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<document id="8E2C171E674392569895ED60E5DFB7CC" ID-CLB-Dataset="264111" ID-DOI="10.5852/ejt.2023.863.2081" ID-GBIF-Dataset="59ed2548-4c91-4485-ae44-08609e1028bb" ID-ISSN="2118-9773" ID-Zenodo-Dep="7767958" ID-ZooBank="10981377-CCE7-4487-A415-4E409E55A507" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1679676818438" checkinUser="felipe" docAuthor="Gibbs, David" docDate="2023" docId="EE3F8791FF2E4C8DFDE03F90D028E569" docLanguage="en" docName="ejt-863_1-162.pdf" docOrigin="European Journal of Taxonomy 863 (1)" docSource="http://zoobank.org/10981377-cce7-4487-a415-4e409e55a507" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Usia grisea Efflatoun 1945" docType="treatment" docVersion="8" lastPageNumber="151" masterDocId="1206FFE9FFB84C1BFFEA3D18D57BE31F" masterDocTitle="A world review of the bee fly tribe Usiini (Diptera, Bombyliidae) - Part 3: Parageron Paramonov s. lat." masterLastPageNumber="162" masterPageNumber="1" pageNumber="151" updateTime="1699477371262" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="84AC0732595666AE7242DB8A0DD11A92">A world review of the bee fly tribe Usiini (Diptera, Bombyliidae) - Part 3: Parageron Paramonov s. lat.</mods:title>
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<mods:title id="E35753250DCC4FE6F4B5249E79813B3C">European Journal of Taxonomy</mods:title>
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<treatment id="EE3F8791FF2E4C8DFDE03F90D028E569" ID-DOI="http://doi.org/10.5281/zenodo.8291115" ID-GBIF-Taxon="212472407" ID-Zenodo-Dep="8291115" LSID="urn:lsid:plazi:treatment:EE3F8791FF2E4C8DFDE03F90D028E569" httpUri="http://treatment.plazi.org/id/EE3F8791FF2E4C8DFDE03F90D028E569" lastPageNumber="151" pageId="150" pageNumber="151">
<subSubSection id="2E8C650CFF2E4C8DFDE03F90D151E1BC" box="[522,1066,648,675]" pageId="150" pageNumber="151" type="nomenclature">
<paragraph id="66293687FF2E4C8DFDE03F90D151E1BC" blockId="150.[522,1066,648,710]" box="[522,1066,648,675]" pageId="150" pageNumber="151">
<heading id="3D6181EBFF2E4C8DFDE03F90D151E1BC" box="[522,1066,648,675]" centered="true" fontSize="11" level="2" pageId="150" pageNumber="151" reason="2">
<taxonomicName id="A1964D04FF2E4C8DFDE03F90D629E1BC" ID-CoL="8FL6B" authority="Efflatoun, 1945" authorityName="Efflatoun" authorityYear="1945" box="[522,850,648,675]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="150" pageNumber="151" phylum="Arthropoda" rank="species" species="grisea">
<emphasis id="54E2EA95FF2E4C8DFDE03F90D7E9E1BD" bold="true" box="[522,658,648,674]" italics="true" pageId="150" pageNumber="151">Usia grisea</emphasis>
<bibRefCitation id="02074B76FF2E4C8DFD733F91D629E1BC" author="Efflatoun H. C." box="[665,850,648,675]" pageId="150" pageNumber="151" pagination="1 - 483" refId="ref74131" refString="Efflatoun H. C. 1945. A monograph of Egyptian Diptera. Part VI. Family Bombyliidae. Section 1: Subfamily Bombyliidae Homoeophthalmae. Bulletin de la Societe Fouad Ier d'Entomologie 29: 1 - 483." type="journal article" year="1945">Efflatoun, 1945</bibRefCitation>
</taxonomicName>
, unplaced
<taxonomicName id="A1964D04FF2E4C8DFC3A3F91D151E1BC" authorityName="Becker" authorityYear="1913" box="[976,1066,649,675]" class="Insecta" family="Bombyliidae" kingdom="Animalia" order="Diptera" pageId="150" pageNumber="158" phylum="Arthropoda" rank="subFamily" subFamily="Usiinae">Usiinae</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2E8C650CFF2E4C8DFD1A3FB4D638E1D9" box="[752,835,684,710]" pageId="150" pageNumber="151" type="description">
<paragraph id="66293687FF2E4C8DFD1A3FB4D638E1D9" blockId="150.[522,1066,648,710]" box="[752,835,684,710]" pageId="150" pageNumber="151">
