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<document id="9981527E0349F6F26AEF7E60C6613757" ID="10.11646/zootaxa.4254.4.3SLASH10855" ID-CLB-Dataset="33201" ID-DOI="10.11646/zootaxa.4254.4.3" ID-GBIF-Dataset="d25360d8-7a6b-4441-8ce3-4fd2b89121f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="556045" ID-ZooBank="29A83502-70C0-4A8F-82D5-CD692FC3219E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1492587321329" checkinUser="plazi" docAuthor="Hara, Marcos R., Bragagnolo, Cibele &amp; Pinto-Da-Rocha, Ricardo" docDate="2017" docId="F179007AFF9EFFCE6CC5CC4DFC41FE47" docLanguage="en" docName="zootaxa.4254.4.3SLASH10855.pdf" docOrigin="Zootaxa 4254 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thaumatocranaus mirabilis Roewer 1932" docType="treatment" docVersion="7" lastPageNumber="463" masterDocId="0D407802FF9DFFC86C52CE0CFFD4FF97" masterDocTitle="Revision of Thaumatocranaus Roewer, 1932, with description of two new species (Opiliones: Laniatores: Gonyleptoidea)" masterLastPageNumber="472" masterPageNumber="457" pageNumber="460" updateTime="1698438960372" updateUser="ExternalLinkService">
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<mods:title id="6162B0F63D95FFC254A9CC982BFBD7E9">Revision of Thaumatocranaus Roewer, 1932, with description of two new species (Opiliones: Laniatores: Gonyleptoidea)</mods:title>
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<mods:namePart id="8C0957D952122EBFB47850447B6C1AF6">Hara, Marcos R.</mods:namePart>
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<treatment id="F179007AFF9EFFCE6CC5CC4DFC41FE47" ID-DOI="http://doi.org/10.5281/zenodo.6010182" ID-GBIF-Taxon="128535416" ID-Zenodo-Dep="6010182" LSID="urn:lsid:plazi:treatment:F179007AFF9EFFCE6CC5CC4DFC41FE47" httpUri="http://treatment.plazi.org/id/F179007AFF9EFFCE6CC5CC4DFC41FE47" lastPageId="6" lastPageNumber="463" pageId="3" pageNumber="460">
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<paragraph id="796FB16CFF9EFFCB6CC5CC4DFD43FDCC" blockId="3.[151,663,577,637]" box="[151,663,577,603]" pageId="3" pageNumber="460">
<heading id="22270600FF9EFFCB6CC5CC4DFD43FDCC" bold="true" box="[151,663,577,603]" fontSize="11" level="1" pageId="3" pageNumber="460" reason="1">
<taxonomicName id="BED0CAEFFF9EFFCB6CC5CC4DFD43FDCC" authority="Roewer, 1932" authorityName="Roewer" authorityYear="1932" box="[151,663,577,603]" class="Arachnida" family="Cranaidae" genus="Thaumatocranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="460" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="4BA46D7EFF9EFFCB6CC5CC4DFD43FDCC" bold="true" box="[151,663,577,603]" pageId="3" pageNumber="460">
<emphasis id="4BA46D7EFF9EFFCB6CC5CC4DFE3CFDCC" bold="true" box="[151,488,577,603]" italics="true" pageId="3" pageNumber="460">Thaumatocranaus mirabilis</emphasis>
<bibRefCitation id="1D41CC9DFF9EFFCB6DA2CC4DFD43FDCC" author="Roewer" box="[496,663,577,603]" pageId="3" pageNumber="460" refString="Roewer, C. F. (1932) Weitere Weberknechte VII (7. Erganzung der Weberknechte der Erde, 1923). Archiv fur Naturgeschichte, 1 (2), 275 - 350." type="journal article" year="1932">Roewer, 1932</bibRefCitation>
</emphasis>
