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<document id="3DE3C68E05F06A85EFAAEA152AE58675" ID-DOI="10.5281/zenodo.5276330" ID-ISSN="0253-116X" ID-Zenodo-Dep="5276330" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630012189340" checkinUser="carolina" docAuthor="Wrase, DW" docDate="2009" docId="EF5F980D0940FFC0FF1B1917730BFB0D" docLanguage="en" docName="LinzerbiolBeitr.2009.41.1.901-935.pdf" docOrigin="Linzer biologische Beiträge 41 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Harpalus sulphuripes subsp. creticus MARAN 1934" docType="treatment" docVersion="3" lastPageNumber="906" masterDocId="1366E0750944FFC5FFAA1A097737FF81" masterDocTitle="New or interesting records of Carabid beetles from Europe, Madeira, northern Africa, Turkey, from the Near East, Iran, Iraq, Kuwait, and Pakistan, with nomenclatorial and taxonomic notes (Coleoptera, Carabidae, Bembidiini, Brachinini, Cyclosomini, Elaphrini, Harpalini, Lebiini, Nebriini, Platynini," masterLastPageNumber="935" masterPageNumber="901" pageNumber="905" updateTime="1699136774480" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="764F129F84B0868C8E6345FE8595A67B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BC0268CCA19360342A3C788AAFEB485E">
<mods:title id="DF669D29C317C4139BD3CBB45DF76121">New or interesting records of Carabid beetles from Europe, Madeira, northern Africa, Turkey, from the Near East, Iran, Iraq, Kuwait, and Pakistan, with nomenclatorial and taxonomic notes (Coleoptera, Carabidae, Bembidiini, Brachinini, Cyclosomini, Elaphrini, Harpalini, Lebiini, Nebriini, Platynini,</mods:title>
</mods:titleInfo>
<mods:name id="8C561690E5AE884FF364F024A6E2A83A" type="personal">
<mods:role id="321DC90E54869671894EED617B1240BB">
<mods:roleTerm id="507813507E5A695DC1D11300AF5BDBC6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4CF901A5439B9AB75212D383362C12B8">Wrase, DW</mods:namePart>
</mods:name>
<mods:typeOfResource id="53558746AC9C735DE77D23AF850753B2">text</mods:typeOfResource>
<mods:relatedItem id="CA4FD4EE6F8892A84DF0B395F1A59F88" type="host">
<mods:titleInfo id="967012020F48A8AA9460189C07928436">
<mods:title id="BAB663CFA5B6EF9D3ADBCA3D74393356">Linzer biologische Beiträge</mods:title>
</mods:titleInfo>
<mods:part id="E6D05C989CAA89242814A5492C8D503E">
<mods:date id="29B7A27DB2C52432FFE683E2DCDC15CF">2009</mods:date>
<mods:detail id="4EA3DC7AAA11589B776F5A7E8873C007" type="pubDate">
<mods:number id="61AABA01E7D3F5B36EB834A77C21B9E9">2009-08-30</mods:number>
</mods:detail>
<mods:detail id="789837F01E24197FEB0D5E4B5BBDEC4F" type="volume">
<mods:number id="23C41BE5D1284650887200C5203EAEAF">41</mods:number>
</mods:detail>
<mods:detail id="D9EBFD352B10D4D89BD87CFF510E46FD" type="issue">
<mods:number id="496219031B2511110B0BD64FA09D9A10">1</mods:number>
</mods:detail>
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<mods:start id="83830C7CE29290944C219953B606C6B0">901</mods:start>
<mods:end id="5F8EDB158DA5548A8EB4974E539BD7B3">935</mods:end>
</mods:extent>
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<mods:classification id="2D726BCBED3FC989A639E3696C560DC4">journal article</mods:classification>
<mods:identifier id="1F5C20F3BE74C12FFBECB38D6B784EB8" type="DOI">10.5281/zenodo.5276330</mods:identifier>
<mods:identifier id="67AE4D0787F2F1F5FD79F799BAD825ED" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="E0896BE2B98D35A8F7494C145E78A613" type="Zenodo-Dep">5276330</mods:identifier>
</mods:mods>
<treatment id="EF5F980D0940FFC0FF1B1917730BFB0D" LSID="urn:lsid:plazi:treatment:EF5F980D0940FFC0FF1B1917730BFB0D" httpUri="http://treatment.plazi.org/id/EF5F980D0940FFC0FF1B1917730BFB0D" lastPageId="5" lastPageNumber="906" pageId="4" pageNumber="905">
<subSubSection id="2FEC7A900940FFC1FF1B19177499FCB5" box="[177,942,798,820]" pageId="4" pageNumber="905" type="nomenclature">
<paragraph id="6749291B0940FFC1FF1B19177499FCB5" blockId="4.[177,942,798,852]" box="[177,942,798,820]" pageId="4" pageNumber="905">
<heading id="3C019E770940FFC1FF1B19177499FCB5" box="[177,942,798,820]" fontSize="8" level="2" pageId="4" pageNumber="905" reason="2">
<emphasis id="5582F5090940FFC1FF1B19177499FCB5" bold="true" box="[177,942,798,820]" pageId="4" pageNumber="905">
<taxonomicName id="A0F652980940FFC1FF1B191775CFFCB5" ID-CoL="8MJCT" authority="MARAN 1934" authorityName="MARAN" authorityYear="1934" box="[177,760,798,820]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="905" phylum="Arthropoda" rank="subSpecies" species="sulphuripes" subSpecies="creticus">
<emphasis id="5582F5090940FFC1FF1B19177624FCB5" bold="true" box="[177,275,798,820]" italics="true" pageId="4" pageNumber="905">Harpalus</emphasis>
(s.str.)
