132 lines
13 KiB
XML
132 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.271.4062" ID-GBIF-Dataset="c93c3a93-d2e1-4c85-8ece-bf3aa1a5109d" ID-PMC="PMC3652427" ID-Pensoft-Pub="1313-2970-271-1" ID-PubMed="23717185" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-271-1" ModsDocOrigin="ZooKeys 271" ModsDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" checkinTime="1451247655320" checkinUser="pensoft" docAuthor="Caterino, Michael S. & Tishechkin, Alexey K." docDate="2013" docId="EDFE6ED51B98EEAEAE71FEE8CA1B944E" docLanguage="en" docName="ZooKeys 271: 1-401" docOrigin="ZooKeys 271" docSource="http://dx.doi.org/10.3897/zookeys.271.4062" docTitle="Operclipygus bosquesecus Caterino & Tishechkin, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="327" masterDocId="4403FF80FFE7FF95FFDA5D0E29259B2F" masterDocTitle="A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)" masterLastPageNumber="401" masterPageNumber="1" pageNumber="326" updateTime="1668155382655" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Caterino, Michael S.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tishechkin, Alexey K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>271</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>401</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.271.4062</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.271.4062</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-271-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152040851" LSID="urn:lsid:zoobank.org:act:30E6D513-4F57-4784-8B40-F231C17220A9" httpUri="http://treatment.plazi.org/id/EDFE6ED51B98EEAEAE71FEE8CA1B944E" lastPageId="326" lastPageNumber="327" pageId="325" pageNumber="326">
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<subSubSection pageId="325" pageNumber="326" type="nomenclature">
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<paragraph pageId="325" pageNumber="326">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:30E6D513-4F57-4784-8B40-F231C17220A9" class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus bosquesecus" order="Coleoptera" pageId="325" pageNumber="326" phylum="Arthropoda" rank="species" species="bosquesecus">Operclipygus bosquesecus</taxonomicName>
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<taxonomicNameLabel pageId="325" pageNumber="326">sp. n.</taxonomicNameLabel>
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Figs 88
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<normalizedToken originalValue="C–E90A–B">C-E90A-B</normalizedToken>
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, E, GMap 31
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</paragraph>
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</subSubSection>
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<subSubSection pageId="325" pageNumber="326" type="type locality">
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<paragraph pageId="325" pageNumber="326">Type locality.</paragraph>
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<paragraph pageId="325" pageNumber="326">
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ECUADOR:
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<normalizedToken originalValue="Manabí">Manabi</normalizedToken>
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: Lalo Loor Reserve [
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<geoCoordinate direction="south" orientation="latitude" precision="5" value="-0.0832">0.0832°S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-80.152">80.1520°W</geoCoordinate>
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].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="325" pageNumber="326" type="type material">
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<paragraph pageId="325" pageNumber="326">Type material.</paragraph>
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<paragraph pageId="325" pageNumber="326">
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Holotype male: "ECUADOR:
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<normalizedToken originalValue="Manabí">Manabi</normalizedToken>
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<geoCoordinate direction="south" orientation="latitude" precision="5" value="-0.0832">0.0832°S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-80.152">80.1520°W</geoCoordinate>
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Lalo Loor Bosque Seco Res., 21-27.v.2011, FIT, AT1304, M. Caterino, A. Tishechkin" / "Caterino/Tishechkin
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<taxonomicName lsidName="" pageId="325" pageNumber="326" rank="tribe" tribe="Exosternini">Exosternini</taxonomicName>
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Voucher EXO-00599" (FMNH). Paratypes (9): ECUADOR: Los
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<normalizedToken originalValue="Ríos">Rios</normalizedToken>
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: 1: CCRP, 8.vi.1980, palma abierta, S. Sandoval (CHSM);
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<normalizedToken originalValue="Manabí">Manabi</normalizedToken>
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: 6: Bosque Seco Lalo Loor,
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<geoCoordinate direction="south" orientation="latitude" precision="5" value="-0.0832">0.0832°S</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="5" value="-80.152">80.1520°W</geoCoordinate>
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, 21-27.v.2011, FIT, M.S. Caterino & A.K. Tishechkin, DNA Extract MSC-2169 (SBMNH, MSCC, AKTC, USFQ); Pichincha: 2: Rio Palenque Sta., 47km S Santo Domingo, 250m, 25.ii.1979, pan trap, S.A. Marshall (CHSM).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="326" lastPageNumber="327" pageId="325" pageNumber="326" type="description">
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<paragraph pageId="325" pageNumber="326">Diagnostic description.</paragraph>
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<paragraph lastPageId="326" lastPageNumber="327" pageId="325" pageNumber="326">
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Length: 1.81-2.22 mm, width: 1.59-1.93 mm; body piceus, broadly rounded, widest near middle of elytra; frons flat, more or less parallel-sided, with sides of frontal stria weakly rounded; central portion of frontal stria outwardly arcuate, complete or narrowly detached from sides; supraorbital stria present, detached from frontal stria; epistoma convex, labrum short, very weakly asymmetrical, with left side more strongly produced than right; left mandible untoothed, right with small, acute basal tooth; pronotal disk with small, irregular but distinct prescutellar impression, with inconspicuous ground punctation and ~10 coarser, elongate punctures toward sides; marginal stria interrupted for width of head; lateral submar
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<pageBreakToken pageId="326" pageNumber="327" start="start">ginal</pageBreakToken>