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<paragraph id="796FB16CFF9EFFCB6CC5CC68FE88FDEB" blockId="3.[151,663,577,637]" box="[151,348,612,637]" pageId="3" pageNumber="460">
(
<figureCitation id="E1EBADE9FF9EFFCB6CCDCC68FF31FDEB" box="[159,229,612,637]" captionStart="FIGURE 1" captionStartId="4.[151,250,1857,1879]" captionTargetBox="[207,1331,223,1803]" captionTargetId="figure@4.[199,1365,193,1834]" captionTargetPageId="4" captionText="FIGURE 1. Thaumatocranaus mirabilis Roewer. Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting small tubercles); C, ditto, right lateral view; D, ocularium, anterior view; E, right ozopore area, dorsolateral view; F, right pedipalp, retrolateral view; G, ditto, apex of femur tarsus prolateral view. Abbreviations: A 1 3, scutal areas I III; Fh, frontal hump; Ft 1 3, free tergites I III; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum; Pr, prosoma (Ca, carapace = fused dorsal tergites of prosoma). A, C at same scale; F, G at same scale. Scale bars: A C: 1 mm; F, G, 0.5 mm; D, E: 0.25 mm." httpUri="https://zenodo.org/record/556046/files/figure.png" pageId="3" pageNumber="460">Figs 1</figureCitation>
,
<figureCitation id="E1EBADE9FF9EFFCB6CA1CC68FED7FDEB" box="[243,259,612,636]" captionStart="FIGURE 2" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[214,1391,186,1893]" captionTargetId="figure@5.[176,1411,167,1906]" captionTargetPageId="5" captionText="FIGURE 2. Thaumatocranaus mirabilis Roewer. Male (holotype): A, right leg I, apex of metatarsus tarsus I, prolateral view; B, right femur III, prolateral view; C, right trochanter IV, dorsal view; D, apex of right femur IV, dorsal view; E, anal operculum, posterior view; F, right trochanter tarsus IV, dorsal view; G, ditto, ventral view. Abbreviations: Pro, prolateral; Ret, retrolateral. A, B at same scale; F, G at same scale. Scale bars: A, B, F, G, 1 mm; C E, 0.5 mm." httpUri="https://zenodo.org/record/556047/files/figure.png" pageId="3" pageNumber="460">2</figureCitation>
,
<figureCitation id="E1EBADE9FF9EFFCB6D5DCC68FEC8FDEB" box="[271,284,612,636]" captionStart="FIGURE 7" captionStartId="12.[151,250,1877,1899]" captionTargetBox="[260,1351,245,1808]" captionTargetId="figure@12.[218,1385,200,1840]" captionTargetPageId="12" captionText="FIGURE 7. Distal part of Thaumatocranaus spp. penises: A, T. mirabilis, dorsal view, B, ditto, right lateral view, C, ditto, ventral view; D, T. magnificus sp. nov., dorsal view, E, ditto, left lateral view; F, T. splendidus sp. nov., dorsal view, G, ditto, left lateral view. Abbreviations: Gl, glans; MS A E, ventral plate macrosetae A E; St, glans stylus; Vpl, ventral plate. A C at same scale. D, E at same scale. F, G at same scale. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/556052/files/figure.png" pageId="3" pageNumber="460">7</figureCitation>
AC)
</paragraph>
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<paragraph id="796FB16CFF9EFFCB6CC5CCA0FAE9FD55" blockId="3.[151,1341,684,706]" box="[151,1341,684,706]" pageId="3" pageNumber="460">
<treatmentCitationGroup id="59C09642FF9EFFCB6CC5CCA0FAE9FD55" box="[151,1341,684,706]" pageId="3" pageNumber="460">