<figureCitation id="FEAD2A02FF2E4C8DFD1A3FB4D638E1D9" box="[752,835,684,710]" captionStart="Fig" captionStartId="151.[189,232,1998,2024]" captionTargetBox="[189,1398,563,1960]" captionTargetId="figure-54@151.[189,1398,560,1963]" captionTargetPageId="151" captionText="Fig. 68. Usia grisea Efflatoun, 1945, habitus, Smithsonian Institution Archives. Image SIA2012-7883." figureDoi="http://doi.org/10.5281/zenodo.7768083" httpUri="https://zenodo.org/record/7768083/files/figure.png" pageId="150" pageNumber="151">Fig. 68</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2E8C650CFF2E4C8DFF573FF6D73DE018" box="[189,582,749,776]" pageId="150" pageNumber="151" type="reference_group">
<paragraph id="66293687FF2E4C8DFF573FF6D73DE018" blockId="150.[189,582,749,776]" box="[189,582,749,776]" pageId="150" pageNumber="151">
<treatmentCitationGroup id="468611A9FF2E4C8DFF573FF6D73DE018" box="[189,582,749,776]" pageId="150" pageNumber="151">
<treatmentCitation id="E7371096FF2E4C8DFF573FF6D739E018" author="Efflatoun H. C." box="[189,578,749,776]" page="214" pageId="150" pageNumber="151" year="1945">
<taxonomicName id="A1964D04FF2E4C8DFF573FF6D739E018" ID-CoL="8FL6B" authority="Efflatoun, 1945: 214" authorityName="Efflatoun" authorityPageNumber="214" authorityYear="1945" box="[189,578,749,776]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="150" pageNumber="151" phylum="Arthropoda" rank="species" species="grisea">
<emphasis id="54E2EA95FF2E4C8DFF573FF6D43EE018" box="[189,325,750,775]" italics="true" pageId="150" pageNumber="151">Usia grisea</emphasis>
<bibRefCitation id="02074B76FF2E4C8DFEA63FF5D739E018" author="Efflatoun H. C." box="[332,578,749,776]" pageId="150" pageNumber="151" pagination="1 - 483" refId="ref74131" refString="Efflatoun H. C. 1945. A monograph of Egyptian Diptera. Part VI. Family Bombyliidae. Section 1: Subfamily Bombyliidae Homoeophthalmae. Bulletin de la Societe Fouad Ier d'Entomologie 29: 1 - 483." type="journal article" year="1945">Efflatoun, 1945: 214</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="2E8C650CFF2E4C8DFF573E36D43AE700" pageId="150" pageNumber="151" type="materials_examined">
<paragraph id="66293687FF2E4C8DFF573E36D417E057" blockId="150.[189,364,814,840]" box="[189,364,814,840]" pageId="150" pageNumber="151">
<heading id="3D6181EBFF2E4C8DFF573E36D417E057" bold="true" box="[189,364,814,840]" fontSize="11" level="3" pageId="150" pageNumber="151" reason="3">
<emphasis id="54E2EA95FF2E4C8DFF573E36D417E057" bold="true" box="[189,364,814,840]" pageId="150" pageNumber="151">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="66293687FF2E4C8DFF003E44D76DE068" blockId="150.[189,1398,860,956]" box="[234,534,860,887]" pageId="150" pageNumber="151">
<emphasis id="54E2EA95FF2E4C8DFF003E44D427E069" bold="true" box="[234,348,860,886]" pageId="150" pageNumber="151">
<typeStatus id="B92D8825FF2E4C8DFF003E44D427E069" box="[234,348,860,886]" pageId="150" pageNumber="151" type="holotype">Holotype</typeStatus>
</emphasis>
(not examined)
</paragraph>
<paragraph id="66293687FF2E4C8DFF573E67D497E0A3" blockId="150.[189,1398,860,956]" pageId="150" pageNumber="151">
<materialsCitation id="D6FE3CDAFF2E4C8DFF573E67D497E0A3" ID-GBIF-Occurrence="4401245423" collectingDate="1938-02-28" collectionCode="ESEC" collectorName="Mohamed Tewfik Effendi" country="Egypt" location="Wadi Ehameib" municipality="West Elba Mountain" pageId="150" pageNumber="151" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<collectingCountry id="1E817617FF2E4C8DFF573E67D460E086" box="[189,283,895,921]" name="Egypt" pageId="150" pageNumber="151">EGYPT</collectingCountry>
•
<specimenCount id="7090FD0EFF2E4C8DFEDF3E67D435E086" box="[309,334,895,921]" pageId="150" pageNumber="151" type="male">♂</specimenCount>
; “
<location id="6349605CFF2E4C8DFE8F3E67D76DE086" LSID="urn:lsid:plazi:treatment:EE3F8791FF2E4C8DFDE03F90D028E569:6349605CFF2E4C8DFE8F3E67D76DE086" box="[357,534,895,921]" country="Egypt" municipality="West Elba Mountain" name="Wadi Ehameib" pageId="150" pageNumber="151">Wadi Ehameib</location>
(
<collectingMunicipality id="864DACFDFF2E4C8DFDCC3E67D65BE085" box="[550,800,895,922]" pageId="150" pageNumber="151">West Elba Mountain</collectingMunicipality>
).
