416 lines
69 KiB
XML
416 lines
69 KiB
XML
<document id="C52FFF1AFBA7129D57630749B107F984" ID-DOI="10.11646/zootaxa.5020.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="5224139" ID-ZooBank="CCF5FB8C-2552-4299-B0AF-8673230E7609" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629392338490" checkinUser="plazi" docAuthor="Mielke, Carlos G. C., Grehan, John R. & Koike, Ricardo M." docDate="2021" docId="F17187DFEE3DFFA3FF10ECB42115FE26" docLanguage="en" docName="zootaxa.5020.3.7.pdf" docOrigin="Zootaxa 5020 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Agripialus Mielke & Grehan & Koike 2021, gen. n." docType="treatment" docVersion="3" lastPageNumber="564" masterDocId="0D48FFA7EE3CFFA0FF87EF662528FFD3" masterDocTitle="Descriptions of two new genera and six new species of ghost-moths (Lepidoptera Hepialoidea: Hepialidae) from south-eastern and southern Brazil" masterLastPageNumber="580" masterPageNumber="561" pageNumber="562" updateTime="1699071641799" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="2D513047909DE9DB6E293800665A069D">Descriptions of two new genera and six new species of ghost-moths (Lepidoptera Hepialoidea: Hepialidae) from south-eastern and southern Brazil</mods:title>
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<mods:namePart id="E0635129D8A47D50C192BE3DFE655BA8">Mielke, Carlos G. C.</mods:namePart>
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<mods:nameIdentifier id="DA74CAC784CB67A0C8FAFF6F0B6DEABF" type="ORCID">0000-0003-0091-0840</mods:nameIdentifier>
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<mods:affiliation id="E42D461925AF11FC7264DB2BB0159854">Caixa postal 1206, 84.145 - 000 Carambeí, Paraná, Brazil. & Research Associate, McGuire Center for Lepidoptera & Biodiversity, Gainesville, FL 32611, USA. & cmielke 1 @ uol. com. br; https: // orcid. org / 0000 - 0003 - 0091 - 0840</mods:affiliation>
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<mods:namePart id="9A2707465159BF8B069450BA84F77A4D">Grehan, John R.</mods:namePart>
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<mods:nameIdentifier id="38014B7E80741410738A3A84AA08A7E4" type="ORCID">0000-0002-3119-1140</mods:nameIdentifier>
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<mods:affiliation id="71DB2045920EE176A88317C995720438">Research Associate, McGuire Center for Lepidoptera & Biodiversity, Gainesville, FL 32611, USA. & calabar. John @ gmail. com; https: // orcid. org / 0000 - 0002 - 3119 - 1140</mods:affiliation>
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<mods:nameIdentifier id="DCD7533C3CB9FAE6ED12EBD0223F0A0E" type="email">ohn@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0AD772AE7746C18384D166AA443939C1">Koike, Ricardo M.</mods:namePart>
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<mods:affiliation id="531795DFBCB19002D76835E37318633A">Estrada das Canjicas, 2500, 07424 - 000, Arujá, São Paulo, Brazil. & ricardokoike. jp @ gmail. com; https: // orcid. org / 0000 - 0002 - 6330 - 6416</mods:affiliation>
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<mods:nameIdentifier id="02AF9442FC4DC8E32C7A2E870D763EC8" type="email">ricardokoike.jp@gmail.com</mods:nameIdentifier>
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<mods:date id="89706AC643B22B369142F9127602831B">2021</mods:date>
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<mods:number id="A01BACF743710884BDF2F605B0C29261">2021-08-13</mods:number>
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<taxonomicName id="BED84D4AEE3DFFA1FF10ECB4243EFC3E" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[151,278,978,1005]" class="Insecta" family="Hepialidae" genus="Agripialus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="4BACEADBEE3DFFA1FF10ECB4243EFC3E" bold="true" box="[151,278,978,1005]" italics="true" pageId="1" pageNumber="562">Agripialus</emphasis>
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<paragraph id="796736C9EE3DFFA1FF10EC912683FBC2" blockId="1.[151,939,978,1041]" box="[151,939,1014,1041]" pageId="1" pageNumber="562">
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(
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<figureCitation id="E1E32A4CEE3DFFA1FF18EC91242CFBC2" box="[159,260,1015,1041]" captionStart-0="FIGURES 1−4" captionStart-1="FIGURES 5−7" captionStartId-0="2.[151,264,1423,1447]" captionStartId-1="4.[151,264,1819,1843]" captionTargetBox-0="[152,1436,181,1399]" captionTargetBox-1="[150,1438,597,1795]" captionTargetId-0="figure-21@2.[151,1436,181,1399]" captionTargetId-1="figure-184@4.[150,1438,597,1795]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 1−4. Agripialus gen. n. spp. Adults. 1, A. campos sp. n.: HT ♀: dorsal view (1a), ventral view (1b) (DZUP). 2–4, A. variabilis sp. n.: 2, HT ♀: dorsal view (2a), ventral view (2b) (DZUP); 3, PT ♀ (CGCM 38.764): dorsal view (DZUP); 4, PT ♀ (CGCM 38.744): dorsal view (DZUP)." captionText-1="FIGURES 5−7. Agripialus gen. n. spp. Adults. 5-6, A. itatiaia sp. n.: 5, HT ♂: dorsal view (5a), ventral view (5b) (CEIOC); 6, PT ♀ (CGCM 25.532): dorsal view (6a), ventral view (6b) (CGCM). 7, A. caparao sp. n.: HT ♂: dorsal view (7a), ventral view (7b) (DZUP)." figureDoi-0="http://doi.org/10.5281/zenodo.5224141" figureDoi-1="http://doi.org/10.5281/zenodo.5224143" httpUri-0="https://zenodo.org/record/5224141/files/figure.png" httpUri-1="https://zenodo.org/record/5224143/files/figure.png" pageId="1" pageNumber="562">Figs 1–7</figureCitation>
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,
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<figureCitation id="E1E32A4CEE3DFFA1FE97EC912471FBC2" box="[272,345,1015,1041]" captionStart="FIGURES 16−24" captionStartId="10.[151,264,1874,1898]" captionTargetBox="[242,1332,181,1847]" captionTargetId="figure-21@10.[227,1361,181,1850]" captionTargetPageId="10" captionText="FIGURES 16−24. Transverse scales and legs. Figs 16–19. Transverse scales (arrowed). 16, Mutipialus dilatus sp. n. PT ♂ (CGCM 23.689). 17, M. monticolus sp. n., HT ♀. 18, Parapielus luteicornis ♂ (CGCM 28.544, Argentina, Rio Grande; CGCM). 19, Hepialiscus htayaungi ♂ (CGCM 33.586, Myanmar; CGCM). Figs 20–24. Legs. 20, Agripialus campos sp. n., PT ♀ (CGCM 31.875). 21, A. variabilis sp. n., HT ♀. 22, A. itatiaia sp. n., HT ♂. 23, A. caparao sp. n., HT ♂. 24, Mutipialus dilatus sp. n., PT ♂ (CGCM 27.435)." figureDoi="http://doi.org/10.5281/zenodo.5224149" httpUri="https://zenodo.org/record/5224149/files/figure.png" pageId="1" pageNumber="562">20–23</figureCitation>
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,
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<figureCitation id="E1E32A4CEE3DFFA1FEE3EC9024FEFBC2" box="[356,470,1014,1041]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="1" pageNumber="562">25–26, 29</figureCitation>
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,
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<figureCitation id="E1E32A4CEE3DFFA1FE65EC912702FBC2" box="[482,554,1015,1041]" captionStart="FIGURES 31−35" captionStartId="12.[151,264,1743,1767]" captionTargetBox="[155,1434,184,1706]" captionTargetId="figure-21@12.[151,1435,181,1718]" captionTargetPageId="12" captionText="FIGURES 31−35. Female abdomen (upper side, tergites; lower side, sternites). 31, Agripialus campos sp. n., PT (CGCM 31.875). 32, A. variabilis sp. n., HT. 33, A. itatiaia sp. n., PT (CGCM 25.532). 34, Mutipialus dilatus sp. n., HT (CGCM 5.801). 35, M. monticolus sp. n., HT." figureDoi="http://doi.org/10.5281/zenodo.5224155" httpUri="https://zenodo.org/record/5224155/files/figure.png" pageId="1" pageNumber="562">31–33</figureCitation>