<taxonomicName id="BED0CAEFFF9EFFCB6CC5CCA0FDA3FD55" authority="Roewer, 1932: 308" authorityName="Roewer" authorityPageNumber="308" authorityYear="1932" box="[151,631,684,706]" class="Arachnida" family="Cranaidae" genus="Thaumatocranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="460" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="4BA46D7EFF9EFFCB6CC5CCA0FE65FD55" box="[151,433,684,706]" italics="true" pageId="3" pageNumber="460">Thaumatocranaus mirabilis</emphasis>
<treatmentCitation id="F871977DFF9EFFCB6DE5CCA1FDA3FD55" author="Roewer" box="[439,631,684,706]" page="308" pageId="3" pageNumber="460" year="1932">
<bibRefCitation id="1D41CC9DFF9EFFCB6DE5CCA1FD9CFD55" author="Roewer" box="[439,584,684,706]" pageId="3" pageNumber="460" refString="Roewer, C. F. (1932) Weitere Weberknechte VII (7. Erganzung der Weberknechte der Erde, 1923). Archiv fur Naturgeschichte, 1 (2), 275 - 350." type="journal article" year="1932">Roewer, 1932</bibRefCitation>
: 308
</treatmentCitation>
</taxonomicName>
(desc), fig. 24;
<treatmentCitation id="F871977DFF9EFFCB6F4ACCA0FBFFFD55" author="Soares" box="[792,1067,684,706]" page="619" pageId="3" pageNumber="460" year="1948">
<bibRefCitation id="1D41CC9DFF9EFFCB6F4ACCA0FC28FD55" author="Soares" box="[792,1020,684,706]" pageId="3" pageNumber="460" refString="Soares, B. A. M. &amp; Soares, H. E. M. (1948) Monografia dos generos de opilioes neotropicos I. Arquivos de Zoologia do Estado de Sao Paulo, 5 (9), 553 - 636." type="journal article" year="1948">Soares &amp; Soares 1948</bibRefCitation>
: 619
</treatmentCitation>
(cat);
<treatmentCitation id="F871977DFF9EFFCB683FCCA1FAD4FD55" author="Kury" box="[1133,1280,684,706]" page="99" pageId="3" pageNumber="460" year="2003">
<bibRefCitation id="1D41CC9DFF9EFFCB683FCCA1FB09FD55" author="Kury" box="[1133,1245,684,706]" pageId="3" pageNumber="460" refString="Kury, A. B. (2003) Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, Special Volume, Monografico n o 1, 5 - 337." type="journal article" year="2003">Kury 2003</bibRefCitation>
: 99
</treatmentCitation>
(cat).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="31CAE2E7FF9EFFCB6CC5CCE2FE79FCBC" pageId="3" pageNumber="460" type="materials_examined">
<paragraph id="796FB16CFF9EFFCB6CC5CCE2FE79FCBC" blockId="3.[151,1437,750,2035]" pageId="3" pageNumber="460">
<materialsCitation id="C9B8BB31FF9EFFCB6CC5CCE2FE79FCBC" ID-GBIF-Occurrence="1457229158" collectionCode="SMF" country="Ecuador" location="Material" pageId="3" pageNumber="460" specimenCode="SMF 1405" specimenCount="1" stateProvince="Chimborazo">
<emphasis id="4BA46D7EFF9EFFCB6CC5CCE2FE52FC90" bold="true" box="[151,390,750,775]" pageId="3" pageNumber="460">
<location id="7C0FE7B7FF9EFFCB6CC5CCE2FF2BFC90" LSID="urn:lsid:plazi:treatment:F179007AFF9EFFCE6CC5CC4DFC41FE47:7C0FE7B7FF9EFFCB6CC5CCE2FF2BFC90" box="[151,255,750,775]" country="Ecuador" name="Material" pageId="3" pageNumber="460" stateProvince="Chimborazo">Material</location>
examined.
</emphasis>
<collectingCountry id="01C7F1FCFF9EFFCB6DC0CCE3FDC9FC90" box="[402,541,751,775]" name="Ecuador" pageId="3" pageNumber="460">ECUADOR</collectingCountry>
.