<emphasis id="5582F5090940FFC1FECF191776E8FCB5" bold="true" box="[357,479,798,820]" italics="true" pageId="4" pageNumber="905">sulphuripes</emphasis>
ssp.
<emphasis id="5582F5090940FFC1FDBE19177552FCB5" bold="true" box="[532,613,798,820]" italics="true" pageId="4" pageNumber="905">creticus</emphasis>
<bibRefCitation id="036754EA0940FFC1FDC61917758AFCB2" author="MARAN J" box="[620,701,798,820]" pageId="4" pageNumber="905" pagination="125 - 128" refId="ref25534" refString="MARAN J. (1934): Nove druhy a formy Carabidu z ostrova Hrety. De novis carabidarum speciebus formis-que ex insula Creta. - Sbornik Entomologickeho oddeleni Narodniho Musea v Praze 12: 125 - 128." type="journal article" year="1934">MAŘAN</bibRefCitation>
1934
</taxonomicName>
, new assignment
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2FEC7A900940FFC1FF1B19497668FBE7" pageId="4" pageNumber="905" type="reference_group">
<paragraph id="6749291B0940FFC1FF1B194975B5FCD5" blockId="4.[177,942,798,852]" box="[177,642,831,852]" pageId="4" pageNumber="905">
<treatmentCitationGroup id="47E60E350940FFC1FF1B194975B5FCD5" box="[177,642,831,852]" pageId="4" pageNumber="905">
<treatmentCitation id="E6570F0A0940FFC1FF1B194975B5FCD5" author="MARAN J" box="[177,642,831,852]" page="126" pageId="4" pageNumber="905" year="1934">
<taxonomicName id="A0F652980940FFC1FF1B194975B5FCD5" authority="MARAN 1934: 126" authorityName="MARAN" authorityPageNumber="126" authorityYear="1934" box="[177,642,831,852]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="905" phylum="Arthropoda" rank="subSpecies" species="honestus" subSpecies="creticus">
<emphasis id="5582F5090940FFC1FF1B1949766BFCD5" box="[177,348,832,852]" italics="true" pageId="4" pageNumber="905">Harpalus honestus</emphasis>
ssp.
<emphasis id="5582F5090940FFC1FE20194976E5FCD5" box="[394,466,832,852]" italics="true" pageId="4" pageNumber="905">creticus</emphasis>
<bibRefCitation id="036754EA0940FFC1FE72194975B5FCD5" author="MARAN J" box="[472,642,831,852]" pageId="4" pageNumber="905" pagination="125 - 128" refId="ref25534" refString="MARAN J. (1934): Nove druhy a formy Carabidu z ostrova Hrety. De novis carabidarum speciebus formis-que ex insula Creta. - Sbornik Entomologickeho oddeleni Narodniho Musea v Praze 12: 125 - 128." type="journal article" year="1934">MAŘAN 1934: 126</bibRefCitation>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
<paragraph id="6749291B0940FFC1FF1B196B7668FBE7" blockId="4.[177,1086,865,1555]" pageId="4" pageNumber="905">
T y p e m a t e r i a l:
<typeStatus id="B84D97B90940FFC1FE0C196B76CAFCF7" box="[422,509,866,886]" pageId="4" pageNumber="905" type="lectotype">Lectotype</typeStatus>
3, herewith designated, labelled: &quot;Nida – Ida montes /
<collectingRegion id="A532E7F90940FFC1FC47196B7329FCF7" box="[1005,1054,866,886]" country="Greece" name="Kriti" pageId="4" pageNumber="905">Creta</collectingRegion>
, V, 1934 / Coll. Bartoň&quot; (black print on white label); &quot;TYPUS&quot; (black printed and framed on red label); &quot;
<taxonomicName id="A0F652980940FFC1FF4D199B7667FC27" authorityName="MARAN" authorityYear="1934" box="[231,336,914,934]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="905" phylum="Arthropoda" rank="subSpecies" species="honestus" subSpecies="creticus">ssp. creticus</taxonomicName>
m. / Dr. Mařan det.&quot; (white label, first line written by Mařan, second one printed); &quot;
<taxonomicName id="A0F652980940FFC1FB8D199B7616FC3F" authorityName="MARAN" authorityYear="1934" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="905" phylum="Arthropoda" rank="species" species="creticus">H. creticus</taxonomicName>