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,
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<figureCitation id="E1E32A4CEE3DFFA1FDB1EC9127D4FBC2" box="[566,764,1015,1041]" captionStart="FIGURES 36−46" captionStartId="13.[151,264,1743,1767]" captionTargetBox="[251,1340,182,1714]" captionTargetId="figure-20@13.[247,1340,181,1719]" captionTargetPageId="13" captionText="FIGURES 36−46. Abdomen. Figs 36–39. Female segments VI (lower)–VIII (upper). 36, Agripialus campos sp. n., PT (CGCM 31.875); arrows pointing to the sclerotised sternites VII and VIII. 37, A. variabilis sp. n., HT. 38, Mutipialus dilatus sp. n., HT (CGCM 5.801). 39, M. monticolus sp. n., HT; arrow pointing to the membranous sternite VIII. Figs 40–42. Male abdomen (upper side, tergites; lower side, sternites). 40, Agripialus caparao sp. n., HT. 41, M. dilatus sp. n., PT (CGCM 27.435). 42, M. monticolus sp. n., PT (CGCM 29.697). Figs 43–46. Male segments VII (lower)–VIII (upper). 43, Agripialus itatiaia sp. n., HT; arrows pointing to the two plates of the sternite VII. 44, A. caparao sp. n., HT; arrows pointing to the two plates of the sternite VII. 45, Mutipialus dilatus sp. n., PT (CGCM 27.435); arrow pointing to the sternite VIII. 46, M. monticolus sp. n., PT (CGCM 29.697); arrow pointing to the sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.5224157" httpUri="https://zenodo.org/record/5224157/files/figure.png" pageId="1" pageNumber="562">36–37, 40, 43–44</figureCitation>
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,
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<figureCitation id="E1E32A4CEE3DFFA1FC8FEC912679FBC2" box="[776,849,1015,1041]" captionStart="FIGURES 47−50" captionStartId="14.[151,264,1413,1437]" captionTargetBox="[159,1421,185,1378]" captionTargetId="figure-21@14.[151,1436,181,1389]" captionTargetPageId="14" captionText="FIGURES 47−50. Male genitalia. 47, Agripialus itatiaia sp. n., HT: ventral (47a), dorsal (internal; 47b); arrows pointing to the posterior sclerotisation of the tergal lobe and pseudotegumen and the notch in the posterior margin of the saccus. 48, A. caparao sp. n., HT: ventral (48a), dorsal (internal; 48b); arrow pointing to the notch in the posterior margin of the saccus. 49, Mutipialus dilatus sp. n., PT (CGCM 27.435): lateral (49a), ventral (49b), dorsal (internal; 49c); arrow pointing to the entrance of the pocket-like saccus. 50, M. monticolus sp. n., PT (CGCM 29.697): lateral (50a), ventral (50b), dorsal (internal; 50c); arrow pointing to the entrance of the pocket-like caecum." figureDoi="http://doi.org/10.5281/zenodo.5224159" httpUri="https://zenodo.org/record/5224159/files/figure.png" pageId="1" pageNumber="562">47–48</figureCitation>
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,
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<figureCitation id="E1E32A4CEE3DFFA1FCDBEC90268BFBC2" box="[860,931,1014,1041]" captionStart="FIGURES 51−57" captionStartId="15.[151,264,1627,1651]" captionTargetBox="[162,1427,289,1591]" captionTargetId="figure-49@15.[151,1437,272,1603]" captionTargetPageId="15" captionText="FIGURES 51−57. Female genitalia. 51, Agripialus campos sp. n., PT: posterior view and ductus and corpus bursae; arrow pointing to W-shaped lamella antevaginalis. 52, A. variabilis sp. n., HT: posterior view and ductus and corpus bursae. 53, A. itatiaia sp. n., PT (CGCM 25.532): posterior view and ductus and corpus bursae. 54, Mutipialus dilatus sp. n., HT (CGCM 5.801): posterior (54a), lateral (54b); arrow pointing to slight mesal projection of the lamella antevaginalis. 55, M. monticolus sp. n., HT: posterior (55a), lateral (55b); arrow pointing to slight mesal projection of the lamella antevaginalis. 56, Mutipialus dilatus sp. n., HT (CGCM 5.801): ductus and corpus bursae. 57, M. monticolus sp. n., HT: ductus and corpus bursae." figureDoi="http://doi.org/10.5281/zenodo.5224161" httpUri="https://zenodo.org/record/5224161/files/figure.png" pageId="1" pageNumber="562">51–53</figureCitation>
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)
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</paragraph>
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<typeStatus id="A663886BEE3DFFA1FF10EB5A25E1FB87" box="[151,201,1084,1108]" pageId="1" pageNumber="562">Type</typeStatus>
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species:
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<taxonomicName id="BED84D4AEE3DFFA1FEA1EB5D24C9FB80" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[294,481,1083,1107]" class="Insecta" family="Hepialidae" genus="Agripialus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="itatiaia" status="sp. nov.">
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<emphasis id="4BACEADBEE3DFFA1FEA1EB5D24C9FB80" box="[294,481,1083,1107]" italics="true" pageId="1" pageNumber="562">Agripialus itatiaia</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="509F57A0EE3DFFA1FE60EB5D2711FB80" box="[487,569,1083,1107]" pageId="1" pageNumber="562" rank="species">
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<emphasis id="4BACEADBEE3DFFA1FE60EB5D24D7FB80" bold="true" box="[487,511,1083,1107]" pageId="1" pageNumber="562">sp</emphasis>
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.
|
||
<emphasis id="4BACEADBEE3DFFA1FD8BEB5D271BFB80" bold="true" box="[524,563,1083,1107]" pageId="1" pageNumber="562">nov</emphasis>
|
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.
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||
</taxonomicNameLabel>
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, by present designation.
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||
</paragraph>
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<paragraph id="796736C9EE3DFFA1FF10EB3C2195FBA2" blockId="1.[151,1213,1083,1138]" box="[151,1213,1114,1138]" pageId="1" pageNumber="562">
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||
Included species:
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||
<taxonomicName id="BED84D4AEE3DFFA1FECAEB3C249EFBA1" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[333,438,1114,1138]" class="Insecta" family="Hepialidae" genus="Agripialus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="campos" status="sp. nov.">
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<emphasis id="4BACEADBEE3DFFA1FECAEB3C2474FBA2" box="[333,348,1114,1137]" italics="true" pageId="1" pageNumber="562">A</emphasis>
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.
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<emphasis id="4BACEADBEE3DFFA1FEEEEB3D249EFBA1" box="[361,438,1115,1138]" italics="true" pageId="1" pageNumber="562">campos</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="509F57A0EE3DFFA1FE3AEB3C2727FBA2" box="[445,527,1114,1138]" pageId="1" pageNumber="562" rank="species">
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<emphasis id="4BACEADBEE3DFFA1FE3AEB3C24FDFBA1" bold="true" box="[445,469,1114,1138]" pageId="1" pageNumber="562">sp</emphasis>
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.
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<emphasis id="4BACEADBEE3DFFA1FE66EB3C2720FBA1" bold="true" box="[481,520,1114,1138]" pageId="1" pageNumber="562">nov</emphasis>
|
||
.
