordinate id="9C72A7ECC28101854AFA4F8424BD7FB7" degrees="39" direction="north" minutes="50" orientation="latitude" precision="925" value="39.833332">39°50'N</geoCoordinate>
,
<geoCoordinate id="077C3D1938AEE73FC4E056C1D335AD5C" degrees="45" direction="east" minutes="19" orientation="longitude" precision="925" value="45.316666">45°19'E</geoCoordinate>
,
<collectingDate id="366E9525652F1C14D7989EC6C1EBBD6B" value="2021-05-09">9.05.2021</collectingDate>
(
<collectorName id="6140FF1FCA8A2CF5C636535386601E26">Y.M. Marusik</collectorName>
)
</materialsCitation>
;
<materialsCitation id="452BFF2D9679611B109E3A58B82F7654" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" country="Armenia" location="Yerevan" specimenCount="16" specimenCount-female="9" specimenCount-male="7" stateProvince="Yerevan">
<emphasis id="0AA66048135A81456ABC666CE79C9F77" italics="true" pageId="0" pageNumber="117">
<collectingRegion id="0B7137D1F13991C0D477C92F1C31992B" country="Armenia" name="Yerevan">Yerevan</collectingRegion>
</emphasis>
:
<specimenCount id="D6EA4CE687C1303B40A8AA45CEAC7460" type="male">7♂</specimenCount>
<specimenCount id="8D953CBB8E9F98D99DB181CCD73E2975" type="female">9♀</specimenCount>
(ZMMU)
</materialsCitation>
,
<materialsCitation id="83FF5D8892DF2A70C452C18FBD23A026" collectingDate="2021-05-17" collectorName="Y. M. Marusik" country="Armenia" latitude="40.211945" location="Yerevan" longLatPrecision="19" longitude="44.555832" specimenCount="1" stateProvince="Yerevan">
<collectingRegion id="510DDD75F4037186512605DB594B105A" country="Armenia" name="Yerevan">Yerevan</collectingRegion>
, botanical garden,
<geoCoordinate id="5FCFFD000E05D6E612BD1BAC5DE0AD78" degrees="40" direction="north" minutes="12" orientation="latitude" precision="15" seconds="43" value="40.211945">40°12'43&quot;N</geoCoordinate>
,
<geoCoordinate id="3817F34732BA44C7091716CF753928CB" degrees="44" direction="east" minutes="33" orientation="longitude" precision="15" seconds="21" value="44.555832">44°33'21&quot;E</geoCoordinate>
,
<collectingDate id="ADAEE4832A9FC58650E59CB6E31CD53C" value="2021-05-17">17.05.2021</collectingDate>
(
<collectorName id="470E4A5CA029EF92CBF73728397DF541">Y.M. Marusik</collectorName>
)
</materialsCitation>
.
<materialsCitation id="6452B2902CD9A5FDC42B0779E4A97291" collectingDate="1975-06-05" collectorName="A. Senglet" country="Iran" latitude="38.35" location="NE of Sofian" longLatPrecision="1175" longitude="45.833332" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="East Azerbaijan Province">
<collectingCountry id="F092435E296F8AAE3599CB9234B83B47" name="Iran">Iran</collectingCountry>
:
<emphasis id="47231436ED37481EB5D03ADC6D775E3D" italics="true" pageId="0" pageNumber="117">
<collectingRegion id="968305DF89836ADA1741CD20647265A9" country="Iran" name="Azarbayjan-e Sharqi">East Azerbaijan Province</collectingRegion>
</emphasis>
:
<specimenCount id="5A9E1B3670532657BC0C944A6FEEEF83" type="male">1♂</specimenCount>
<specimenCount id="DD78C914E2F2ED18B408A6023790B8B8" type="female">3♀</specimenCount>
(MHNG),
<location id="BFE7621818F5037C15DFE8506B667273" LSID="urn:lsid:plazi:treatment:F1F60027744556EF968FBDD4C67717A7:BFE7621818F5037C15DFE8506B667273" country="Iran" latitude="38.35" longLatPrecision="1175" longitude="45.833332" name="NE of Sofian" stateProvince="East Azerbaijan Province">NE of Sofian</location>
,
<geoCoordinate id="529C8CEDCA5320AAEC0EF9D84DA6391E" degrees="38" direction="north" minutes="21" orientation="latitude" precision="925" value="38.35">38°21'N</geoCoordinate>
,
<geoCoordinate id="C90329D2A0A3956C499C7EF7E7190E3D" degrees="45" direction="east" minutes="50" orientation="longitude" precision="925" value="45.833332">45°50'E</geoCoordinate>