<collectingRegion id="BB147F8EFF9EFFCB6E7FCCE2FD63FC90" box="[557,695,750,775]" country="Ecuador" name="Chimborazo" pageId="3" pageNumber="460">Chimborazo</collectingRegion>
: Alausi. Without further data of date or collector name, male holoype (
<specimenCode id="29761917FF9EFFCB6D52CD1FFEAFFCBC" box="[256,379,786,811]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="460">
<collectionCode id="1FC129A9FF9EFFCB6D52CD1FFEEEFCBC" box="[256,314,787,811]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="3" pageNumber="460">SMF</collectionCode>
1405
</specimenCode>
/16).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="31CAE2E7FF9EFFCB6C95CD3AFD03FC2C" pageId="3" pageNumber="460" type="diagnosis">
<paragraph id="796FB16CFF9EFFCB6C95CD3AFD03FC2C" blockId="3.[151,1437,750,2035]" pageId="3" pageNumber="460">
<emphasis id="4BA46D7EFF9EFFCB6C95CD3AFE95FCD8" bold="true" box="[199,321,822,847]" pageId="3" pageNumber="460">Diagnosis.</emphasis>
<taxonomicName id="BED0CAEFFF9EFFCB6D15CD3BFE1CFCD8" box="[327,456,822,847]" class="Arachnida" family="Cranaidae" genus="Thaumatocranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="460" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="4BA46D7EFF9EFFCB6D15CD3BFE1CFCD8" box="[327,456,822,847]" italics="true" pageId="3" pageNumber="460">T. mirabilis</emphasis>
</taxonomicName>
can be distinguished from the other two species of the genus by: the free tergite III with robust and bifid apophysis; coxa IV with bifid retroapical apophysis; trochanter IV with a retrodorsal, acuminated, enlarged apical tubercle; metatarsus I inflated on distal third; tarsi IIIIV with the three basalmost tarsomeres inflated; three pairs of MS B on penis ventral plate.
</paragraph>
</subSubSection>
<subSubSection id="31CAE2E7FF9EFFCE6C95CDCAFE4AFE1F" lastPageId="6" lastPageNumber="463" pageId="3" pageNumber="460" type="description">
<paragraph id="796FB16CFF9EFFCB6C95CDCAFEBCFBF8" blockId="3.[151,1437,750,2035]" pageId="3" pageNumber="460">
<emphasis id="4BA46D7EFF9EFFCB6C95CDCAFEA0FC48" bold="true" box="[199,372,966,991]" pageId="3" pageNumber="460">Redescription.</emphasis>
<emphasis id="4BA46D7EFF9EFFCB6D29CDCAFE61FC48" box="[379,437,966,991]" italics="true" pageId="3" pageNumber="460">Male</emphasis>
(
<typeStatus id="A66B0FCEFF9EFFCB6D96CDCAFDF9FC48" box="[452,557,966,991]" pageId="3" pageNumber="460" type="holotype">holotype</typeStatus>
): Dorsum (
<figureCitation id="E1EBADE9FF9EFFCB6EFBCDCAFD26FC48" box="[681,754,966,991]" captionStart="FIGURE 1" captionStartId="4.[151,250,1857,1879]" captionTargetBox="[207,1331,223,1803]" captionTargetId="figure@4.[199,1365,193,1834]" captionTargetPageId="4" captionText="FIGURE 1. Thaumatocranaus mirabilis Roewer. Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting small tubercles); C, ditto, right lateral view; D, ocularium, anterior view; E, right ozopore area, dorsolateral view; F, right pedipalp, retrolateral view; G, ditto, apex of femur tarsus prolateral view. Abbreviations: A 1 3, scutal areas I III; Fh, frontal hump; Ft 1 3, free tergites I III; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum; Pr, prosoma (Ca, carapace = fused dorsal tergites of prosoma). A, C at same scale; F, G at same scale. Scale bars: A C: 1 mm; F, G, 0.5 mm; D, E: 0.25 mm." httpUri="https://zenodo.org/record/556046/files/figure.png" pageId="3" pageNumber="460">Figs. 1</figureCitation>
A, CE, 2E): Measurements: DSL 3.55; DSW 3.25; LI 6.95; LII 12.30; LIII 8.40; LIV 11.90. Carapace and scutal areas IIII with tubercles sparsely distributed. Median longitudinal groove of scutal area I wide and conspicuous. Lateral margin of dorsal scutum with an external row of tubercles increasing in size posteriorly, becoming slightly large and blunt. Free tergite III with a robust, bifid median apophysis.