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||
</taxonomicNameLabel>
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,
|
||
<taxonomicName id="BED84D4AEE3DFFA1FD9BEB3C27A4FBA1" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[540,652,1114,1138]" class="Insecta" family="Hepialidae" genus="Agripialus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="caparao" status="sp. nov.">
|
||
<emphasis id="4BACEADBEE3DFFA1FD9BEB3C2703FBA2" box="[540,555,1114,1137]" italics="true" pageId="1" pageNumber="562">A</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3DFFA1FDBFEB3D27A4FBA1" box="[568,652,1115,1138]" italics="true" pageId="1" pageNumber="562">caparao</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="509F57A0EE3DFFA1FD15EB3C27CDFBA2" box="[658,741,1114,1138]" pageId="1" pageNumber="562" rank="species">
|
||
<emphasis id="4BACEADBEE3DFFA1FD15EB3C2782FBA1" bold="true" box="[658,682,1114,1138]" pageId="1" pageNumber="562">sp</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3DFFA1FD30EB3C27F6FBA1" bold="true" box="[695,734,1114,1138]" pageId="1" pageNumber="562">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="BED84D4AEE3DFFA1FD75EB3C267FFBA2" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[754,855,1114,1137]" class="Insecta" family="Hepialidae" genus="Agripialus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="itatiaia" status="sp. nov.">
|
||
<emphasis id="4BACEADBEE3DFFA1FD75EB3C2629FBA2" box="[754,769,1114,1137]" italics="true" pageId="1" pageNumber="562">A</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3DFFA1FC89EB3C267FFBA2" box="[782,855,1114,1137]" italics="true" pageId="1" pageNumber="562">itatiaia</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="509F57A0EE3DFFA1FCDAEB3C2687FBA2" box="[861,943,1114,1138]" pageId="1" pageNumber="562" rank="species">
|
||
<emphasis id="4BACEADBEE3DFFA1FCDAEB3C265DFBA1" bold="true" box="[861,885,1114,1138]" pageId="1" pageNumber="562">sp</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3DFFA1FC05EB3C2681FBA1" bold="true" box="[898,937,1114,1138]" pageId="1" pageNumber="562">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, and
|
||
<taxonomicName id="BED84D4AEE3DFFA1FC60EB3C214CFBA1" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[999,1124,1114,1138]" class="Insecta" family="Hepialidae" genus="Agripialus" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
|
||
<emphasis id="4BACEADBEE3DFFA1FC60EB3C26DEFBA2" box="[999,1014,1114,1137]" italics="true" pageId="1" pageNumber="562">A</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3DFFA1FB84EB3C214CFBA1" box="[1027,1124,1114,1138]" italics="true" pageId="1" pageNumber="562">variabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="509F57A0EE3DFFA1FBECEB3C2195FBA2" box="[1131,1213,1114,1138]" pageId="1" pageNumber="562" rank="species">
|
||
<emphasis id="4BACEADBEE3DFFA1FBECEB3C21ABFBA1" bold="true" box="[1131,1155,1114,1138]" pageId="1" pageNumber="562">sp</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3DFFA1FB08EB3C219EFBA1" bold="true" box="[1167,1206,1114,1138]" pageId="1" pageNumber="562">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31C26542EE3DFFA1FF10EBC62498FA40" pageId="1" pageNumber="562" type="diagnosis">
|
||
<paragraph id="796736C9EE3DFFA1FF10EBC62498FA40" blockId="1.[151,1437,1184,2038]" pageId="1" pageNumber="562">
|
||
<emphasis id="4BACEADBEE3DFFA1FF10EBC62422FB69" bold="true" box="[151,266,1184,1210]" pageId="1" pageNumber="562">Diagnosis</emphasis>
|
||
. Distinguished from all other
|
||
<taxonomicName id="BED84D4AEE3DFFA1FDDBEBC627FDFB69" box="[604,725,1184,1210]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="562" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
|
||
by (i) sternite VII in males (when known) forming two distinct plates (
|
||
<figureCitation id="E1E32A4CEE3DFFA1FF6FEBA22444FB0D" box="[232,364,1220,1246]" captionStart="FIGURES 36−46" captionStartId="13.[151,264,1743,1767]" captionTargetBox="[251,1340,182,1714]" captionTargetId="figure-20@13.[247,1340,181,1719]" captionTargetPageId="13" captionText="FIGURES 36−46. Abdomen. Figs 36–39. Female segments VI (lower)–VIII (upper). 36, Agripialus campos sp. n., PT (CGCM 31.875); arrows pointing to the sclerotised sternites VII and VIII. 37, A. variabilis sp. n., HT. 38, Mutipialus dilatus sp. n., HT (CGCM 5.801). 39, M. monticolus sp. n., HT; arrow pointing to the membranous sternite VIII. Figs 40–42. Male abdomen (upper side, tergites; lower side, sternites). 40, Agripialus caparao sp. n., HT. 41, M. dilatus sp. n., PT (CGCM 27.435). 42, M. monticolus sp. n., PT (CGCM 29.697). Figs 43–46. Male segments VII (lower)–VIII (upper). 43, Agripialus itatiaia sp. n., HT; arrows pointing to the two plates of the sternite VII. 44, A. caparao sp. n., HT; arrows pointing to the two plates of the sternite VII. 45, Mutipialus dilatus sp. n., PT (CGCM 27.435); arrow pointing to the sternite VIII. 46, M. monticolus sp. n., PT (CGCM 29.697); arrow pointing to the sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.5224157" httpUri="https://zenodo.org/record/5224157/files/figure.png" pageId="1" pageNumber="562">Figs 43–44</figureCitation>
|
||
), (ii) pseudotegumen partially membranous (
|
||
<figureCitation id="E1E32A4CEE3DFFA1FCE0EBA226C0FB0D" box="[871,1000,1220,1246]" captionStart="FIGURES 47−50" captionStartId="14.[151,264,1413,1437]" captionTargetBox="[159,1421,185,1378]" captionTargetId="figure-21@14.[151,1436,181,1389]" captionTargetPageId="14" captionText="FIGURES 47−50. Male genitalia. 47, Agripialus itatiaia sp. n., HT: ventral (47a), dorsal (internal; 47b); arrows pointing to the posterior sclerotisation of the tergal lobe and pseudotegumen and the notch in the posterior margin of the saccus. 48, A. caparao sp. n., HT: ventral (48a), dorsal (internal; 48b); arrow pointing to the notch in the posterior margin of the saccus. 49, Mutipialus dilatus sp. n., PT (CGCM 27.435): lateral (49a), ventral (49b), dorsal (internal; 49c); arrow pointing to the entrance of the pocket-like saccus. 50, M. monticolus sp. n., PT (CGCM 29.697): lateral (50a), ventral (50b), dorsal (internal; 50c); arrow pointing to the entrance of the pocket-like caecum." figureDoi="http://doi.org/10.5281/zenodo.5224159" httpUri="https://zenodo.org/record/5224159/files/figure.png" pageId="1" pageNumber="562">Figs 47–48</figureCitation>
|
||
) and (iii) the lamella antevaginalis Wshaped and not connected to tergum IX (
|
||
<figureCitation id="E1E32A4CEE3DFFA1FDDFEB8E27F3FAD0" box="[600,731,1256,1283]" captionStart="FIGURES 51−57" captionStartId="15.[151,264,1627,1651]" captionTargetBox="[162,1427,289,1591]" captionTargetId="figure-49@15.[151,1437,272,1603]" captionTargetPageId="15" captionText="FIGURES 51−57. Female genitalia. 51, Agripialus campos sp. n., PT: posterior view and ductus and corpus bursae; arrow pointing to W-shaped lamella antevaginalis. 52, A. variabilis sp. n., HT: posterior view and ductus and corpus bursae. 53, A. itatiaia sp. n., PT (CGCM 25.532): posterior view and ductus and corpus bursae. 54, Mutipialus dilatus sp. n., HT (CGCM 5.801): posterior (54a), lateral (54b); arrow pointing to slight mesal projection of the lamella antevaginalis. 55, M. monticolus sp. n., HT: posterior (55a), lateral (55b); arrow pointing to slight mesal projection of the lamella antevaginalis. 56, Mutipialus dilatus sp. n., HT (CGCM 5.801): ductus and corpus bursae. 57, M. monticolus sp. n., HT: ductus and corpus bursae." figureDoi="http://doi.org/10.5281/zenodo.5224161" httpUri="https://zenodo.org/record/5224161/files/figure.png" pageId="1" pageNumber="562">Figs 51–53</figureCitation>