,
<collectingDate id="124854AC17EF65DDE3ADC8A60F7470B9" value="1975-06-05">5.06.1975</collectingDate>
(
<collectorName id="3AAB9650C518FB0373C960B685CB7103">A. Senglet</collectorName>
)
</materialsCitation>
;
<materialsCitation id="D9409807CF46588812237F4D8B3BF6BA" collectingDate="1973-07-05" collectingDateMax="1975-06-11" collectingDateMin="1973-07-05" collectorName="Senglet, A. Senglet" country="Iran" latitude="37.183334" location="Lahidjan" longLatPrecision="1183" longitude="49.9" municipality="Asalem" specimenCount="8" specimenCount-female="6" specimenCount-male="2" stateProvince="Gilan Province">
<emphasis id="C83AFBB634D13853F7B86C60F732D5DF" italics="true" pageId="0" pageNumber="117">
<collectingRegion id="F988B1040B2A154E9AC7D6B1E090EAC0" country="Iran" name="Gilan">Gilan Province</collectingRegion>
</emphasis>
:
<specimenCount id="CBAA7783A5637E2EE7CCC067F65A6D40" type="male">1♂</specimenCount>
<specimenCount id="EF57AB5A11182636FA228C5D286C24E9" type="female">3♀</specimenCount>
(MHNG),
<collectingMunicipality id="050D5AC8C0466FFA7B50BE68A43B0323">Asalem</collectingMunicipality>
,
<geoCoordinate id="91006851EC6CC43C121BA85906BF3EEB" degrees="37" direction="north" minutes="45" orientation="latitude" precision="925" value="37.75">37°45'N</geoCoordinate>
,
<geoCoordinate id="B20A55FFE253D9D52ADEF9DCA60B6869" degrees="48" direction="east" minutes="57" orientation="longitude" precision="925" value="48.95">48°57'E</geoCoordinate>
,
<collectingDate id="2F0391C7D00B8FE11E12806017E65B98" value="1975-06-11">11.06.1975</collectingDate>
(A.
<collectorName id="CD1A353B937865EFE261F9899AA80111">Senglet</collectorName>
);
<specimenCount id="09587290DF5115032145AD585C6453A7" type="male">1♂</specimenCount>
<specimenCount id="6B1B123DBEC879D055A636C4A791CCBE" type="female">3♀</specimenCount>
(MHNG),
<location id="D29538368A21F446AECB8BD9EE6C5E5C" LSID="urn:lsid:plazi:treatment:F1F60027744556EF968FBDD4C67717A7:D29538368A21F446AECB8BD9EE6C5E5C" country="Iran" latitude="37.183334" longLatPrecision="1183" longitude="49.9" municipality="Asalem" name="Lahidjan" stateProvince="Gilan Province">Lahidjan</location>
,
<geoCoordinate id="297340DB8603865DB3CF8B85F9A5CD4A" degrees="37" direction="north" minutes="11" orientation="latitude" precision="925" value="37.183334">37°11'N</geoCoordinate>
,
<geoCoordinate id="0B763C8C3AB792C89E406F533A083A91" degrees="49" direction="east" minutes="54" orientation="longitude" precision="925" value="49.9">49°54'E</geoCoordinate>
,
<collectingDate id="9ED583AFDB9B26A54D369443DE058937" value="1973-07-05">5.07.1973</collectingDate>
(
<collectorName id="02E2968D9075F05EA1E13F8235525392">A. Senglet</collectorName>
)
</materialsCitation>
;
<materialsCitation id="F27A2FFBF0B51BAB6F1EB3BF18F5DA18" collectingDate="1974-08-07" collectorName="A. Senglet" country="Iran" elevation="1200" latitude="36.916668" location="Djavaherdeh" longLatPrecision="1185" longitude="50.55" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Mazandaran Province">
<emphasis id="CCF10E8E908A4AAB1E80159F3B7E4838" italics="true" pageId="0" pageNumber="117">
<collectingRegion id="161846AA8A8C6E072F9FB18C52379AFE" country="Iran" name="Mazandaran">Mazandaran Province</collectingRegion>
</emphasis>
:
<specimenCount id="8DFE2BB849FBB1C521EE5170588C7698" type="male">1♂</specimenCount>
<specimenCount id="46F4D7AA097AC10386AB6AECA0470777" type="female">3♀</specimenCount>
(MHNG), road to