m. 3 / Dr. J. Mařan det. 1934&quot; (white label, first line handwritten by Mařan, second one printed; &quot;34&quot; handwritten); two subsequent labels: &quot;
<collectingRegion id="A532E7F90940FFC1FD3519CB75FDFC57" box="[671,714,962,982]" country="Turkey" name="Mus" pageId="4" pageNumber="905">Mus</collectingRegion>
. Nat. Pragae / Inv. 857 (printed on antique pink, number handwritten), and &quot;
<typeStatus id="B84D97B90940FFC1FE5119D37563FC6F" box="[507,596,986,1006]" pageId="4" pageNumber="905" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="A0F652980940FFC1FDF319D375CFFC6F" ID-CoL="3JSLH" baseAuthorityName="DUFTSCHMIDT" baseAuthorityYear="1812" box="[601,760,986,1006]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="905" phylum="Arthropoda" rank="species" species="honestus">Harpalus honestus</taxonomicName>
<taxonomicName id="A0F652980940FFC1FD5719D37321FC6F" authority="MARAN, 1934 WRASE" authorityName="MARAN" authorityYear="1934" box="[765,1046,985,1006]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="905" phylum="Arthropoda" rank="species" species="creticus">
creticus
<bibRefCitation id="036754EA0940FFC1FCEF19D374F4FC6F" author="MARAN J" box="[837,963,985,1006]" pageId="4" pageNumber="905" pagination="125 - 128" refId="ref25534" refString="MARAN J. (1934): Nove druhy a formy Carabidu z ostrova Hrety. De novis carabidarum speciebus formis-que ex insula Creta. - Sbornik Entomologickeho oddeleni Narodniho Musea v Praze 12: 125 - 128." type="journal article" year="1934">MAŘAN, 1934</bibRefCitation>
WRASE
</taxonomicName>
des. 2008&quot; (black print on red label).
<typeStatus id="B84D97B90940FFC1FDAB19FB75B5FB87" box="[513,642,1010,1030]" pageId="4" pageNumber="905" type="paralectotype">Paralectotypes</typeStatus>
: 13,
<specimenCount id="71F0E2920940FFC1FD1F19FB75E4FB8A" box="[693,723,1010,1035]" pageId="4" pageNumber="905" type="female">1♀</specimenCount>
, with same data as
<typeStatus id="B84D97B90940FFC1FC2819FB74E6FB87" box="[898,977,1010,1030]" pageId="4" pageNumber="905" type="lectotype">lectotype</typeStatus>
but without third label, fourth label for female specimen with a female sign instead of male sign, and with inventory number &quot;858&quot; (male), and &quot;895&quot; (female), and with
<typeStatus id="B84D97B90940FFC1FC801E2B74A9FBB7" box="[810,926,1058,1078]" pageId="4" pageNumber="905" type="paralectotype">paralectotype</typeStatus>
labels. All
<specimenCount id="71F0E2920940FFC1FBB81E2B760EFBCF" pageId="4" pageNumber="905" type="generic">three specimens</specimenCount>
with label &quot;
<taxonomicName id="A0F652980940FFC1FE0A1E33756EFBCF" ID-CoL="3JSRQ" authorityName="GERMAR" authorityYear="1824" box="[416,601,1082,1102]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="905" phylum="Arthropoda" rank="species" species="sulphuripes">Harpalus sulphuripes</taxonomicName>
<taxonomicName id="A0F652980940FFC1FDF71E337442FBCF" authority="MARAN, 1934 WRASE" authorityName="MARAN" authorityYear="1934" box="[605,885,1081,1102]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="905" phylum="Arthropoda" rank="species" species="creticus">
creticus
<bibRefCitation id="036754EA0940FFC1FD0F1E337415FBCF" author="MARAN J" box="[677,802,1081,1102]" pageId="4" pageNumber="905" pagination="125 - 128" refId="ref25534" refString="MARAN J. (1934): Nove druhy a formy Carabidu z ostrova Hrety. De novis carabidarum speciebus formis-que ex insula Creta. - Sbornik Entomologickeho oddeleni Narodniho Musea v Praze 12: 125 - 128." type="journal article" year="1934">MAŘAN, 1934</bibRefCitation>
WRASE
</taxonomicName>
det. 2008&quot; (black print on white label.