|
||
). Also by the following combination of characters: i) transverse tuft over the eyes emerging just below the antennal socket, ii) labial palpus trisegmented, iii) antenna lamellate, iv) ‘hepialine’ venation, v)
|
||
<specimenCount id="6FDEFD40EE3DFFA1FE23EA572491FA98" box="[420,441,1329,1355]" pageId="1" pageNumber="562" type="male">♂</specimenCount>
|
||
hindwing 1A and 2A complete (
|
||
<figureCitation id="E1E32A4CEE3DFFA1FCAEEA562655FA98" box="[809,893,1328,1355]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="1" pageNumber="562">Fig. 25</figureCitation>
|
||
), vi)
|
||
<specimenCount id="6FDEFD40EE3DFFA1FC3EEA5726E6FA98" box="[953,974,1329,1355]" pageId="1" pageNumber="562" type="female">♀</specimenCount>
|
||
hindwing 1A, 2A and 3A complete (
|
||
<figureCitation id="E1E32A4CEE3DFFA1FAE8EA56259EFABC" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="1" pageNumber="562">Fig. 26</figureCitation>
|
||
), vii)
|
||
<specimenCount id="6FDEFD40EE3DFFA1FF7EEA332426FABC" box="[249,270,1365,1391]" pageId="1" pageNumber="562" type="female">♀</specimenCount>
|
||
hindwing 1A and 2A closer at basal half (
|
||
<figureCitation id="E1E32A4CEE3DFFA1FD62EA322610FABC" box="[741,824,1364,1391]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="1" pageNumber="562">Fig. 26</figureCitation>
|
||
), viii) arolium present, and ix) tergite and sternite VIII sclerotised in both sexes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31C26542EE3DFFA2FF40EAFA26BFF8E0" lastPageId="2" lastPageNumber="563" pageId="1" pageNumber="562" type="description">
|
||
<paragraph id="796736C9EE3DFFA1FF40EAFA2647FA65" blockId="1.[151,1437,1184,2038]" box="[199,879,1436,1463]" pageId="1" pageNumber="562">
|
||
<emphasis id="4BACEADBEE3DFFA1FF40EAFA2479FA65" bold="true" box="[199,337,1436,1462]" pageId="1" pageNumber="562">Description</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3DFFA1FEE7EAFA24B5FA65" bold="true" box="[352,413,1436,1462]" pageId="1" pageNumber="562">Male</emphasis>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3DFFA1FE2CEAFA2738FA64" box="[427,528,1436,1463]" captionStart="FIGURES 5−7" captionStartId="4.[151,264,1819,1843]" captionTargetBox="[150,1438,597,1795]" captionTargetId="figure-184@4.[150,1438,597,1795]" captionTargetPageId="4" captionText="FIGURES 5−7. Agripialus gen. n. spp. Adults. 5-6, A. itatiaia sp. n.: 5, HT ♂: dorsal view (5a), ventral view (5b) (CEIOC); 6, PT ♀ (CGCM 25.532): dorsal view (6a), ventral view (6b) (CGCM). 7, A. caparao sp. n.: HT ♂: dorsal view (7a), ventral view (7b) (DZUP)." figureDoi="http://doi.org/10.5281/zenodo.5224143" httpUri="https://zenodo.org/record/5224143/files/figure.png" pageId="1" pageNumber="562">Figs 5, 7</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3DFFA1FD9BEAFA274DFA65" box="[540,613,1436,1462]" captionStart="FIGURES 16−24" captionStartId="10.[151,264,1874,1898]" captionTargetBox="[242,1332,181,1847]" captionTargetId="figure-21@10.[227,1361,181,1850]" captionTargetPageId="10" captionText="FIGURES 16−24. Transverse scales and legs. Figs 16–19. Transverse scales (arrowed). 16, Mutipialus dilatus sp. n. PT ♂ (CGCM 23.689). 17, M. monticolus sp. n., HT ♀. 18, Parapielus luteicornis ♂ (CGCM 28.544, Argentina, Rio Grande; CGCM). 19, Hepialiscus htayaungi ♂ (CGCM 33.586, Myanmar; CGCM). Figs 20–24. Legs. 20, Agripialus campos sp. n., PT ♀ (CGCM 31.875). 21, A. variabilis sp. n., HT ♀. 22, A. itatiaia sp. n., HT ♂. 23, A. caparao sp. n., HT ♂. 24, Mutipialus dilatus sp. n., PT ♂ (CGCM 27.435)." figureDoi="http://doi.org/10.5281/zenodo.5224149" httpUri="https://zenodo.org/record/5224149/files/figure.png" pageId="1" pageNumber="562">22–23</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3DFFA1FDF7EAFA27A6FA64" box="[624,654,1436,1463]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="1" pageNumber="562">25</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3DFFA1FD1DEAFA2624FA65" box="[666,780,1436,1462]" captionStart="FIGURES 36−46" captionStartId="13.[151,264,1743,1767]" captionTargetBox="[251,1340,182,1714]" captionTargetId="figure-20@13.[247,1340,181,1719]" captionTargetPageId="13" captionText="FIGURES 36−46. Abdomen. Figs 36–39. Female segments VI (lower)–VIII (upper). 36, Agripialus campos sp. n., PT (CGCM 31.875); arrows pointing to the sclerotised sternites VII and VIII. 37, A. variabilis sp. n., HT. 38, Mutipialus dilatus sp. n., HT (CGCM 5.801). 39, M. monticolus sp. n., HT; arrow pointing to the membranous sternite VIII. Figs 40–42. Male abdomen (upper side, tergites; lower side, sternites). 40, Agripialus caparao sp. n., HT. 41, M. dilatus sp. n., PT (CGCM 27.435). 42, M. monticolus sp. n., PT (CGCM 29.697). Figs 43–46. Male segments VII (lower)–VIII (upper). 43, Agripialus itatiaia sp. n., HT; arrows pointing to the two plates of the sternite VII. 44, A. caparao sp. n., HT; arrows pointing to the two plates of the sternite VII. 45, Mutipialus dilatus sp. n., PT (CGCM 27.435); arrow pointing to the sternite VIII. 46, M. monticolus sp. n., PT (CGCM 29.697); arrow pointing to the sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.5224157" httpUri="https://zenodo.org/record/5224157/files/figure.png" pageId="1" pageNumber="562">40, 43–44</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3DFFA1FC9FEAFA264AFA65" box="[792,866,1436,1462]" captionStart="FIGURES 47−50" captionStartId="14.[151,264,1413,1437]" captionTargetBox="[159,1421,185,1378]" captionTargetId="figure-21@14.[151,1436,181,1389]" captionTargetPageId="14" captionText="FIGURES 47−50. Male genitalia. 47, Agripialus itatiaia sp. n., HT: ventral (47a), dorsal (internal; 47b); arrows pointing to the posterior sclerotisation of the tergal lobe and pseudotegumen and the notch in the posterior margin of the saccus. 48, A. caparao sp. n., HT: ventral (48a), dorsal (internal; 48b); arrow pointing to the notch in the posterior margin of the saccus. 49, Mutipialus dilatus sp. n., PT (CGCM 27.435): lateral (49a), ventral (49b), dorsal (internal; 49c); arrow pointing to the entrance of the pocket-like saccus. 50, M. monticolus sp. n., PT (CGCM 29.697): lateral (50a), ventral (50b), dorsal (internal; 50c); arrow pointing to the entrance of the pocket-like caecum." figureDoi="http://doi.org/10.5281/zenodo.5224159" httpUri="https://zenodo.org/record/5224159/files/figure.png" pageId="1" pageNumber="562">47–48</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3DFFA1FF40EAA62133F995" blockId="1.[151,1437,1184,2038]" pageId="1" pageNumber="562">
|
||
<emphasis id="4BACEADBEE3DFFA1FF40EAA6242CFA09" box="[199,260,1472,1498]" italics="true" pageId="1" pageNumber="562">Head</emphasis>
|
||
. Clypeus glabrous, projected, and differentiated from frons. Frons with piliform and porrect scales, transverse ocular scale tuft from medial eye margin. Vertex scales as for frons. Eyes large, occupying 3/5 of the head in anterior view. Labial palps trisegmented. Antenna lamellate, each segment ventrally projected, scattered sensilla caetica and sensilla trichodea present; scape and pedicel with scales as for frons.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3DFFA1FF40E9362424F929" blockId="1.[151,1437,1184,2038]" pageId="1" pageNumber="562">
|
||
<emphasis id="4BACEADBEE3DFFA1FF40E9362408F9B9" box="[199,288,1616,1642]" italics="true" pageId="1" pageNumber="562">Thorax.</emphasis>
|
||
Legs (
|
||