<date id="12281047FF2E4C8DFCD93E67D17CE086" box="[819,1031,895,921]" pageId="150" pageNumber="151" value="1938-02-28">
<collectingDate id="026CE9AFFF2E4C8DFCD93E67D17CE086" box="[819,1031,895,921]" pageId="150" pageNumber="151" value="1938-02-28">28 February 1938</collectingDate>
</date>
, leg.
<collectorName id="CB635351FF2E4C8DFBAD3E67D00DE086" box="[1095,1398,895,921]" pageId="150" pageNumber="151">Mohamed Tewfik Effendi</collectorName>
(
<bibRefCitation id="02074B76FF2E4C8DFF2F3EB9D400E0A3" author="Efflatoun H. C." box="[197,379,929,956]" pageId="150" pageNumber="151" pagination="1 - 483" refId="ref74131" refString="Efflatoun H. C. 1945. A monograph of Egyptian Diptera. Part VI. Family Bombyliidae. Section 1: Subfamily Bombyliidae Homoeophthalmae. Bulletin de la Societe Fouad Ier d'Entomologie 29: 1 - 483." type="journal article" year="1945">Efflatoun 1945</bibRefCitation>
)”;
<collectionCode id="0087AE42FF2E4C8DFE763EBAD493E0A3" box="[412,488,930,956]" country="Egypt" httpUri="http://grbio.org/cool/4cts-0w01" name="Entomological Society of Egypt" pageId="150" pageNumber="151">ESEC</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="66293687FF2E4C8DFF573EFAD43AE700" blockId="150.[189,1399,994,1055]" pageId="150" pageNumber="151">
The
<typeStatus id="B92D8825FF2E4C8DFF193EFAD421E0E3" box="[243,346,994,1020]" pageId="150" pageNumber="151" type="holotype">holotype</typeStatus>
male is in
<collectionCode id="0087AE42FF2E4C8DFE343EFBD752E0E2" box="[478,553,995,1021]" country="Egypt" httpUri="http://grbio.org/cool/4cts-0w01" name="Entomological Society of Egypt" pageId="150" pageNumber="151">ESEC</collectionCode>
, photographs suggest that it still exists but are not sufficient to read the data labels.
</paragraph>
</subSubSection>
<subSubSection id="2E8C650CFF2E4C8DFF57395ED6A0E518" pageId="150" pageNumber="151" type="discussion">
<paragraph id="66293687FF2E4C8DFF57395ED454E77F" blockId="150.[189,303,1094,1120]" box="[189,303,1094,1120]" pageId="150" pageNumber="151">
<heading id="3D6181EBFF2E4C8DFF57395ED454E77F" bold="true" box="[189,303,1094,1120]" fontSize="11" level="3" pageId="150" pageNumber="151" reason="3">
<emphasis id="54E2EA95FF2E4C8DFF57395ED454E77F" bold="true" box="[189,303,1094,1120]" pageId="150" pageNumber="151">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="66293687FF2E4C8DFF57396CD1D9E641" blockId="150.[189,1399,1140,1375]" pageId="150" pageNumber="151">
With the
<typeStatus id="B92D8825FF2E4C8DFECB396CD4FCE791" box="[289,391,1140,1166]" pageId="150" pageNumber="151" type="holotype">holotype</typeStatus>
being inaccessible, and no non-type material running to this species nor fitting
<bibRefCitation id="02074B76FF2E4C8DFB1D396CD471E7AE" author="Efflatoun H. C." pageId="150" pageNumber="151" pagination="1 - 483" refId="ref74131" refString="Efflatoun H. C. 1945. A monograph of Egyptian Diptera. Part VI. Family Bombyliidae. Section 1: Subfamily Bombyliidae Homoeophthalmae. Bulletin de la Societe Fouad Ier d'Entomologie 29: 1 - 483." type="journal article" year="1945">Efflatoun’s (1945)</bibRefCitation>
description, the placement of this taxon is difficult. The plate in the Smithsonian archive [Image no. SIA2012-7883] (
<figureCitation id="FEAD2A02FF2E4C8DFE4539A1D778E7CB" box="[431,515,1209,1236]" captionStart="Fig" captionStartId="151.[189,232,1998,2024]" captionTargetBox="[189,1398,563,1960]" captionTargetId="figure-54@151.[189,1398,560,1963]" captionTargetPageId="151" captionText="Fig. 68. Usia grisea Efflatoun, 1945, habitus, Smithsonian Institution Archives. Image SIA2012-7883." figureDoi="http://doi.org/10.5281/zenodo.7768083" httpUri="https://zenodo.org/record/7768083/files/figure.png" pageId="150" pageNumber="151">Fig. 68</figureCitation>
) seems to show a typical