</paragraph>
<paragraph id="796FB16CFF9EFFCB6C95CA77FAA4FB04" blockId="3.[151,1437,750,2035]" box="[199,1392,1146,1171]" pageId="3" pageNumber="460">
Venter (
<figureCitation id="E1EBADE9FF9EFFCB6D4DCA76FEB4FB04" box="[287,352,1146,1171]" captionStart="FIGURE 1" captionStartId="4.[151,250,1857,1879]" captionTargetBox="[207,1331,223,1803]" captionTargetId="figure@4.[199,1365,193,1834]" captionTargetPageId="4" captionText="FIGURE 1. Thaumatocranaus mirabilis Roewer. Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting small tubercles); C, ditto, right lateral view; D, ocularium, anterior view; E, right ozopore area, dorsolateral view; F, right pedipalp, retrolateral view; G, ditto, apex of femur tarsus prolateral view. Abbreviations: A 1 3, scutal areas I III; Fh, frontal hump; Ft 1 3, free tergites I III; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum; Pr, prosoma (Ca, carapace = fused dorsal tergites of prosoma). A, C at same scale; F, G at same scale. Scale bars: A C: 1 mm; F, G, 0.5 mm; D, E: 0.25 mm." httpUri="https://zenodo.org/record/556046/files/figure.png" pageId="3" pageNumber="460">Fig. 1</figureCitation>
B): Coxae IIV tuberculate; coxa I with median rows of tubercles increasing in size apically.
</paragraph>
<paragraph id="796FB16CFF9EFFCB6C95CA92FF01FB4C" blockId="3.[151,1437,750,2035]" pageId="3" pageNumber="460">
Chelicerae (
<figureCitation id="E1EBADE9FF9EFFCB6D1CCA92FE5BFB20" box="[334,399,1182,1207]" captionStart="FIGURE 1" captionStartId="4.[151,250,1857,1879]" captionTargetBox="[207,1331,223,1803]" captionTargetId="figure@4.[199,1365,193,1834]" captionTargetPageId="4" captionText="FIGURE 1. Thaumatocranaus mirabilis Roewer. Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting small tubercles); C, ditto, right lateral view; D, ocularium, anterior view; E, right ozopore area, dorsolateral view; F, right pedipalp, retrolateral view; G, ditto, apex of femur tarsus prolateral view. Abbreviations: A 1 3, scutal areas I III; Fh, frontal hump; Ft 1 3, free tergites I III; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum; Pr, prosoma (Ca, carapace = fused dorsal tergites of prosoma). A, C at same scale; F, G at same scale. Scale bars: A C: 1 mm; F, G, 0.5 mm; D, E: 0.25 mm." httpUri="https://zenodo.org/record/556046/files/figure.png" pageId="3" pageNumber="460">Fig. 1</figureCitation>
A): Segment I with few dorsal tubercles; movable finger with 45 teeth; fixed finger with 5 teeth.
</paragraph>
<paragraph id="796FB16CFF9EFFCB6C95CAEAFCA2FAFC" blockId="3.[151,1437,750,2035]" pageId="3" pageNumber="460">
Pedipalps (
<figureCitation id="E1EBADE9FF9EFFCB6D14CAEAFE5CFB68" box="[326,392,1254,1279]" captionStart="FIGURE 1" captionStartId="4.[151,250,1857,1879]" captionTargetBox="[207,1331,223,1803]" captionTargetId="figure@4.[199,1365,193,1834]" captionTargetPageId="4" captionText="FIGURE 1. Thaumatocranaus mirabilis Roewer. Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting small tubercles); C, ditto, right lateral view; D, ocularium, anterior view; E, right ozopore area, dorsolateral view; F, right pedipalp, retrolateral view; G, ditto, apex of femur tarsus prolateral view. Abbreviations: A 1 3, scutal areas I III; Fh, frontal hump; Ft 1 3, free tergites I III; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum; Pr, prosoma (Ca, carapace = fused dorsal tergites of prosoma). A, C at same scale; F, G at same scale. Scale bars: A C: 1 mm; F, G, 0.5 mm; D, E: 0.25 mm." httpUri="https://zenodo.org/record/556046/files/figure.png" pageId="3" pageNumber="460">Fig. 1</figureCitation>
F, G): Trochanter dorsal face smooth, unarmed; ventral face with 12, enlarged setiferous tubercles (prolateral largest). Femur with few, sparsely distributed dorsal tubercles, 3 ventral tubercles. Patella tarsus with few dorsolateral scattered setiferous tubercles. Tibial setation: prolateral I, retrolateral?i (broken). Tarsal setation: prolateral and retrolateral IIi placed on distal half.