<figureCitation id="E1E32A4CEE3DFFA1FEEDE93624C5F9B9" box="[362,493,1616,1642]" captionStart="FIGURES 16−24" captionStartId="10.[151,264,1874,1898]" captionTargetBox="[242,1332,181,1847]" captionTargetId="figure-21@10.[227,1361,181,1850]" captionTargetPageId="10" captionText="FIGURES 16−24. Transverse scales and legs. Figs 16–19. Transverse scales (arrowed). 16, Mutipialus dilatus sp. n. PT ♂ (CGCM 23.689). 17, M. monticolus sp. n., HT ♀. 18, Parapielus luteicornis ♂ (CGCM 28.544, Argentina, Rio Grande; CGCM). 19, Hepialiscus htayaungi ♂ (CGCM 33.586, Myanmar; CGCM). Figs 20–24. Legs. 20, Agripialus campos sp. n., PT ♀ (CGCM 31.875). 21, A. variabilis sp. n., HT ♀. 22, A. itatiaia sp. n., HT ♂. 23, A. caparao sp. n., HT ♂. 24, Mutipialus dilatus sp. n., PT ♂ (CGCM 27.435)." figureDoi="http://doi.org/10.5281/zenodo.5224149" httpUri="https://zenodo.org/record/5224149/files/figure.png" pageId="1" pageNumber="562">Figs 22–23</figureCitation>
|
||
): epiphysis and arolium present. Venation (
|
||
<figureCitation id="E1E32A4CEE3DFFA1FC4BE9362137F9B8" box="[972,1055,1616,1643]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="1" pageNumber="562">Fig. 25</figureCitation>
|
||
): fore- and hindwing without Sc1; hindwing CuP absent, 1A and 2A complete; hindwing Sc and Rs separated. Dorsal forewing ornamentation slightly marked by transverse blackish patch between cubital veins which sometimes extends disto-anteriorly, in parallel to outer margin. Ventral forewing, dorsal and ventral hindwing uniformly coloured, sometimes darker distally, unornamented.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3DFFA1FF40E86227DFF859" blockId="1.[151,1437,1184,2038]" pageId="1" pageNumber="562">
|
||
<emphasis id="4BACEADBEE3DFFA1FF40E8622419F8CD" box="[199,305,1796,1822]" italics="true" pageId="1" pageNumber="562">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3DFFA1FEC7E86224C5F8CD" box="[320,493,1796,1822]" captionStart="FIGURES 36−46" captionStartId="13.[151,264,1743,1767]" captionTargetBox="[251,1340,182,1714]" captionTargetId="figure-20@13.[247,1340,181,1719]" captionTargetPageId="13" captionText="FIGURES 36−46. Abdomen. Figs 36–39. Female segments VI (lower)–VIII (upper). 36, Agripialus campos sp. n., PT (CGCM 31.875); arrows pointing to the sclerotised sternites VII and VIII. 37, A. variabilis sp. n., HT. 38, Mutipialus dilatus sp. n., HT (CGCM 5.801). 39, M. monticolus sp. n., HT; arrow pointing to the membranous sternite VIII. Figs 40–42. Male abdomen (upper side, tergites; lower side, sternites). 40, Agripialus caparao sp. n., HT. 41, M. dilatus sp. n., PT (CGCM 27.435). 42, M. monticolus sp. n., PT (CGCM 29.697). Figs 43–46. Male segments VII (lower)–VIII (upper). 43, Agripialus itatiaia sp. n., HT; arrows pointing to the two plates of the sternite VII. 44, A. caparao sp. n., HT; arrows pointing to the two plates of the sternite VII. 45, Mutipialus dilatus sp. n., PT (CGCM 27.435); arrow pointing to the sternite VIII. 46, M. monticolus sp. n., PT (CGCM 29.697); arrow pointing to the sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.5224157" httpUri="https://zenodo.org/record/5224157/files/figure.png" pageId="1" pageNumber="562">Figs 40, 43–44</figureCitation>
|
||
). Tergosternal sclerite with strongly curved tergosternal bar curving disto-anteriorly, intermediate zone weakly sclerotized with robust posterior edge angled ventrally almost to tergosternal bar separated by narrow ‘notch’; lateral ridge anterior to tergosternal bar short, dorsal ridge not fused with anterior ridge of tergum I. Tergite and sternite VII and VIII sclerotized.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3DFFA1FF40E8F2243FF825" blockId="1.[151,1437,1184,2038]" pageId="1" pageNumber="562">
|
||
<emphasis id="4BACEADBEE3DFFA1FF40E8F22419F87D" box="[199,305,1940,1966]" italics="true" pageId="1" pageNumber="562">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3DFFA1FEC5E8F224E0F87D" box="[322,456,1940,1966]" captionStart="FIGURES 47−50" captionStartId="14.[151,264,1413,1437]" captionTargetBox="[159,1421,185,1378]" captionTargetId="figure-21@14.[151,1436,181,1389]" captionTargetPageId="14" captionText="FIGURES 47−50. Male genitalia. 47, Agripialus itatiaia sp. n., HT: ventral (47a), dorsal (internal; 47b); arrows pointing to the posterior sclerotisation of the tergal lobe and pseudotegumen and the notch in the posterior margin of the saccus. 48, A. caparao sp. n., HT: ventral (48a), dorsal (internal; 48b); arrow pointing to the notch in the posterior margin of the saccus. 49, Mutipialus dilatus sp. n., PT (CGCM 27.435): lateral (49a), ventral (49b), dorsal (internal; 49c); arrow pointing to the entrance of the pocket-like saccus. 50, M. monticolus sp. n., PT (CGCM 29.697): lateral (50a), ventral (50b), dorsal (internal; 50c); arrow pointing to the entrance of the pocket-like caecum." figureDoi="http://doi.org/10.5281/zenodo.5224159" httpUri="https://zenodo.org/record/5224159/files/figure.png" pageId="1" pageNumber="562">Figs 47–48</figureCitation>
|
||
). Tegumen fused to pseudotegumen. Tergal lobes and posterior portion of pseudotegumen membranous. Pseudotegumen fused anteriorly. Fultura superior membranous. Valva elbowed and heavily sclerotised.
|
||
</paragraph>
|
||
<caption id="2DA76641EE3EFFA2FF10EAE9270AFA3C" ID-DOI="http://doi.org/10.5281/zenodo.5224141" ID-Zenodo-Dep="5224141" httpUri="https://zenodo.org/record/5224141/files/figure.png" pageId="2" pageNumber="563" startId="2.[151,264,1423,1447]" targetBox="[152,1436,181,1399]" targetPageId="2">
|
||
<paragraph id="796736C9EE3EFFA2FF10EAE9270AFA3C" blockId="2.[151,1436,1423,1519]" pageId="2" pageNumber="563">
|
||
<emphasis id="4BACEADBEE3EFFA2FF10EAE9241EFA74" bold="true" box="[151,310,1423,1447]" pageId="2" pageNumber="563">FIGURES 1−4</emphasis>
|
||
.
|
||
<taxonomicName id="BED84D4AEE3EFFA2FEC5EAE92485FA74" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[322,429,1423,1447]" class="Insecta" family="Hepialidae" genus="Agripialus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="4BACEADBEE3EFFA2FEC5EAE92485FA74" box="[322,429,1423,1447]" italics="true" pageId="2" pageNumber="563">Agripialus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="509F57A0EE3EFFA2FE35EAE924D1FA75" box="[434,505,1423,1447]" pageId="2" pageNumber="563" rank="genus">
|
||
<emphasis id="4BACEADBEE3EFFA2FE35EAE924F0FA74" bold="true" box="[434,472,1423,1447]" pageId="2" pageNumber="563">gen</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3EFFA2FE62EAE924DBFA74" bold="true" box="[485,499,1423,1447]" pageId="2" pageNumber="563">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
spp. Adults.
|
||
<emphasis id="4BACEADBEE3EFFA2FDFAEAE927A2FA74" bold="true" box="[637,650,1423,1447]" pageId="2" pageNumber="563">1</emphasis>
|
||
,
|
||
<taxonomicName id="BED84D4AEE3EFFA2FD11EAE927D6FA74" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[662,766,1423,1447]" class="Insecta" family="Hepialidae" genus="Agripialus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="species" species="campos" status="sp. nov.">
|
||
<emphasis id="4BACEADBEE3EFFA2FD11EAE9278DFA75" box="[662,677,1423,1446]" italics="true" pageId="2" pageNumber="563">A</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3EFFA2FD36EAF627D6FA74" box="[689,766,1424,1447]" italics="true" pageId="2" pageNumber="563">campos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="509F57A0EE3EFFA2FC82EAE92616FA75" box="[773,830,1423,1447]" pageId="2" pageNumber="563" rank="species">
|
||
<emphasis id="4BACEADBEE3EFFA2FC82EAE92635FA74" bold="true" box="[773,797,1423,1447]" pageId="2" pageNumber="563">sp</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3EFFA2FCAEEAE9261FFA74" bold="true" box="[809,823,1423,1447]" pageId="2" pageNumber="563">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
: HT ♀: dorsal view (1a), ventral view (1b) (DZUP).