<taxonomicName id="A1964D04FF2E4C8DFCC539A2D6D7E7CC" authorityName="Paramonov" authorityYear="1929" box="[815,940,1210,1235]" class="Insecta" family="Bombyliidae" genus="Parageron" kingdom="Animalia" order="Diptera" pageId="150" pageNumber="151" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="54E2EA95FF2E4C8DFCC539A2D6D7E7CC" box="[815,940,1210,1235]" italics="true" pageId="150" pageNumber="151">Parageron</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFF2E4C8DFC5939A2D68AE7CC" box="[947,1009,1209,1236]" pageId="150" pageNumber="151" sensu="lato">s. lat.</taxonomicNameLabel>
(
<taxonomicName id="A1964D04FF2E4C8DFBE839A2D1FBE7CC" authority="Gibbs, 2023" authorityName="Gibbs" authorityYear="2023" box="[1026,1152,1210,1235]" class="Insecta" family="Bombyliidae" genus="Protypusia" kingdom="Animalia" order="Diptera" pageId="150" pageNumber="151" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="54E2EA95FF2E4C8DFBE839A2D1FBE7CC" box="[1026,1152,1210,1235]" italics="true" pageId="150" pageNumber="151">Protypusia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFF2E4C8DFB6D39A2D18EE7CB" box="[1159,1269,1210,1236]" pageId="150" pageNumber="151" rank="genus">gen. nov.</taxonomicNameLabel>
), relatively robust, short postpedicel, long hair across mesonotum, short discal cell, A
<subScript id="FA1234C2FF2E4C8DFBCC39F7D155E7E2" attach="left" box="[1062,1070,1263,1277]" fontSize="6" pageId="150" pageNumber="151">1</subScript>
+CuA
<subScript id="FA1234C2FF2E4C8DFB9F39F7D105E7E2" attach="left" box="[1141,1150,1263,1277]" fontSize="6" pageId="150" pageNumber="151">2</subScript>
(petiole of anal cell) short and A
<subScript id="FA1234C2FF2E4C8DFEAE380AD436E63F" attach="left" box="[324,333,1298,1312]" fontSize="6" pageId="150" pageNumber="151">1</subScript>
angled where it meets CuA
<subScript id="FA1234C2FF2E4C8DFD7B380AD7E1E63F" attach="left" box="[657,666,1298,1312]" fontSize="6" pageId="150" pageNumber="151">2</subScript>
just before wing margin (left wing only). On the other hand, it is a tiny species and Efflatoun keyed it close to
<taxonomicName id="A1964D04FF2E4C8DFD06383AD636E624" authorityName="Effl., Det. Efflatoun" authorityYear="1945" box="[748,845,1313,1340]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="150" pageNumber="151" phylum="Arthropoda" rank="species" species="elbae">
<emphasis id="54E2EA95FF2E4C8DFD06383AD636E624" box="[748,845,1313,1340]" italics="true" pageId="150" pageNumber="151">A. elbae</emphasis>
</taxonomicName>
, compares it to
<taxonomicName id="A1964D04FF2E4C8DFBE1383AD110E624" authorityName="Effl., Det. Efflatoun" authorityYear="1945" box="[1035,1131,1313,1340]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="150" pageNumber="151" phylum="Arthropoda" rank="species" species="elbae">
<emphasis id="54E2EA95FF2E4C8DFBE1383AD110E624" box="[1035,1131,1313,1340]" italics="true" pageId="150" pageNumber="151">A. elbae</emphasis>
</taxonomicName>
in the description, and appears to illustrate the
<taxonomicName id="A1964D04FF2E4C8DFE3B385CD748E641" authorityName="Loew" authorityYear="1860" box="[465,563,1348,1374]" class="Insecta" family="Bombyliidae" genus="Apolysis" kingdom="Animalia" order="Diptera" pageId="150" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="54E2EA95FF2E4C8DFE3B385CD748E641" box="[465,563,1348,1374]" italics="true" pageId="150" pageNumber="151">Apolysis</emphasis>
</taxonomicName>
articulating arista in
<bibRefCitation id="02074B76FF2E4C8DFCC3385CD69FE640" author="Efflatoun H. C." box="[809,996,1348,1375]" pageId="150" pageNumber="151" pagination="1 - 483" refId="ref74131" refString="Efflatoun H. C. 1945. A monograph of Egyptian Diptera. Part VI. Family Bombyliidae. Section 1: Subfamily Bombyliidae Homoeophthalmae. Bulletin de la Societe Fouad Ier d'Entomologie 29: 1 - 483." type="journal article" year="1945">Efflatoun (1945</bibRefCitation>
: pl. 20 fig. 319).