</paragraph>
</subSubSection>
<subSubSection id="2FEC7A900940FFC0FF1B1E7B730BFB0D" lastPageId="5" lastPageNumber="906" pageId="4" pageNumber="905" type="description">
<paragraph id="6749291B0940FFC1FF1B1E7B754AFAAF" blockId="4.[177,1086,865,1555]" pageId="4" pageNumber="905">
A d d i t i o n a l m a t e r i a l: G r e e c e:
<collectingRegion id="A532E7F90940FFC1FD381E7B7589FB07" box="[658,702,1138,1158]" country="Greece" name="Kriti" pageId="4" pageNumber="905">Kríti</collectingRegion>
: Lasithíou: Díkti Mts., Màles env.,
<quantity id="A00E84FE0940FFC1FBA81E7B730AFB07" box="[1026,1085,1138,1158]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="905" unit="km" value="15.0">15 km</quantity>
NW Ierápetra,
<date id="13480FDB0940FFC1FEC91E8376FEFB1F" box="[355,457,1162,1182]" pageId="4" pageNumber="905" value="2001-06-12">12.VI.2001</date>
, K. Orszulik leg. (
<specimenCount id="71F0E2920940FFC1FDDA1E8375BBFB22" box="[624,652,1162,1187]" pageId="4" pageNumber="905" type="female">1♀</specimenCount>
, cORSZ). - Haníon: W
<collectingRegion id="A532E7F90940FFC1FCCF1E8374A1FB1F" box="[869,918,1162,1182]" country="Greece" name="Kriti" pageId="4" pageNumber="905">Crete</collectingRegion>
, Lefka Óri Mts.,
<quantity id="A00E84FE0940FFC1FB981E8377CCFB37" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="905" unit="km" value="4.0">4 km</quantity>
S Omalos,
<quantity id="A00E84FE0940FFC1FEC81EAB7690FB37" box="[354,423,1186,1206]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="905" unit="m" value="1400.0">1400 m</quantity>
,
<date id="13480FDB0940FFC1FE1B1EAB753CFB37" box="[433,523,1186,1206]" pageId="4" pageNumber="905" value="1990-04-01">1.IV.1990</date>
, R.Borovec leg. (13, cWR).
<collectingRegion id="A532E7F90940FFC1FCA41EAB7408FB37" box="[782,831,1186,1206]" country="Greece" name="Kriti" pageId="4" pageNumber="905">Crete</collectingRegion>
, Levka Ori, E Omalos,
<quantity id="A00E84FE0940FFC1FBBB1EAB77C6FB4F" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="905" unit="m" value="1600.0">1600 m</quantity>
, N 35°19’15’’/E
<geoCoordinate id="02C24FDC0940FFC1FE2F1EB376C3FB4F" box="[389,500,1210,1230]" degrees="23" direction="east" minutes="55" orientation="longitude" pageId="4" pageNumber="905" precision="15" seconds="56" value="23.932222">23°55’56’’E</geoCoordinate>
,
<date id="13480FDB0940FFC1FE541EB3756BFB4F" box="[510,604,1210,1230]" pageId="4" pageNumber="905" value="2004-05-27">27.V.2004</date>
, Chr. Bayer leg. (233, cWR).
<collectingRegion id="A532E7F90940FFC1FCDD1EB3749FFB4F" box="[887,936,1210,1230]" country="Greece" name="Kriti" pageId="4" pageNumber="905">Crete</collectingRegion>
, Lefka Óri Mts., Omalós,
<date id="13480FDB0940FFC1FE871EDB76A5FB67" box="[301,402,1234,1254]" pageId="4" pageNumber="905" value="1994-04-28">28.IV.1994</date>
, R. Kmeco leg. (
<specimenCount id="71F0E2920940FFC1FD8C1EDB7575FB6A" box="[550,578,1234,1259]" pageId="4" pageNumber="905" type="female">1♀</specimenCount>
, cKM). - Rethímnis:
<collectingRegion id="A532E7F90940FFC1FD511EDB7419FB67" box="[763,814,1234,1254]" country="Greece" name="Kriti" pageId="4" pageNumber="905">Kreta</collectingRegion>
, Ida-Gebirge, Umg. Ida-Höhle,
<quantity id="A00E84FE0940FFC1FF741EE37610FB7F" box="[222,295,1258,1278]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="905" unit="m" value="1500.0">1500 m</quantity>
,
<date id="13480FDB0940FFC1FE9E1EE376A5FB7F" box="[308,402,1258,1278]" pageId="4" pageNumber="905" value="1989-05-24">24.V.1989</date>
, H. Korge leg. (13, cKORG).