</paragraph>
<paragraph id="796FB16CFF9EFFCB6C95CB7BFEF4F864" blockId="3.[151,1437,750,2035]" pageId="3" pageNumber="460">
Legs (
<figureCitation id="E1EBADE9FF9EFFCB6D5FCB7AFE9BFA18" box="[269,335,1398,1423]" captionStart="FIGURE 2" captionStartId="5.[151,250,1921,1943]" captionTargetBox="[214,1391,186,1893]" captionTargetId="figure@5.[176,1411,167,1906]" captionTargetPageId="5" captionText="FIGURE 2. Thaumatocranaus mirabilis Roewer. Male (holotype): A, right leg I, apex of metatarsus tarsus I, prolateral view; B, right femur III, prolateral view; C, right trochanter IV, dorsal view; D, apex of right femur IV, dorsal view; E, anal operculum, posterior view; F, right trochanter tarsus IV, dorsal view; G, ditto, ventral view. Abbreviations: Pro, prolateral; Ret, retrolateral. A, B at same scale; F, G at same scale. Scale bars: A, B, F, G, 1 mm; C E, 0.5 mm." httpUri="https://zenodo.org/record/556047/files/figure.png" pageId="3" pageNumber="460">Fig. 2</figureCitation>
AD, F, G): Coxa IV with 1 prolateral and 1 retrolateral conical, apical apophyses, retrolateral one bifid and shortest. Trochanters IIV with 1 basoventral median enlarged tubercle; trochanters IIII retrolateral face with 1 basal and 1 apical enlarged tubercles (apical ones acuminated on trochanters IIIII). Trochanter IV with 1 prodorsal, acuminated, enlarged subapical tubercle pointing dorsally; retrolateral face with 1 median and 1 apical enlarged tubercles (apical largest). Femora III ventral face with 2 rows of slightly enlarged, pointed tubercles; femur I dorsal face unarmed; femur II dorsal face with 1 enlarged retroapical spine. Femur III with 1 pair of dorsoapical spines (retrolateral largest); ventral face with 1 prolateral row of tubercles increasing in size subapically, alternating high, acuminated tubercles with low ones on basal half, 1 retrolateral row of tubercles increasing in size apically, becoming acuminated; ventroapical face with 1 prolateral enlarged tubercle, 1 retrolateral spine. Femur IV roughly straight, with 1 pair of dorsoapical spines (retrolateral largest); ventral face with 2 rows of acuminated tubercles increasing in size apically, retrolateral one with enlarged tubercles; 1 pair of ventroapical tubercles (prolateral largest). Patellae III unarmed; patella III with 1 retrodorsal apical enlarged tubercle. Patella IV with 1 retrodorsal, acuminated, enlarged apical tubercle and few retroventral acuminated tubercles. Tibiae IIII unarmed. Tibia IV dorso- and ventroapical face unarmed, 1 retrolateral row of slightly enlarged tubercles (subapical ones largest); ventral face with 2 rows of tubercles, retrolateral one with 45 acuminated, large tubercles. Metatarsus I distal ¼ inflated; metatarsus IV basal part dorso-ventrally curved, with subapical constriction, apical part inflated. Tarsi
<date id="0D6E97ACFF9EFFCB6EE4C9BBFCC6F858" box="[694,786,1975,1999]" pageId="3" pageNumber="460">IIIIV 3</date>
basal most tarsomeres inflated. Tarsal segmentation: 5(3), 89(3), 6, 6.
</paragraph>
<caption id="2DAFE1E4FF99FFCC6CC5C94DFC14F866" httpUri="https://zenodo.org/record/556046/files/figure.png" pageId="4" pageNumber="461" targetBox="[207,1331,223,1803]" targetPageId="4">
<paragraph id="796FB16CFF99FFCC6CC5C94DFC14F866" blockId="4.[151,1436,1857,2033]" pageId="4" pageNumber="461">
<emphasis id="4BA46D7EFF99FFCC6CC5C94DFEC0F8C1" bold="true" box="[151,276,1857,1879]" pageId="4" pageNumber="461">FIGURE 1.</emphasis>
<taxonomicName id="BED0CAEFFF99FFCC6D4EC94DFD40F8C0" authority="Roewer." authorityName="Roewer." box="[284,660,1857,1879]" class="Arachnida" family="Cranaidae" genus="Thaumatocranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="461" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="4BA46D7EFF99FFCC6D4EC94DFDECF8C0" box="[284,568,1857,1879]" italics="true" pageId="4" pageNumber="461">Thaumatocranaus mirabilis</emphasis>
Roewer.
</taxonomicName>
Male (holotype): A, habitus, dorsal view; B, ditto, ventral view (schematic illustration, omitting small tubercles); C, ditto, right lateral view; D, ocularium, anterior view; E, right ozopore area, dorsolateral view; F, right pedipalp, retrolateral view; G, ditto, apex of femurtarsus prolateral view. Abbreviations: A13, scutal areas IIII; Fh, frontal hump; Ft13, free tergites IIII; Oc, ocularium; Op, opisthosoma; Pac, prolateral apical apophysis of coxa IV; Pm, posterior margin of dorsal scutum; Pr, prosoma (Ca, carapace = fused dorsal tergites of prosoma). A, C at same scale; F, G at same scale. Scale bars: AC: 1 mm; F, G, 0.5 mm; D, E: 0.25 mm.
</paragraph>
</caption>
<caption id="2DAFE1E4FF98FFCD6CC5C98DFB8EF864" httpUri="https://zenodo.org/record/556047/files/figure.png" pageId="5" pageNumber="462" targetBox="[214,1391,186,1893]" targetPageId="5">
<paragraph id="796FB16CFF98FFCD6CC5C98DFB8EF864" blockId="5.[151,1436,1921,2035]" pageId="5" pageNumber="462">
<emphasis id="4BA46D7EFF98FFCD6CC5C98DFEC7F801" bold="true" box="[151,275,1921,1943]" pageId="5" pageNumber="462">FIGURE 2.</emphasis>
<taxonomicName id="BED0CAEFFF98FFCD6D48C98DFD44F800" authority="Roewer." authorityName="Roewer." box="[282,656,1921,1943]" class="Arachnida" family="Cranaidae" genus="Thaumatocranaus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="462" phylum="Arthropoda" rank="species" species="mirabilis">
<emphasis id="4BA46D7EFF98FFCD6D48C98DFDE1F800" box="[282,565,1921,1943]" italics="true" pageId="5" pageNumber="462">Thaumatocranaus mirabilis</emphasis>
Roewer.
</taxonomicName>
Male (holotype): A, right leg I, apex of metatarsustarsus I, prolateral view; B, right femur III, prolateral view; C, right trochanter IV, dorsal view; D, apex of right femur IV, dorsal view; E, anal operculum, posterior view; F, right trochantertarsus IV, dorsal view; G, ditto, ventral view. Abbreviations: Pro, prolateral; Ret, retrolateral. A, B at same scale; F, G at same scale. Scale bars: A, B, F, G, 1 mm; CE, 0.5 mm.