<location id="3C80B5A83F574AB5E4CC42B69F5C97FF" LSID="urn:lsid:plazi:treatment:F1F60027744556EF968FBDD4C67717A7:3C80B5A83F574AB5E4CC42B69F5C97FF" country="Iran" latitude="36.916668" longLatPrecision="1185" longitude="50.55" name="Djavaherdeh" stateProvince="Mazandaran Province">Djavaherdeh</location>
,
<geoCoordinate id="E9168F6ECF9308E90085C97506D99860" degrees="36" direction="north" minutes="55" orientation="latitude" precision="925" value="36.916668">36°55'N</geoCoordinate>
,
<geoCoordinate id="E3F02397276B2585C6F12604954231C2" degrees="50" direction="east" minutes="33" orientation="longitude" precision="925" value="50.55">50°33'E</geoCoordinate>
,
<quantity id="6DF63634D36AFE0D69304DFF903B9AED" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
<elevation id="A82909E06289EC4FDF512BF699C5986A" metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
</quantity>
,
<collectingDate id="C9A2D392FD4149E04B4A71D13A01EC8E" value="1974-08-07">7.08.1974</collectingDate>
(
<collectorName id="7765B5F26C8E3CE1F9095387AD1D8BAC">A. Senglet</collectorName>
)
</materialsCitation>
;
<materialsCitation id="863E882A27D889F35E4935E3829FEB5A" collectingDate="1975-05-31" collectorName="A. Senglet" country="Iran" latitude="39.066666" location="Qara Kelisa" longLatPrecision="1171" longitude="44.633335" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="West Azerbaijan Province">
<emphasis id="4B1A5D000C4BAB8C78ECE5EE5B14E92D" italics="true" pageId="0" pageNumber="117">
<collectingRegion id="25C6F54F01B3357D002194F990850F3C" country="Iran" name="Azarbayjan-e Gharbi">West Azerbaijan Province</collectingRegion>
</emphasis>
:
<specimenCount id="768E7E2670E1708B796AD50A20CC29A1" type="male">1♂</specimenCount>
<specimenCount id="3EBF430A306A95B104338FF1304EF3AB" type="female">3♀</specimenCount>
(MHNG),
<location id="4C3F5EA1B47093683146663FBD972AA1" LSID="urn:lsid:plazi:treatment:F1F60027744556EF968FBDD4C67717A7:4C3F5EA1B47093683146663FBD972AA1" country="Iran" latitude="39.066666" longLatPrecision="1171" longitude="44.633335" name="Qara Kelisa" stateProvince="West Azerbaijan Province">Qara Kelisa</location>
,
<geoCoordinate id="2253C80B26EF161100490BC3D8D5ED3B" degrees="39" direction="north" minutes="04" orientation="latitude" precision="925" value="39.066666">39°04'N</geoCoordinate>
,
<geoCoordinate id="873286C84760560908DB878A7DCF663D" degrees="44" direction="east" minutes="38" orientation="longitude" precision="925" value="44.633335">44°38'E</geoCoordinate>
,
<collectingDate id="94D2C8F445BCF05188B1EA1395240145" value="1975-05-31">31.05.1975</collectingDate>
(
<collectorName id="937522CA23B012230E6863FA9331AE7C">A. Senglet</collectorName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F5D682D0C017A52967713AA44BFA7A5C" pageId="0" pageNumber="117" type="comments">
<paragraph id="BCDEE552A7033457097A81F436DE9CAF" pageId="0" pageNumber="117">Comments.</paragraph>
<paragraph id="FEEA45CDF8B0D88E5E3DFE48A14CC0E9" pageId="0" pageNumber="117">
This species is most similar to
<taxonomicName id="EE6BF0AADBF771C05BA58BC22D241260" lsidName="H. secunda" pageId="0" pageNumber="117" rank="species" species="secunda">
<emphasis id="1DE2E4D208B74D9CA48EF39331AA8D63" italics="true" pageId="0" pageNumber="117">H. secunda</emphasis>
</taxonomicName>
Dunin, 1989, a species restricted to northern Armenia, but can be differentiated from it by the base of embolus (
<emphasis id="B309EAAAA04793F947EE11A98F96264D" italics="true" pageId="0" pageNumber="117">Eb</emphasis>
) (slightly and roundly bent, vs sharply bent to about 70°). It is possible that this small difference is only an intraspecific variation; in order to verify this, it is necessary to collect additional material from northern Armenia or examine the type specimens of
<taxonomicName id="1EFA95074C690B613D458A834ABD44A1" lsidName="H. secunda" pageId="0" pageNumber="117" rank="species" species="secunda">
<emphasis id="E9F19957F96BF7832655BAA5CC7BAD27" italics="true" pageId="0" pageNumber="117">H. secunda</emphasis>
</taxonomicName>
. Furthermore, it is likely that the female illustrated by
<bibRefCitation id="6BA6500E7E84E4E5BA250BFEA899E431" author="Dunin, PM" journalOrPublisher="Zoologicheskiĭ Zhurnal" pageId="0" pageNumber="117" pagination="142 - 145" refId="B8" refString="Dunin, PM, 1989. New spider species of the Harpactea genus from Armenia (Aranei, Dysderidae). Zoologicheskiĭ Zhurnal 68 (7): 142 - 145" title="New spider species of the Harpactea genus from Armenia (Aranei, Dysderidae)." volume="68" year="1989">Dunin (1989</bibRefCitation>
,
<bibRefCitation id="5F11AF717996DE93F044B1B4B710E3BE" author="Dunin, PM" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="117" pagination="35 - 76" refId="B11" refString="Dunin, PM, 1992. The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta 1 (3): 35 - 76" title="The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae)." volume="1" year="1992">1992</bibRefCitation>
) belongs to another species. All females collected in Armenia and Iran examined here have a longer receptacle, longer than anterior diverticulum (
<emphasis id="A308FFF2974ED9C1E7EB4A74B4612838" italics="true" pageId="0" pageNumber="117">Ad</emphasis>
)
<emphasis id="5B49C6097CE9590CDF910DF2B4CE0CF7" italics="true" pageId="0" pageNumber="117">sensu</emphasis>
<bibRefCitation id="CC64B89EF24BAD98D661AD373A06679B" author="Dunin, PM" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="117" pagination="35 - 76" refId="B11" refString="Dunin, PM, 1992. The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae). Arthropoda Selecta 1 (3): 35 - 76" title="The spider family Dysderidae of the Caucasian fauna (Arachnida Aranei Haplogynae)." volume="1" year="1992">Dunin (1992)</bibRefCitation>
, and lack bulbous thickening of the receptacle (
<emphasis id="FF737179C318A012A47BDB73490C9E8B" italics="true" pageId="0" pageNumber="117">Re</emphasis>
, Fig.
<figureCitation id="D430B112210E03D611776CAAF85ABA83" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Copulatory organs of Harpactea armenica A-C bulb, prolateral, retrolateral and ventral D endogyne, dorsal. Abbreviations: Ad - anterior diverticulum, Eb - base of embolus, Re - receptacle. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1172.107112.figure9" httpUri="https://binary.pensoft.net/fig/883884" pageId="0" pageNumber="117">9D</figureCitation>
).
</paragraph>
<caption id="953836F629F281713137CB2088FA4124" doi="10.3897/zookeys.1172.107112.figure8" httpUri="https://binary.pensoft.net/fig/883883" pageId="0" pageNumber="117" start="Figure 8" startId="F8">
<paragraph id="9A6541D0D761DBA139CD8570D687EEAF" pageId="0" pageNumber="117">
<emphasis id="FBCABBF4EE8C156DD5304044A974B3A6" bold="true" pageId="0" pageNumber="117">Figure 8.</emphasis>
Habitus of
<taxonomicName id="8628188F7B05C592A2B2408DF1E019C9" authorityName="Dunin" authorityYear="1989" class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea armenica" order="Aranei" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="armenica">
<emphasis id="2416E8C5499D520430197D1F38778B0F" italics="true" pageId="0" pageNumber="117">Harpactea armenica</emphasis>
</taxonomicName>
, male (
<emphasis id="2EE79294327EC6EADB6C4B7E86CB3C5E" bold="true" pageId="0" pageNumber="117">A</emphasis>
) and female (
<emphasis id="D318BF02DE1AA561E2CDA8DE7746DD9A" bold="true" pageId="0" pageNumber="117">B</emphasis>
), dorsal.