|
||
<emphasis id="4BACEADBEE3EFFA2FAD3EAE92052FA74" bold="true" box="[1364,1402,1423,1447]" pageId="2" pageNumber="563">2–4</emphasis>
|
||
,
|
||
<taxonomicName id="BED84D4AEE3EFFA2FA01EAE925D0FA18" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" class="Insecta" family="Hepialidae" genus="Agripialus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="species" species="variabilis" status="sp. nov.">
|
||
<emphasis id="4BACEADBEE3EFFA2FA01EAE920BDFA75" box="[1414,1429,1423,1446]" italics="true" pageId="2" pageNumber="563">A</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3EFFA2FF10EAD525D0FA18" box="[151,248,1459,1483]" italics="true" pageId="2" pageNumber="563">variabilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="509F57A0EE3EFFA2FF79EAD52410FA19" box="[254,312,1459,1483]" pageId="2" pageNumber="563" rank="species">
|
||
<emphasis id="4BACEADBEE3EFFA2FF79EAD5243EFA18" bold="true" box="[254,278,1459,1483]" pageId="2" pageNumber="563">sp</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3EFFA2FEA4EAD52419FA18" bold="true" box="[291,305,1459,1483]" pageId="2" pageNumber="563">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
:
|
||
<emphasis id="4BACEADBEE3EFFA2FEC3EAD52479FA18" bold="true" box="[324,337,1459,1483]" pageId="2" pageNumber="563">2</emphasis>
|
||
, HT ♀: dorsal view (2a), ventral view (2b) (DZUP);
|
||
<emphasis id="4BACEADBEE3EFFA2FCEEEAD5265EFA18" bold="true" box="[873,886,1459,1483]" pageId="2" pageNumber="563">3</emphasis>
|
||
, PT ♀ (CGCM 38.764): dorsal view (DZUP);
|
||
<emphasis id="4BACEADBEE3EFFA2FACAEAD52072FA18" bold="true" box="[1357,1370,1459,1483]" pageId="2" pageNumber="563">4</emphasis>
|
||
, PT ♀ (CGCM 38.744): dorsal view (DZUP).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="796736C9EE3EFFA2FF40E9272648F98F" blockId="2.[151,1437,1601,2024]" box="[199,864,1601,1628]" pageId="2" pageNumber="563">
|
||
<emphasis id="4BACEADBEE3EFFA2FF40E9272436F988" bold="true" box="[199,286,1601,1627]" pageId="2" pageNumber="563">Female</emphasis>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3EFFA2FEAAE92724BAF988" box="[301,402,1601,1627]" captionStart="FIGURES 1−4" captionStartId="2.[151,264,1423,1447]" captionTargetBox="[152,1436,181,1399]" captionTargetId="figure-21@2.[151,1436,181,1399]" captionTargetPageId="2" captionText="FIGURES 1−4. Agripialus gen. n. spp. Adults. 1, A. campos sp. n.: HT ♀: dorsal view (1a), ventral view (1b) (DZUP). 2–4, A. variabilis sp. n.: 2, HT ♀: dorsal view (2a), ventral view (2b) (DZUP); 3, PT ♀ (CGCM 38.764): dorsal view (DZUP); 4, PT ♀ (CGCM 38.744): dorsal view (DZUP)." figureDoi="http://doi.org/10.5281/zenodo.5224141" httpUri="https://zenodo.org/record/5224141/files/figure.png" pageId="2" pageNumber="563">Figs 1–4</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3EFFA2FE19E9272486F988" box="[414,430,1601,1627]" captionStart="FIGURES 5−7" captionStartId="4.[151,264,1819,1843]" captionTargetBox="[150,1438,597,1795]" captionTargetId="figure-184@4.[150,1438,597,1795]" captionTargetPageId="4" captionText="FIGURES 5−7. Agripialus gen. n. spp. Adults. 5-6, A. itatiaia sp. n.: 5, HT ♂: dorsal view (5a), ventral view (5b) (CEIOC); 6, PT ♀ (CGCM 25.532): dorsal view (6a), ventral view (6b) (CGCM). 7, A. caparao sp. n.: HT ♂: dorsal view (7a), ventral view (7b) (DZUP)." figureDoi="http://doi.org/10.5281/zenodo.5224143" httpUri="https://zenodo.org/record/5224143/files/figure.png" pageId="2" pageNumber="563">6</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3EFFA2FE3DE927272AF988" box="[442,514,1601,1627]" captionStart="FIGURES 16−24" captionStartId="10.[151,264,1874,1898]" captionTargetBox="[242,1332,181,1847]" captionTargetId="figure-21@10.[227,1361,181,1850]" captionTargetPageId="10" captionText="FIGURES 16−24. Transverse scales and legs. Figs 16–19. Transverse scales (arrowed). 16, Mutipialus dilatus sp. n. PT ♂ (CGCM 23.689). 17, M. monticolus sp. n., HT ♀. 18, Parapielus luteicornis ♂ (CGCM 28.544, Argentina, Rio Grande; CGCM). 19, Hepialiscus htayaungi ♂ (CGCM 33.586, Myanmar; CGCM). Figs 20–24. Legs. 20, Agripialus campos sp. n., PT ♀ (CGCM 31.875). 21, A. variabilis sp. n., HT ♀. 22, A. itatiaia sp. n., HT ♂. 23, A. caparao sp. n., HT ♂. 24, Mutipialus dilatus sp. n., PT ♂ (CGCM 27.435)." figureDoi="http://doi.org/10.5281/zenodo.5224149" httpUri="https://zenodo.org/record/5224149/files/figure.png" pageId="2" pageNumber="563">20–21</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3EFFA2FD89E927277EF988" box="[526,598,1601,1627]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="2" pageNumber="563">26, 29</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3EFFA2FDE5E9272783F988" box="[610,683,1601,1627]" captionStart="FIGURES 31−35" captionStartId="12.[151,264,1743,1767]" captionTargetBox="[155,1434,184,1706]" captionTargetId="figure-21@12.[151,1435,181,1718]" captionTargetPageId="12" captionText="FIGURES 31−35. Female abdomen (upper side, tergites; lower side, sternites). 31, Agripialus campos sp. n., PT (CGCM 31.875). 32, A. variabilis sp. n., HT. 33, A. itatiaia sp. n., PT (CGCM 25.532). 34, Mutipialus dilatus sp. n., HT (CGCM 5.801). 35, M. monticolus sp. n., HT." figureDoi="http://doi.org/10.5281/zenodo.5224155" httpUri="https://zenodo.org/record/5224155/files/figure.png" pageId="2" pageNumber="563">31–33</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3EFFA2FD31E92727D6F988" box="[694,766,1601,1627]" captionStart="FIGURES 36−46" captionStartId="13.[151,264,1743,1767]" captionTargetBox="[251,1340,182,1714]" captionTargetId="figure-20@13.[247,1340,181,1719]" captionTargetPageId="13" captionText="FIGURES 36−46. Abdomen. Figs 36–39. Female segments VI (lower)–VIII (upper). 36, Agripialus campos sp. n., PT (CGCM 31.875); arrows pointing to the sclerotised sternites VII and VIII. 37, A. variabilis sp. n., HT. 38, Mutipialus dilatus sp. n., HT (CGCM 5.801). 39, M. monticolus sp. n., HT; arrow pointing to the membranous sternite VIII. Figs 40–42. Male abdomen (upper side, tergites; lower side, sternites). 40, Agripialus caparao sp. n., HT. 41, M. dilatus sp. n., PT (CGCM 27.435). 42, M. monticolus sp. n., PT (CGCM 29.697). Figs 43–46. Male segments VII (lower)–VIII (upper). 43, Agripialus itatiaia sp. n., HT; arrows pointing to the two plates of the sternite VII. 44, A. caparao sp. n., HT; arrows pointing to the two plates of the sternite VII. 45, Mutipialus dilatus sp. n., PT (CGCM 27.435); arrow pointing to the sternite VIII. 46, M. monticolus sp. n., PT (CGCM 29.697); arrow pointing to the sternite VIII." figureDoi="http://doi.org/10.5281/zenodo.5224157" httpUri="https://zenodo.org/record/5224157/files/figure.png" pageId="2" pageNumber="563">36–37</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3EFFA2FC8DE927267DF98F" box="[778,853,1601,1628]" captionStart="FIGURES 51−57" captionStartId="15.[151,264,1627,1651]" captionTargetBox="[162,1427,289,1591]" captionTargetId="figure-49@15.[151,1437,272,1603]" captionTargetPageId="15" captionText="FIGURES 51−57. Female genitalia. 51, Agripialus campos sp. n., PT: posterior view and ductus and corpus bursae; arrow pointing to W-shaped lamella antevaginalis. 52, A. variabilis sp. n., HT: posterior view and ductus and corpus bursae. 53, A. itatiaia sp. n., PT (CGCM 25.532): posterior view and ductus and corpus bursae. 54, Mutipialus dilatus sp. n., HT (CGCM 5.801): posterior (54a), lateral (54b); arrow pointing to slight mesal projection of the lamella antevaginalis. 55, M. monticolus sp. n., HT: posterior (55a), lateral (55b); arrow pointing to slight mesal projection of the lamella antevaginalis. 56, Mutipialus dilatus sp. n., HT (CGCM 5.801): ductus and corpus bursae. 57, M. monticolus sp. n., HT: ductus and corpus bursae." figureDoi="http://doi.org/10.5281/zenodo.5224161" httpUri="https://zenodo.org/record/5224161/files/figure.png" pageId="2" pageNumber="563">51–53</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3EFFA2FF40E90324B4F9AC" blockId="2.[151,1437,1601,2024]" box="[199,412,1637,1664]" pageId="2" pageNumber="563">
|
||
<emphasis id="4BACEADBEE3EFFA2FF40E903242CF9AC" box="[199,260,1637,1663]" italics="true" pageId="2" pageNumber="563">Head</emphasis>