</paragraph>
<paragraph id="66293687FF2E4C8DFF57389DD6A0E518" blockId="150.[189,1399,1413,1543]" pageId="150" pageNumber="151">
Given the contradictory evidence, and the fact that the
<typeStatus id="B92D8825FF2E4C8DFCBE389DD6C0E680" box="[852,955,1413,1439]" pageId="150" pageNumber="151" type="holotype">holotype</typeStatus>
persists, assigning it to a genus here cannot be justified. The possibility that it is conspecific with one of the species of
<taxonomicName id="A1964D04FF2E4C8DFB6E38B0D07FE6DE" authority="Gibbs, 2023" authorityName="Gibbs" authorityYear="2023" box="[1156,1284,1448,1473]" class="Insecta" family="Bombyliidae" genus="Protypusia" kingdom="Animalia" order="Diptera" pageId="150" pageNumber="151" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="54E2EA95FF2E4C8DFB6E38B0D07FE6DE" box="[1156,1284,1448,1473]" italics="true" pageId="150" pageNumber="151">Protypusia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4FD157EEFF2E4C8DFAE638B0D00CE6DD" box="[1292,1399,1448,1474]" pageId="150" pageNumber="151" rank="genus">gen. nov.</taxonomicNameLabel>
treated above has been researched, but none fit the description of
<taxonomicName id="A1964D04FF2E4C8DFC3938D3D139E6FB" authorityName="Efflatoun" authorityYear="1945" box="[979,1090,1483,1508]" class="Insecta" family="Bombyliidae" genus="Usia" kingdom="Animalia" order="Diptera" pageId="150" pageNumber="151" phylum="Arthropoda" rank="species" species="grisea">
<emphasis id="54E2EA95FF2E4C8DFC3938D3D139E6FB" box="[979,1090,1483,1508]" italics="true" pageId="150" pageNumber="151">U. grisea</emphasis>
</taxonomicName>
very closely. However, it could well belong in
<taxonomicName id="A1964D04FF2E4C8DFE5F38F6D74EE518" authorityName="Gibbs" authorityYear="2023" box="[437,565,1518,1543]" class="Insecta" family="Bombyliidae" genus="Protypusia" kingdom="Animalia" order="Diptera" pageId="150" pageNumber="151" phylum="Arthropoda" rank="genus">
<emphasis id="54E2EA95FF2E4C8DFE5F38F6D74EE518" box="[437,565,1518,1543]" italics="true" pageId="150" pageNumber="151">Protypusia</emphasis>
</taxonomicName>
and this is accounted for in the key.
</paragraph>
</subSubSection>
<subSubSection id="2E8C650CFF2E4C8DFF573B35D028E569" box="[189,1363,1581,1654]" pageId="150" pageNumber="151" type="distribution">
<paragraph id="66293687FF2E4C8DFF573B35D42DE558" blockId="150.[189,342,1581,1607]" box="[189,342,1581,1607]" pageId="150" pageNumber="151">
<heading id="3D6181EBFF2E4C8DFF573B35D42DE558" bold="true" box="[189,342,1581,1607]" fontSize="11" level="3" pageId="150" pageNumber="151" reason="3">
<emphasis id="54E2EA95FF2E4C8DFF573B35D42DE558" bold="true" box="[189,342,1581,1607]" pageId="150" pageNumber="151">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="66293687FF2E4C8DFF573B44D028E569" blockId="150.[189,1363,1628,1654]" box="[189,1363,1628,1654]" pageId="150" pageNumber="151">The single male was collected in Wadi Ehameib, West Elba Mountain, close to the Sudanese border.</paragraph>
</subSubSection>
</treatment>
</document>
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