<collectingRegion id="A532E7F90940FFC1FD121EE375DEFB7F" box="[696,745,1258,1278]" country="Greece" name="Kriti" pageId="4" pageNumber="905">Crete</collectingRegion>
, Anogia,
<date id="13480FDB0940FFC1FCEF1EE374A3FB7F" box="[837,916,1258,1278]" pageId="4" pageNumber="905" value="2003-06">VI 2003</date>
, R. Hergovits leg. (233,
<specimenCount id="71F0E2920940FFC1FE8E1F0B7663FA9A" box="[292,340,1282,1307]" pageId="4" pageNumber="905" type="female">2♀♀</specimenCount>
, cLOH, cWR). W
<collectingRegion id="A532E7F90940FFC1FDA01F0B750CFA97" box="[522,571,1282,1302]" country="Greece" name="Kriti" pageId="4" pageNumber="905">Crete</collectingRegion>
, Nida-Hochebene,
<quantity id="A00E84FE0940FFC1FD401F0B7410FA97" box="[746,807,1282,1302]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="905" unit="km" value="15.0">15 km</quantity>
südlich von Anogia,
<quantity id="A00E84FE0940FFC1FC5B1F0B730EFA97" box="[1009,1081,1282,1302]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="905" unit="m" value="1500.0">1500 m</quantity>
,
<date id="13480FDB0940FFC1FF741F137606FAAF" box="[222,305,1306,1326]" pageId="4" pageNumber="905" value="2003-05-04">4.V.2003</date>
, W. Apfel leg. (233, NKME, cWR).
</paragraph>
<paragraph id="6749291B0940FFC1FF1B1F33730AFACE" blockId="4.[177,1086,865,1555]" box="[177,1085,1337,1359]" pageId="4" pageNumber="905">
Mařan’s description was based on about
<specimenCount id="71F0E2920940FFC1FDD51F307427FACE" box="[639,784,1337,1359]" pageId="4" pageNumber="905" type="generic">25 specimens</specimenCount>
coming from the &quot;Montes
</paragraph>
<paragraph id="6749291B0940FFC1FF1B1F5C730BFAEA" blockId="4.[177,1086,865,1555]" box="[177,1084,1365,1387]" pageId="4" pageNumber="905">
Psiloritis (
<collectingRegion id="A532E7F90940FFC1FEB41F5C7661FAEA" box="[286,342,1365,1387]" country="Greece" name="Kriti" pageId="4" pageNumber="905">Creta</collectingRegion>
centralis)&quot;, which were deposited in the collection of Cyril Bartoň-
</paragraph>
<paragraph id="6749291B0940FFC1FF1B1F78730BFA06" blockId="4.[177,1086,865,1555]" box="[177,1084,1393,1415]" pageId="4" pageNumber="905">Dobenín (stored today in NMP) and in NMP. He separated his taxon defined as subspe-</paragraph>
<paragraph id="6749291B0940FFC1FF1B1F84730BFA22" blockId="4.[177,1086,865,1555]" box="[177,1084,1421,1443]" pageId="4" pageNumber="905">
cies of
<taxonomicName id="A0F652980940FFC1FF551F877569FA22" authority="(DUFTSCHMIDT 1812)" baseAuthorityName="DUFTSCHMIDT" baseAuthorityYear="1812" box="[255,606,1421,1443]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="905" phylum="Arthropoda" rank="species" species="honestus">
<emphasis id="5582F5090940FFC1FF551F87764FFA22" box="[255,376,1421,1443]" italics="true" pageId="4" pageNumber="905">H. honestus</emphasis>
(DUFTSCHMIDT 1812)
</taxonomicName>
from the nominotypical
<taxonomicName id="A0F652980940FFC1FCCE1F8774E9FA22" baseAuthorityName="DUFTSCHMIDT" baseAuthorityYear="1812" box="[868,990,1421,1443]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="905" phylum="Arthropoda" rank="species" species="honestus">
<emphasis id="5582F5090940FFC1FCCE1F8774E9FA22" box="[868,990,1421,1443]" italics="true" pageId="4" pageNumber="905">H. honestus</emphasis>
</taxonomicName>
amongst
</paragraph>
<paragraph id="6749291B0940FFC1FF1B1FA0730AFA3E" blockId="4.[177,1086,865,1555]" box="[177,1085,1449,1471]" pageId="4" pageNumber="905">other features by more narrowed pronotum, laterally less sinuate, with anterior angles</paragraph>
<paragraph id="6749291B0940FFC1FF1B1FCF730BFA5A" blockId="4.[177,1086,865,1555]" box="[177,1084,1477,1499]" pageId="4" pageNumber="905">more rounded, and with posterior angles obtuse, by metepisterna distinctly shorter and by</paragraph>
<paragraph id="6749291B0940FFC1FF1B1FE8730AFA76" blockId="4.[177,1086,865,1555]" box="[177,1085,1505,1527]" pageId="4" pageNumber="905">rudimentary hind wings, median lobe stouter, with apical lamella shorter, and by colora-</paragraph>
<paragraph id="6749291B0940FFC1FF1B1FF4745AF992" blockId="4.[177,1086,865,1555]" box="[177,877,1533,1555]" pageId="4" pageNumber="905">tion in males always bluish black, and in females black, respectively.</paragraph>