|
||
. As for male.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3EFFA2FF40E9EF21ADF970" blockId="2.[151,1437,1601,2024]" box="[199,1157,1673,1700]" pageId="2" pageNumber="563">
|
||
<emphasis id="4BACEADBEE3EFFA2FF40E9EF2408F970" box="[199,288,1673,1699]" italics="true" pageId="2" pageNumber="563">Thorax.</emphasis>
|
||
Legs (
|
||
<figureCitation id="E1E32A4CEE3EFFA2FEECE9EF24C7F970" box="[363,495,1673,1699]" captionStart="FIGURES 16−24" captionStartId="10.[151,264,1874,1898]" captionTargetBox="[242,1332,181,1847]" captionTargetId="figure-21@10.[227,1361,181,1850]" captionTargetPageId="10" captionText="FIGURES 16−24. Transverse scales and legs. Figs 16–19. Transverse scales (arrowed). 16, Mutipialus dilatus sp. n. PT ♂ (CGCM 23.689). 17, M. monticolus sp. n., HT ♀. 18, Parapielus luteicornis ♂ (CGCM 28.544, Argentina, Rio Grande; CGCM). 19, Hepialiscus htayaungi ♂ (CGCM 33.586, Myanmar; CGCM). Figs 20–24. Legs. 20, Agripialus campos sp. n., PT ♀ (CGCM 31.875). 21, A. variabilis sp. n., HT ♀. 22, A. itatiaia sp. n., HT ♂. 23, A. caparao sp. n., HT ♂. 24, Mutipialus dilatus sp. n., PT ♂ (CGCM 27.435)." figureDoi="http://doi.org/10.5281/zenodo.5224149" httpUri="https://zenodo.org/record/5224149/files/figure.png" pageId="2" pageNumber="563">Figs 20–21</figureCitation>
|
||
): as for male. Venation (
|
||
<figureCitation id="E1E32A4CEE3EFFA2FD78E9EF267BF977" box="[767,851,1673,1700]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="2" pageNumber="563">Fig. 26</figureCitation>
|
||
): 1A, 2A and 3A complete.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3EFFA2FF40E9CB27A2F914" blockId="2.[151,1437,1601,2024]" box="[199,650,1709,1735]" pageId="2" pageNumber="563">
|
||
<emphasis id="4BACEADBEE3EFFA2FF40E9CB2419F914" box="[199,305,1709,1735]" italics="true" pageId="2" pageNumber="563">Abdomen</emphasis>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3EFFA2FEC7E9CB24BFF914" box="[320,407,1709,1735]" captionStart="FIGURES 25−30" captionStartId="11.[151,264,1909,1933]" captionTargetBox="[212,1387,181,1885]" captionTargetId="figure-20@11.[201,1387,181,1885]" captionTargetPageId="11" captionText="FIGURES 25−30. Wing venation and tergosternal sclerite. Figs 25–28. Wing venation. 25, Agripialus caparao sp. n., ♂. 26, A. itatiaia sp. n., ♀. 27–28, Mutipialus dilatus sp. n., ♂ (27), ♀ (28); arrow pointing to the connection between CuP and A veins in forewing. Figs 29–30. Tergosternal sclerite. 29, Agripialus variabilis sp. n., ♀; arrows pointing to the notch and the posterior portion. 30, Mutipialus dilatus sp. n., ♂." figureDoi="http://doi.org/10.5281/zenodo.5224153" httpUri="https://zenodo.org/record/5224153/files/figure.png" pageId="2" pageNumber="563">Figs 29</figureCitation>
|
||
,
|
||
<figureCitation id="E1E32A4CEE3EFFA2FE24E9CB24C4F914" box="[419,492,1709,1735]" captionStart="FIGURES 31−35" captionStartId="12.[151,264,1743,1767]" captionTargetBox="[155,1434,184,1706]" captionTargetId="figure-21@12.[151,1435,181,1718]" captionTargetPageId="12" captionText="FIGURES 31−35. Female abdomen (upper side, tergites; lower side, sternites). 31, Agripialus campos sp. n., PT (CGCM 31.875). 32, A. variabilis sp. n., HT. 33, A. itatiaia sp. n., PT (CGCM 25.532). 34, Mutipialus dilatus sp. n., HT (CGCM 5.801). 35, M. monticolus sp. n., HT." figureDoi="http://doi.org/10.5281/zenodo.5224155" httpUri="https://zenodo.org/record/5224155/files/figure.png" pageId="2" pageNumber="563">31–33</figureCitation>
|
||
). As for male.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3EFFA2FF40E9B726BFF8E0" blockId="2.[151,1437,1601,2024]" pageId="2" pageNumber="563">
|
||
<emphasis id="4BACEADBEE3EFFA2FF40E9B72419F938" box="[199,305,1745,1771]" italics="true" pageId="2" pageNumber="563">Genitalia</emphasis>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3EFFA2FEB8E9B724EAF93F" box="[319,450,1745,1772]" captionStart="FIGURES 51−57" captionStartId="15.[151,264,1627,1651]" captionTargetBox="[162,1427,289,1591]" captionTargetId="figure-49@15.[151,1437,272,1603]" captionTargetPageId="15" captionText="FIGURES 51−57. Female genitalia. 51, Agripialus campos sp. n., PT: posterior view and ductus and corpus bursae; arrow pointing to W-shaped lamella antevaginalis. 52, A. variabilis sp. n., HT: posterior view and ductus and corpus bursae. 53, A. itatiaia sp. n., PT (CGCM 25.532): posterior view and ductus and corpus bursae. 54, Mutipialus dilatus sp. n., HT (CGCM 5.801): posterior (54a), lateral (54b); arrow pointing to slight mesal projection of the lamella antevaginalis. 55, M. monticolus sp. n., HT: posterior (55a), lateral (55b); arrow pointing to slight mesal projection of the lamella antevaginalis. 56, Mutipialus dilatus sp. n., HT (CGCM 5.801): ductus and corpus bursae. 57, M. monticolus sp. n., HT: ductus and corpus bursae." figureDoi="http://doi.org/10.5281/zenodo.5224161" httpUri="https://zenodo.org/record/5224161/files/figure.png" pageId="2" pageNumber="563">Figs 51–53</figureCitation>
|
||
). Each dorsal plate (tergite IX) dorsally separated, other than a sclerotised dorsal ridge or a thickened membrane; ventrally separated from lamella antevaginalis. Latter W-shaped with mesal fusion of each of lateral plates. Ductus bursae tube-like and corpus bursae globular.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31C26542EE3EFFA3FF40E85B2722FF2A" lastPageId="3" lastPageNumber="564" pageId="2" pageNumber="563" type="etymology">
|
||
<paragraph id="796736C9EE3EFFA2FF40E85B219FF84C" blockId="2.[151,1437,1601,2024]" pageId="2" pageNumber="563">
|
||
<emphasis id="4BACEADBEE3EFFA2FF40E85B2460F884" bold="true" box="[199,328,1853,1879]" pageId="2" pageNumber="563">Etymology</emphasis>
|
||
.
|
||
<taxonomicName id="BED84D4AEE3EFFA2FEDFE85B24E6F884" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[344,462,1853,1879]" class="Insecta" family="Hepialidae" genus="Agripialus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="4BACEADBEE3EFFA2FEDFE85B24E6F884" box="[344,462,1853,1879]" italics="true" pageId="2" pageNumber="563">Agripialus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="509F57A0EE3EFFA2FE50E85B270FF88B" box="[471,551,1853,1880]" pageId="2" pageNumber="563" rank="genus">
|
||
<emphasis id="4BACEADBEE3EFFA2FE50E85B2729F884" bold="true" box="[471,513,1853,1879]" pageId="2" pageNumber="563">gen</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3EFFA2FD96E85B2709F884" bold="true" box="[529,545,1853,1879]" pageId="2" pageNumber="563">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
comes from the Latin prefix
|
||
<emphasis id="4BACEADBEE3EFFA2FCFDE85B2684F884" box="[890,940,1853,1879]" italics="true" pageId="2" pageNumber="563">Agri</emphasis>
|
||
- (field, open land) added ‘
|
||
<emphasis id="4BACEADBEE3EFFA2FB66E85B200DF884" box="[1249,1317,1853,1879]" italics="true" pageId="2" pageNumber="563">pialus</emphasis>
|
||
’ extracted from the first described
|
||
<taxonomicName id="BED84D4AEE3EFFA2FE24E8072734F8A8" box="[419,540,1889,1915]" class="Insecta" family="Hepialidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="family">Hepialidae</taxonomicName>
|
||
genus
|
||
<taxonomicName id="BED84D4AEE3EFFA2FDECE8072651F8A8" authority="Fabricius. The" authorityName="Fabricius. The" authorityYear="1775" box="[619,889,1889,1915]" class="Insecta" family="Hepialidae" genus="Hepialus" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="563" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4BACEADBEE3EFFA2FDECE80727F8F8A8" box="[619,720,1889,1915]" italics="true" pageId="2" pageNumber="563">Hepialus</emphasis>
|
||
Fabricius. The
|
||
</taxonomicName>
|
||
name is a reference of the biotope where all species described below have been found, areas with natural grasslands. The gender is masculine.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3EFFA3FF40E8CF2722FF2A" blockId="2.[151,1437,1601,2024]" lastBlockId="3.[151,1437,150,501]" lastPageId="3" lastPageNumber="564" pageId="2" pageNumber="563">
|
||
<emphasis id="4BACEADBEE3EFFA2FF40E8CF2728F810" bold="true" box="[199,512,1961,1987]" pageId="2" pageNumber="563">Geographical distribution</emphasis>
|
||
.