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stria complete along side, curved inward at front nearly to anterior submarginal stria, the ends of which are perpendicularly recurved, median pronotal gland openings visible laterad its ends, about 8 puncture widths from anterior margin; elytron with three complete epipleural striae, outer subhumeral stria present in apical half only, mostly below a humeral swelling, inner subhumeral stria weakly impressed, present in most of apical half, striae 1-3 complete, 4th stria present in apical two-thirds, 5th stria present in apical half, sutural stria present in apical two-thirds, elytra depressed along suture; prosternal keel truncate at base, carinal striae complete, well separated, converging only slightly to front, connected basally and apically; prosternal lobe with complete marginal stria; mesoventrite with anterior margin straight, marginal stria complete; mesometaventral stria arched forward to near marginal stria, continued by lateral metaventral stria toward outer third of metacoxa; postmesocoxal stria recurved to mesepimeron; 1st abdominal ventrite with single complete (inner) lateral stria and fragments of outer lateral stria, with series of small punctures along posterior margin; propygidium with fine sparse ground punctation, larger shallow punctures uniformly interspersed, separated by about one-half their diameters; pygidium with denser ground punctation and small coarse punctures irregularly but sparsely interspersed; marginal pygidial stria complete, fine, slightly crenulate. Male genitalia (Figs 90
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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, E, G): accessory sclerites present, small; T8 elongate, sides narrowing gradually to apex, basal emargination broadly rounded, basal membrane attachment line distad by about one-half basal emargination depth from its apex, apical emargination narrow; S8 with apical guides moderately well developed, largely constant in width from base to narrowly rounded apices; T9 with apices acute, convergent, opposing; T10 with halves large, separate; S9 narrowest at middle of stem, gradually widened to subtruncate, desclerotized base, with apical emargination distinct, apical flanges small and separate, lateral flanges parallel, shallow; tegmen widest beyond midpoint, with sides rounded to apex, narrowed slightly to base, with narrow,
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<normalizedToken originalValue="‘U’">'U'</normalizedToken>
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-shaped medioventral process projecting beneath about one-third from base, apical third of tegmen moderately curved ventrad; basal piece short, about one-fourth tegmen length; median lobe nearly one-half tegmen length, with gonopore wide, basal apodemes separate throughout their lengths.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="326" pageNumber="327" type="remarks">
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<paragraph pageId="326" pageNumber="327">Remarks.</paragraph>
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<paragraph pageId="326" pageNumber="327">
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This species is best recognized by its small size, the presence of a prescutellar impression, the weak vestige of an inner subhumeral stria, and the strongly arched mesometaventral stria (Fig. 88E). There are also very few other
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus" order="Coleoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="genus">Operclipygus</taxonomicName>
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on the dry western slopes of the Andes where this species occurs.
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</paragraph>
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<caption pageId="326" pageNumber="327">
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<paragraph pageId="326" pageNumber="327">
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Figure 90. Male genitalia of various
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus" order="Coleoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="genus">Operclipygus</taxonomicName>
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species. A T8 of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus bosquesecus" order="Coleoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="bosquesecus">Operclipygus bosquesecus</taxonomicName>
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B S8 of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus bosquesecus" order="Coleoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="bosquesecus">Operclipygus bosquesecus</taxonomicName>
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C T8 of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus arnaudi" order="Coleoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="arnaudi">Operclipygus arnaudi</taxonomicName>
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D S8 of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus arnaudi" order="Coleoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="arnaudi">Operclipygus arnaudi</taxonomicName>
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E S9 of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus bosquesecus" order="Coleoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="bosquesecus">Operclipygus bosquesecus</taxonomicName>
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F S9 of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus arnaudi" order="Coleoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="arnaudi">Operclipygus arnaudi</taxonomicName>
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G Aedeagus, dorsal and lateral views, of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus bosquesecus" order="Coleoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="bosquesecus">Operclipygus bosquesecus</taxonomicName>
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H Aedeagus, dorsal and lateral views, of
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<taxonomicName class="Insecta" family="Histeridae" genus="Operclipygus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Operclipygus arnaudi" order="Coleoptera" pageId="326" pageNumber="327" phylum="Arthropoda" rank="species" species="arnaudi">Operclipygus arnaudi</taxonomicName>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="326" pageNumber="327" type="etymology">
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<paragraph pageId="326" pageNumber="327">Etymology.</paragraph>
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<paragraph pageId="326" pageNumber="327">This species is named for the seasonally dry forest environment (in Spanish, bosque seco) where it is found.</paragraph>
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</subSubSection>
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</treatment>
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</document> |