<paragraph id="6749291B0940FFC0FF1B1C2B7318FEAA" blockId="4.[177,1085,1570,1592]" lastBlockId="5.[177,1085,219,299]" lastPageId="5" lastPageNumber="906" pageId="4" pageNumber="905">
After having asked for the type series I received
<specimenCount id="71F0E2920940FFC1FD691C2B746EF9B9" box="[707,857,1570,1592]" pageId="4" pageNumber="905" type="generic" typeStatus="syntypes">three syntypes</specimenCount>
from NMP, glued to cards and identified by their labels: &quot;Nida – Ida montes /
<collectingRegion id="A532E7F90941FFC0FCAA1AD2740EFF70" box="[768,825,219,241]" country="Greece" name="Kriti" pageId="5" pageNumber="906">Creta</collectingRegion>
, V, 1934 / Coll. Bartoň&quot;. The genitalia of
<specimenCount id="71F0E2920941FFC0FEF11AF0768FFE8F" box="[347,440,248,270]" pageId="5" pageNumber="906" type="male">one male</specimenCount>
were already extracted and glued to card, this male was choosen as
<typeStatus id="B84D97B90941FFC0FF5F1B1C7665FEAA" box="[245,338,277,299]" pageId="5" pageNumber="906" type="lectotype">lectotype</typeStatus>
(body length
<quantity id="A00E84FE0941FFC0FE751B1C751AFEAA" box="[479,557,277,299]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="5" pageNumber="906" unit="mm" value="9.2">9.2 mm</quantity>
) and his median lobe I have embedded in Euparal.
</paragraph>
<paragraph id="6749291B0941FFC0FF1B1B3276C7FD31" blockId="5.[177,1085,315,688]" pageId="5" pageNumber="906">
Some years earlier I have already investigated material from Crete, belonging to the same taxon, and the identity of the median lobe (apical lamella in
<taxonomicName id="A0F652980941FFC0FC801B51730AFEEF" authority="GERMAR 1824" authorityName="GERMAR" authorityYear="1824" box="[810,1085,344,366]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="906" phylum="Arthropoda" rank="species" species="sulphuripes">
<emphasis id="5582F5090941FFC0FC801B517497FEEF" box="[810,928,344,366]" italics="true" pageId="5" pageNumber="906">sulphuripes</emphasis>
GERMAR 1824
</taxonomicName>
only somewhat shorter than in specimens from Crete) and the fine stucture of the internal sac with that one of
<taxonomicName id="A0F652980941FFC0FE211B9A7515FE29" authorityName="GERMAR" authorityYear="1824" box="[395,546,402,424]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="906" phylum="Arthropoda" rank="species" species="sulphuripes">
<emphasis id="5582F5090941FFC0FE211B9A7515FE29" box="[395,546,402,424]" italics="true" pageId="5" pageNumber="906">H. sulphuripes</emphasis>
</taxonomicName>
convinced me that it comes nearest to this species. The investigation of the
<typeStatus id="B84D97B90941FFC0FE1D1BB976DAFE44" box="[439,493,432,453]" pageId="5" pageNumber="906">types</typeStatus>
of
<emphasis id="5582F5090941FFC0FDBC1BB875DCFE47" box="[534,747,432,454]" italics="true" pageId="5" pageNumber="906">
<taxonomicName id="A0F652980941FFC0FDBC1BB875A6FE47" baseAuthorityName="DUFTSCHMIDT" baseAuthorityYear="1812" box="[534,657,432,454]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="906" phylum="Arthropoda" rank="species" species="honestus">H. honestus</taxonomicName>
creticus
</emphasis>
confirmed my supposition, that they belong to
<taxonomicName id="A0F652980941FFC0FEFB1BC776DFFE62" authorityName="GERMAR" authorityYear="1824" box="[337,488,461,483]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="906" phylum="Arthropoda" rank="species" species="sulphuripes">
<emphasis id="5582F5090941FFC0FEFB1BC776DFFE62" box="[337,488,461,483]" italics="true" pageId="5" pageNumber="906">H. sulphuripes</emphasis>
</taxonomicName>
and not to
<taxonomicName id="A0F652980941FFC0FDCF1BC7741EFE62" authority="(DFT.)" baseAuthorityName="DUFTSCHMIDT" baseAuthorityYear="1812" box="[613,809,461,483]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="906" phylum="Arthropoda" rank="species" species="honestus">
<emphasis id="5582F5090941FFC0FDCF1BC775E8FE62" box="[613,735,461,483]" italics="true" pageId="5" pageNumber="906">H. honestus</emphasis>
(DFT.)