</paragraph>
</caption>
<paragraph id="796FB16CFF9BFFCE6C95CE9BFE85FF60" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="463">
Penis (
<figureCitation id="E1EBADE9FF9BFFCE6D48CE9BFEB6FF27" box="[282,354,151,176]" captionStart="FIGURE 7" captionStartId="12.[151,250,1877,1899]" captionTargetBox="[260,1351,245,1808]" captionTargetId="figure@12.[218,1385,200,1840]" captionTargetPageId="12" captionText="FIGURE 7. Distal part of Thaumatocranaus spp. penises: A, T. mirabilis, dorsal view, B, ditto, right lateral view, C, ditto, ventral view; D, T. magnificus sp. nov., dorsal view, E, ditto, left lateral view; F, T. splendidus sp. nov., dorsal view, G, ditto, left lateral view. Abbreviations: Gl, glans; MS A E, ventral plate macrosetae A E; St, glans stylus; Vpl, ventral plate. A C at same scale. D, E at same scale. F, G at same scale. Scale bars: 0.1 mm." httpUri="https://zenodo.org/record/556052/files/figure.png" pageId="6" pageNumber="463">Fig. 7</figureCitation>
AC): Glans stylus sub cylindrical, without trichomes. Ventral plate with 3 pairs of long, subapically curved MS C, 1 pair of short MS D, 3 pairs of short MS E (basalmost pair largest), 2 pairs of MS A and 3 pairs of MS B.
</paragraph>
<paragraph id="796FB16CFF9BFFCE6C95CF08FD3EFEF2" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="463">Coloration: Large median pale beige spot (from distal ½ of scutal area I to III) and many narrow and/or small pale beige spots of depigmented areas on dorsal scutum. Body, patellae IIV, trochanter IV and apophyses dark brown. Remaining parts of the legs and body, brown.</paragraph>
<paragraph id="796FB16CFF9BFFCE6C95CF63FE4AFE1F" blockId="6.[151,1437,151,464]" box="[199,414,367,392]" pageId="6" pageNumber="463">Female: Unknown.</paragraph>
</subSubSection>
<subSubSection id="31CAE2E7FF9BFFCE6C95CF98FC41FE47" pageId="6" pageNumber="463" type="materials_examined">
<paragraph id="796FB16CFF9BFFCE6C95CF98FD6BFE3A" blockId="6.[151,1437,151,464]" box="[199,703,404,429]" pageId="6" pageNumber="463">
<materialsCitation id="C9B8BB31FF9BFFCE6C95CF98FD6BFE3A" ID-GBIF-Occurrence="1457229161" box="[199,703,404,429]" country="Ecuador" location="Chimborazo" pageId="6" pageNumber="463" specimenCount="1" stateProvince="Chimborazo" typeStatus="holotype">
<emphasis id="4BA46D7EFF9BFFCE6C95CF98FEB1FE3A" bold="true" box="[199,357,404,429]" pageId="6" pageNumber="463">
<typeStatus id="A66B0FCEFF9BFFCE6C95CF98FED6FE3A" box="[199,258,404,429]" pageId="6" pageNumber="463">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="01C7F1FCFF9BFFCE6D3ECF98FE1FFE3A" box="[364,459,404,429]" name="Ecuador" pageId="6" pageNumber="463">Ecuador</collectingCountry>
,
<collectingRegion id="BB147F8EFF9BFFCE6D84CF98FDB7FE3A" box="[470,611,404,429]" country="Ecuador" name="Chimborazo" pageId="6" pageNumber="463">Chimborazo</collectingRegion>
, Alausi.
</materialsCitation>
</paragraph>
<paragraph id="796FB16CFF9BFFCE6C95CFBBFC41FE47" blockId="6.[151,1437,151,464]" box="[199,917,439,464]" pageId="6" pageNumber="463">
<emphasis id="4BA46D7EFF9BFFCE6C95CFBBFDD0FE47" bold="true" box="[199,516,439,464]" pageId="6" pageNumber="463">Geographical distribution.</emphasis>
Known only from the type locality.
</paragraph>
</subSubSection>
</treatment>
</document>