</paragraph>
</caption>
<caption id="60BE2502BE08B3885C1941116DB94CAE" doi="10.3897/zookeys.1172.107112.figure9" httpUri="https://binary.pensoft.net/fig/883884" pageId="0" pageNumber="117" start="Figure 9" startId="F9">
<paragraph id="66FB39E65A46AED94E2BFC820C9B8EBA" pageId="0" pageNumber="117">
<emphasis id="EF663F791B242FDFB56073F9E8720D78" bold="true" pageId="0" pageNumber="117">Figure 9.</emphasis>
Copulatory organs of
<taxonomicName id="48736137308220AFD114C0EBDA566F77" authorityName="Dunin" authorityYear="1989" class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea armenica" order="Aranei" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="armenica">
<emphasis id="E25D928029C61E885D76EDF6ABCA03C6" italics="true" pageId="0" pageNumber="117">Harpactea armenica</emphasis>
</taxonomicName>
<emphasis id="DF1E3451C7F7921A69C483D1EBAF2100" bold="true" pageId="0" pageNumber="117">A-C</emphasis>
bulb, prolateral, retrolateral and ventral
<emphasis id="CB4999E96B0AEAF86BA4D849FE4824B8" bold="true" pageId="0" pageNumber="117">D</emphasis>
endogyne, dorsal. Abbreviations:
<emphasis id="A7423F5252F2E695E29F83FCEC128638" italics="true" pageId="0" pageNumber="117">Ad</emphasis>
- anterior diverticulum,
<emphasis id="22F5D513EC1FD82E3BDD40D67A2A11D6" italics="true" pageId="0" pageNumber="117">Eb</emphasis>
- base of embolus,
<emphasis id="90D9F5FF2E4FC066B08FC24BF9F56ECE" italics="true" pageId="0" pageNumber="117">Re</emphasis>
- receptacle. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="D7944530C23A4C732ED423E83D01007B" pageId="0" pageNumber="117" type="distribution">
<paragraph id="6F94FC442703027761D0FA8622146886" pageId="0" pageNumber="117">Distribution.</paragraph>
<paragraph id="625C739831F3563B12691CFA1250B556" pageId="0" pageNumber="117">
This species was previously known from only two localities in Armenia: Yerevan and the southernmost part of Kotayk Province. The new records slightly extend the known range of the species to the north, approximately 5° to the east, and around 4° to the south. Additionally, this species is newly recorded from Iran (Fig.
<figureCitation id="62BDD6A86610E3980F622590DCDDAE49" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution records of Dysdera haykana sp. nov. (circles), D. nakhchivanica (squares), and Harpactea armenica (triangles). Type localities are marked with red." figureDoi="10.3897/zookeys.1172.107112.figure10" httpUri="https://binary.pensoft.net/fig/883885" pageId="0" pageNumber="117">10</figureCitation>
).
</paragraph>
<caption id="D954E7276A1E18F290616138CA0D6CB7" doi="10.3897/zookeys.1172.107112.figure10" httpUri="https://binary.pensoft.net/fig/883885" pageId="0" pageNumber="117" start="Figure 10" startId="F10">
<paragraph id="1E3DA6D3EC8A964A28DCE1A81E839D6B" pageId="0" pageNumber="117">
<emphasis id="4151CBF588DC7F801C101C247584EB20" bold="true" pageId="0" pageNumber="117">Figure 10.</emphasis>
Distribution records of
<taxonomicName id="4695ABBA56C051F9926A09AEBF0640E1" authorityName="Kosyan &amp; Zamani &amp; Marusik" authorityYear="2023" class="Arachnida" family="Dysderidae" genus="Dysdera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Dysdera haykana" order="Araneae" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="haykana">
<emphasis id="20069150D101C35D807CAF8FC77B93E0" italics="true" pageId="0" pageNumber="117">Dysdera haykana</emphasis>
</taxonomicName>
sp. nov. (circles),
<taxonomicName id="F0F461823915002E20E06D1BCCC665E5" lsidName="D. nakhchivanica" pageId="0" pageNumber="117" rank="species" species="nakhchivanica">
<emphasis id="E5543C33AB5B418287C5EA69903A3C1B" italics="true" pageId="0" pageNumber="117">D. nakhchivanica</emphasis>
</taxonomicName>
(squares), and
<taxonomicName id="0E4C9F0E895FF0A9FD8815B932D0F135" authorityName="Dunin" authorityYear="1989" class="Arachnida" family="Dysderidae" genus="Harpactea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Harpactea armenica" order="Aranei" pageId="0" pageNumber="117" phylum="Arthropoda" rank="species" species="armenica">
<emphasis id="C708F917CAB5DF98A4CB0BE93467295B" italics="true" pageId="0" pageNumber="117">Harpactea armenica</emphasis>
</taxonomicName>
(triangles). Type localities are marked with red.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>