</taxonomicName>
. So far the nominotypical form of
<taxonomicName id="A0F652980941FFC0FEAF1BE276D4FD81" authority="GERM." authorityName="GERMAR" authorityYear="1824" box="[261,483,490,512]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="906" phylum="Arthropoda" rank="species" species="sulphuripes">
<emphasis id="5582F5090941FFC0FEAF1BE276ADFD81" box="[261,410,490,512]" italics="true" pageId="5" pageNumber="906">H. sulphuripes</emphasis>
GERM.
</taxonomicName>
is not known from Crete, this fact makes it very likely that the taxon
<taxonomicName id="A0F652980941FFC0FEB3180176FFFD9C" authority="MAR." authorityName="MARAN" authorityYear="1934" box="[281,456,518,541]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="906" phylum="Arthropoda" rank="species" species="creticus">
<emphasis id="5582F5090941FFC0FEB3180176BDFD9C" box="[281,394,520,541]" italics="true" pageId="5" pageNumber="906">H. creticus</emphasis>
MAŘ.
</taxonomicName>
is an isolate, which can, as differing in some important characters, be ranked as one subspecies of the polytypic
<taxonomicName id="A0F652980941FFC0FD7A182F7498FDBB" authority="GERM." authorityName="GERMAR" authorityYear="1824" box="[720,943,549,571]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="906" phylum="Arthropoda" rank="species" species="sulphuripes">
<emphasis id="5582F5090941FFC0FD7A182F7452FDBA" box="[720,869,549,571]" italics="true" pageId="5" pageNumber="906">H. sulphuripes</emphasis>
GERM.
</taxonomicName>
, which forms in the western part of its range a subspecies (in N Africa, S
<collectingCountry id="1FE1698B0941FFC0FCBA184B747BFDD9" box="[784,844,578,600]" name="Spain" pageId="5" pageNumber="906">Spain</collectingCountry>
, S
<collectingCountry id="1FE1698B0941FFC0FCC1184A7485FDD9" box="[875,946,579,600]" name="France" pageId="5" pageNumber="906">France</collectingCountry>
, and Sardinia with ssp.
<emphasis id="5582F5090941FFC0FEB21869753AFDF4" box="[280,525,607,629]" italics="true" pageId="5" pageNumber="906">
<taxonomicName id="A0F652980941FFC0FEB218697687FDF4" authorityName="GERMAR" authorityYear="1824" box="[280,432,607,629]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="906" phylum="Arthropoda" rank="species" species="sulphuripes">H. sulphuripes</taxonomicName>
goudotii
</emphasis>
DEJEAN 1829). The reasons presumably lie also in a brachyptery which limits the spreading power and hence the ability of overcoming geographical considerable barriers.
</paragraph>
<paragraph id="6749291B0941FFC0FF1B18B774EBFCA9" blockId="5.[177,1085,702,808]" pageId="5" pageNumber="906">
Judging from the label: &quot;
<taxonomicName id="A0F652980941FFC0FE1618B77519FD55" authorityName="MARAN" authorityYear="1934" box="[444,558,702,724]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="906" phylum="Arthropoda" rank="species" species="creticus">H. creticus</taxonomicName>
m. 3 / Dr. J. Mařan det. 1934&quot; Mařan probably intended to vote for species status for his
<emphasis id="5582F5090941FFC0FDF418D2759AFD71" box="[606,685,731,752]" italics="true" pageId="5" pageNumber="906">creticus</emphasis>
, though in the description it got a subspecies status. Further investigations may demonstrate that there are two species but for settling of this question more material and genetic research methods are needed.
</paragraph>
<paragraph id="6749291B0941FFC0FF1B193E730AFCCC" blockId="5.[177,1085,823,1164]" box="[177,1085,823,845]" pageId="5" pageNumber="906">
The nominotypical subspecies of
<taxonomicName id="A0F652980941FFC0FDBC193175CAFCCC" authority="GERM." authorityName="GERMAR" authorityYear="1824" box="[534,765,823,845]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="906" phylum="Arthropoda" rank="species" species="sulphuripes">
<emphasis id="5582F5090941FFC0FDBC19317598FCCC" box="[534,687,823,845]" italics="true" pageId="5" pageNumber="906">H. sulphuripes</emphasis>
GERM.
</taxonomicName>
and subspecies
<emphasis id="5582F5090941FFC0FC041931730AFCCC" box="[942,1085,824,845]" italics="true" pageId="5" pageNumber="906">
H. s.
<taxonomicName id="A0F652980941FFC0FC471931730AFCCC" box="[1005,1085,824,845]" pageId="5" pageNumber="906">creticus</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="6749291B0941FFC0FF1B195C76A9FCEB" blockId="5.[177,1085,823,1164]" box="[177,414,851,874]" pageId="5" pageNumber="906">MAŘ. differ as follows:</paragraph>
<paragraph id="6749291B0941FFC0FF1B197C730BFC7D" blockId="5.[177,1085,823,1164]" pageId="5" pageNumber="906">
A Antennae unicolorous light. Penultimate and preceeding abdominal sternites, except the two obligatory setae glabrous, or sometimes with few fine additional setae. Body on average more slender with head small. Apical lamella of median lobe somewhat shorter. Upper surface black, males mostly with weak metallic tinge from blue to green, females black, or with metallic tinge much weaker than in males.
<collectingCountry id="1FE1698B0941FFC0FC0E19D974E4FC64" box="[932,979,976,997]" name="Italy" pageId="5" pageNumber="906">Italy</collectingCountry>
, Balkan Peninsula except Crete,
<collectingCountry id="1FE1698B0941FFC0FE6A19EE7527FC7D" box="[448,528,999,1020]" name="Ukraine" pageId="5" pageNumber="906">Ukraine</collectingCountry>
,
<collectingCountry id="1FE1698B0941FFC0FDB019EE75BCFC7D" box="[538,651,999,1020]" name="Turkey" pageId="5" pageNumber="906">Asia Minor</collectingCountry>
. ............
<taxonomicName id="A0F652980941FFC0FD4A19EE730BFC7D" authority="GERMAR 1824" authorityName="GERMAR" authorityYear="1824" box="[736,1084,999,1020]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="906" phylum="Arthropoda" rank="subSpecies" species="sulphuripes" subSpecies="sulphuripes">
<emphasis id="5582F5090941FFC0FD4A19EE743AFC7D" box="[736,781,999,1020]" italics="true" pageId="5" pageNumber="906">H. s.</emphasis>
ssp.
<emphasis id="5582F5090941FFC0FC9419EE7499FC7D" box="[830,942,999,1020]" italics="true" pageId="5" pageNumber="906">sulphuripes</emphasis>
GERMAR 1824
</taxonomicName>
</paragraph>
<paragraph id="6749291B0941FFC0FF1B1E0F730BFB0D" blockId="5.[177,1085,823,1164]" pageId="5" pageNumber="906">
B Antennae except antennomere 1 strongly darkened, especially middle antennomeres. Penultimate and preceeding abdominal sternites, except the two obligatory setae, seemingly always with numerous long additional setae. Body on average stouter with head larger. Apical lamella of median lobe somewhat longer. Upper surface black, with weak blue-violaceous metallic tinge, females black, or with metallic tinge much weaker than in males.
<collectingRegion id="A532E7F90941FFC0FE1A1E7E76D2FB0D" box="[432,485,1143,1164]" country="Greece" name="Kriti" pageId="5" pageNumber="906">Crete</collectingRegion>
........................ ........................
<taxonomicName id="A0F652980941FFC0FCA61E7E730BFB0D" authority="MARAN 1934" authorityName="MARAN" authorityYear="1934" box="[780,1084,1142,1164]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="906" phylum="Arthropoda" rank="subSpecies" species="honestus" subSpecies="creticus">
<emphasis id="5582F5090941FFC0FCA61E7E740EFB0D" box="[780,825,1143,1164]" italics="true" pageId="5" pageNumber="906">H. s.</emphasis>
ssp.
<emphasis id="5582F5090941FFC0FCC01E7E7481FB0D" box="[874,950,1143,1164]" italics="true" pageId="5" pageNumber="906">creticus</emphasis>
<bibRefCitation id="036754EA0941FFC0FC161E7E730BFB0D" author="MARAN J" box="[956,1084,1142,1164]" pageId="5" pageNumber="906" pagination="125 - 128" refId="ref25534" refString="MARAN J. (1934): Nove druhy a formy Carabidu z ostrova Hrety. De novis carabidarum speciebus formis-que ex insula Creta. - Sbornik Entomologickeho oddeleni Narodniho Musea v Praze 12: 125 - 128." type="journal article" year="1934">MAŘAN 1934</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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