|
||
<materialsCitation id="C9B03C94EE3EFFA3FD88E8CC272EFF2A" country="Brazil" lastPageId="3" lastPageNumber="564" location="Tucuman" pageId="2" pageNumber="563" specimenCount="1" stateProvince="Tucuman">
|
||
Known from eastern Minas Gerais to Rio Grande do Sul in
|
||
<collectingCountry id="01CF7659EE3EFFA2FB38E8CF202BF810" box="[1215,1283,1961,1987]" name="Brazil" pageId="2" pageNumber="563">Brazil</collectingCountry>
|
||
and likely to northeastern (
|
||
<collectingRegion id="BB1CF82BEE3EFFA2FEB6E8AB24B5F834" box="[305,413,1997,2023]" country="Argentina" name="Tucuman" pageId="2" pageNumber="563">Tucumán</collectingRegion>
|
||
) and eastern (
|
||
<collectingRegion id="BB1CF82BEE3EFFA2FDBAE8AB27FFF83B" box="[573,727,1997,2024]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="2" pageNumber="563">Buenos Aires</collectingRegion>
|
||
)
|
||
<collectingCountry id="01CF7659EE3EFFA2FD61E8AB267DF834" box="[742,853,1997,2023]" name="Argentina" pageId="2" pageNumber="563">Argentina</collectingCountry>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3EFFA2FCE1E8AB2694F83B" box="[870,956,1997,2024]" captionStart="FIGURE 58" captionStartId="16.[151,250,1100,1124]" captionTargetBox="[169,1335,181,1076]" captionTargetId="figure-21@16.[169,1418,181,1076]" captionTargetPageId="16" captionText="FIGURE 58. Distribution of Agripialus gen. n. spp. and Mutipialus gen. n. spp. within southern southeastern and southern Brazil and records of Agripialus gen. n. in Argentina (see remarks at the description of the genus)." figureDoi="http://doi.org/10.5281/zenodo.5224165" httpUri="https://zenodo.org/record/5224165/files/figure.png" pageId="2" pageNumber="563">Fig. 58</figureCitation>
|
||
). It appears to be restricted to areas where grasslands are part of the natural vegetation (
|
||
<figureCitation id="E1E32A4CEE3FFFA3FD11EFF12632FF62" box="[662,794,150,177]" captionStart="FIGURES 59−62" captionStartId="17.[151,264,1727,1751]" captionTargetBox="[175,1411,489,1704]" captionTargetId="figure-162@17.[175,1411,489,1704]" captionTargetPageId="17" captionText="FIGURES 59−62. Habitats in Brazil. 59, Caparaó National Park, 2200 m, Espera Feliz, Minas Gerais (Agripialus caparao sp. n.). 60, Lavrinhas, 1900 m, Campos do Jordão, São Paulo (A. campos sp. n., Mutipialus dilatus sp. n., and M. monticolus sp. n.). 61, Itatiaia National Park, 2400 m, Itatiaia, Rio de Janeiro (A. itatiaia sp. n.). 62, São Joaquim National Park, 1600 m, Urubici, Santa Catarina (A. variabilis sp. n.)." figureDoi="http://doi.org/10.5281/zenodo.5224167" httpUri="https://zenodo.org/record/5224167/files/figure.png" pageId="3" pageNumber="564">Figs 59–62</figureCitation>
|
||
). Such biotopes are typical of humid subtropical areas at moderate to high altitudes. Although not examined, two records are known to us from
|
||
<collectingCountry id="01CF7659EE3FFFA3FBE3EFDD21FFFF06" box="[1124,1239,187,213]" name="Argentina" pageId="3" pageNumber="564">Argentina</collectingCountry>
|
||
. Both records are not taxonomically treated herein
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="31C26542EE3FFFA3FF40EE652115FE26" pageId="3" pageNumber="564" type="discussion">
|
||
<paragraph id="796736C9EE3FFFA3FF40EE652155FECE" blockId="3.[151,1437,150,501]" box="[199,1149,259,285]" pageId="3" pageNumber="564">
|
||
<emphasis id="4BACEADBEE3FFFA3FF40EE65241CFECE" bold="true" box="[199,308,259,285]" pageId="3" pageNumber="564">Remarks</emphasis>
|
||
. All
|
||
<taxonomicName id="BED84D4AEE3FFFA3FEECEE6524C9FECE" authority="Mielke & Grehan & Koike, 2021" authorityName="Mielke & Grehan & Koike" authorityYear="2021" box="[363,481,259,285]" class="Insecta" family="Hepialidae" genus="Agripialus" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="564" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="4BACEADBEE3FFFA3FEECEE6524C9FECE" box="[363,481,259,285]" italics="true" pageId="3" pageNumber="564">Agripialus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="509F57A0EE3FFFA3FE6EEE65271FFECE" box="[489,567,259,285]" pageId="3" pageNumber="564" rank="genus">
|
||
<emphasis id="4BACEADBEE3FFFA3FE6EEE65273BFECE" bold="true" box="[489,531,259,285]" pageId="3" pageNumber="564">gen</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3FFFA3FDA6EE652719FECE" bold="true" box="[545,561,259,285]" pageId="3" pageNumber="564">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
species are monovoltine in late spring and summer.
|
||
</paragraph>
|
||
<paragraph id="796736C9EE3FFFA3FF40EE412115FE26" blockId="3.[151,1437,150,501]" pageId="3" pageNumber="564">
|
||
A single female specimen potentially of
|
||
<emphasis id="4BACEADBEE3FFFA3FD15EE412620FE92" box="[658,776,295,321]" italics="true" pageId="3" pageNumber="564">Agripialus</emphasis>
|
||
<taxonomicNameLabel id="509F57A0EE3FFFA3FC97EE412678FE92" box="[784,848,295,321]" pageId="3" pageNumber="564" rank="species">
|
||
<emphasis id="4BACEADBEE3FFFA3FC97EE412602FE92" bold="true" box="[784,810,295,321]" pageId="3" pageNumber="564">sp</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3FFFA3FCBEEE412661FE92" bold="true" box="[825,841,295,321]" pageId="3" pageNumber="564">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from northeastern
|
||
<collectingCountry id="01CF7659EE3FFFA3FBA0EE4121BEFE92" box="[1063,1174,295,321]" name="Argentina" pageId="3" pageNumber="564">Argentina</collectingCountry>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3FFFA3FB20EE4121D2FE92" box="[1191,1274,294,321]" captionStart="FIGURE 58" captionStartId="16.[151,250,1100,1124]" captionTargetBox="[169,1335,181,1076]" captionTargetId="figure-21@16.[169,1418,181,1076]" captionTargetPageId="16" captionText="FIGURE 58. Distribution of Agripialus gen. n. spp. and Mutipialus gen. n. spp. within southern southeastern and southern Brazil and records of Agripialus gen. n. in Argentina (see remarks at the description of the genus)." figureDoi="http://doi.org/10.5281/zenodo.5224165" httpUri="https://zenodo.org/record/5224165/files/figure.png" pageId="3" pageNumber="564">Fig. 58</figureCitation>
|
||
, white circle), not dissected, was not included because this is an article dedicated to the Brazilian fauna. If indeed confirmed, the distribution of the genus would not only be greatly expanded but also disjunct between the Andes and Atlantic coast. A second female from another potential
|
||
<emphasis id="4BACEADBEE3FFFA3FDD4EEF527E1FE7E" box="[595,713,403,429]" italics="true" pageId="3" pageNumber="564">Agripialus</emphasis>
|
||
<taxonomicNameLabel id="509F57A0EE3FFFA3FD48EEF52625FE7E" box="[719,781,403,429]" pageId="3" pageNumber="564" rank="species">
|
||
<emphasis id="4BACEADBEE3FFFA3FD48EEF527C1FE7E" bold="true" box="[719,745,403,429]" pageId="3" pageNumber="564">sp</emphasis>
|
||
.
|
||
<emphasis id="4BACEADBEE3FFFA3FD70EEF5262FFE7E" bold="true" box="[759,775,403,429]" pageId="3" pageNumber="564">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
from south of
|
||
<collectingRegion id="BB1CF82BEE3FFFA3FC28EEF5216CFE7E" box="[943,1092,403,429]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="3" pageNumber="564">Buenos Aires</collectingRegion>
|
||
in
|
||
<collectingCountry id="01CF7659EE3FFFA3FBE3EEF521FBFE7E" box="[1124,1235,403,429]" name="Argentina" pageId="3" pageNumber="564">Argentina</collectingCountry>
|
||
(
|
||
<figureCitation id="E1E32A4CEE3FFFA3FB66EEF5201AFE7E" box="[1249,1330,402,429]" captionStart="FIGURE 58" captionStartId="16.[151,250,1100,1124]" captionTargetBox="[169,1335,181,1076]" captionTargetId="figure-21@16.[169,1418,181,1076]" captionTargetPageId="16" captionText="FIGURE 58. Distribution of Agripialus gen. n. spp. and Mutipialus gen. n. spp. within southern southeastern and southern Brazil and records of Agripialus gen. n. in Argentina (see remarks at the description of the genus)." figureDoi="http://doi.org/10.5281/zenodo.5224165" httpUri="https://zenodo.org/record/5224165/files/figure.png" pageId="3" pageNumber="564">Fig. 58</figureCitation>
|
||
, question mark) was presented to us by Ezequiel Núñez Bustos (
|
||
<collectingRegion id="BB1CF82BEE3FFFA3FD7EEED126BAFE01" box="[761,914,439,466]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="3" pageNumber="564">Buenos Aires</collectingRegion>
|
||
) through an image only. Such a record, almost at sea level, would represent a further range boundary within the grassland biotope.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |