221 lines
21 KiB
XML
221 lines
21 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.4762202" ID-GBIF-Dataset="df79feb7-c7e6-4267-b324-91597bc182fd" ID-GBIF-Taxon="182070961" ID-ISSN="1854-0392" ID-Zenodo-Dep="4762202" checkinTime="1620963448601" checkinUser="felipe" docAuthor="Sivec, Ignac & Stark, Bill P." docDate="2011" docId="ED2FE866E4617D60FC66E5E72C718AB5" docLanguage="en" docName="Illiesia.7.24.264-279.pdf" docOrigin="Illiesia 7 (24)" docStyle="DocumentStyle:3ED4ECBC8873166DC296BE619562C54F.3:Illiesia.2009-.journal_article" docStyleId="3ED4ECBC8873166DC296BE619562C54F" docStyleName="Illiesia.2009-.journal_article" docStyleVersion="3" docTitle="Phanoperla circumspina Sivec & Stark 2011, sp. n." docType="treatment" docVersion="6" lastPageId="13" lastPageNumber="275" masterDocId="1116901EE46B7D6DFFDAE1242D7E8B32" masterDocTitle="New Species Of Neoperla Needham And Phanoperla Banks (Plecoptera: Perlidae) From The Philippine Islands" masterLastPageNumber="279" masterPageNumber="264" pageId="10" pageNumber="274" updateTime="1632023202857" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New Species Of Neoperla Needham And Phanoperla Banks (Plecoptera: Perlidae) From The Philippine Islands</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sivec, Ignac</mods:namePart>
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<mods:affiliation>Slovenian Museum of Natural History, Prešernova 20, P. O. Box 290, SLO- 1001 Ljubljana, Slovenia E-mail: isivec @ pms-lj. si</mods:affiliation>
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<mods:nameIdentifier type="email">isivec@pms-lj.si</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Stark, Bill P.</mods:namePart>
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<mods:affiliation>Box 4045, Department of Biology, Mississippi College, Clinton, Mississippi, U. S. A. 39058 E-mail: stark @ mc. edu</mods:affiliation>
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<mods:nameIdentifier type="email">stark@mc.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Illiesia</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>7</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>24</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>264</mods:start>
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<mods:end>279</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.4760560</mods:identifier>
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<mods:identifier type="GBIF-Dataset">df79feb7-c7e6-4267-b324-91597bc182fd</mods:identifier>
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<mods:identifier type="ISSN">1854-0392</mods:identifier>
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<mods:identifier type="Zenodo-Dep">4760560</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.4762202" ID-GBIF-Taxon="182070961" ID-Zenodo-Dep="4762202" LSID="urn:lsid:plazi:treatment:ED2FE866E4617D60FC66E5E72C718AB5" httpUri="http://treatment.plazi.org/id/ED2FE866E4617D60FC66E5E72C718AB5" lastPageId="13" lastPageNumber="275" pageId="10" pageNumber="274">
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<subSubSection box="[956,1322,1219,1244]" pageId="10" pageNumber="274" type="nomenclature">
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<paragraph blockId="10.[956,1322,1219,1244]" box="[956,1322,1219,1244]" pageId="10" pageNumber="274">
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<heading bold="true" box="[956,1322,1219,1244]" centered="true" fontSize="10" level="2" pageId="10" pageNumber="274" reason="2">
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<emphasis bold="true" box="[956,1322,1219,1244]" pageId="10" pageNumber="274">
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<taxonomicName authority="Sivec & Stark, 2011" authorityName="Sivec & Stark" authorityYear="2011" box="[956,1244,1219,1243]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="10" pageNumber="274" phylum="Arthropoda" rank="species" species="circumspina" status="sp. n.">
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<emphasis bold="true" box="[956,1244,1219,1243]" italics="true" pageId="10" pageNumber="274">Phanoperla circumspina</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1258,1322,1220,1244]" pageId="10" pageNumber="274" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[1068,1211,1255,1279]" pageId="10" pageNumber="274" type="description">
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<paragraph blockId="10.[1068,1211,1255,1279]" box="[1068,1211,1255,1279]" pageId="10" pageNumber="274">
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(
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<figureCitation box="[1076,1203,1255,1279]" captionStart-0="Figs" captionStart-1="Figs" captionStartId-0="12.[189,241,829,853]" captionStartId-1="12.[189,241,1894,1918]" captionTargetBox-0="[209,1426,197,826]" captionTargetBox-1="[249,1383,925,1847]" captionTargetId-0="figure-112@12.[631,1001,204,742]" captionTargetId-1="figure-137@12.[249,1383,925,1847]" captionTargetPageId-0="12" captionTargetPageId-1="12" captionText-0="Figs. 45-50. Phanoperla circumspina. 45. Head and pronotum. 46. Male terminalia. 47-50. Penis tube and everted sac. 47. Ventral aspect. 48. Dorsal aspect. 49. Lateral aspect. 50. Apical aspect." captionText-1="Figs. 51-54. Phanoperla circumspina egg. 51. Entire egg. 52. Collar end. 53. Chorionic detail. 54. Anterior end." figureDoi-0="http://doi.org/10.5281/zenodo.4760587" figureDoi-1="http://doi.org/10.5281/zenodo.4760589" httpUri-0="https://zenodo.org/record/4760587/files/figure.png" httpUri-1="https://zenodo.org/record/4760589/files/figure.png" pageId="10" pageNumber="274">Figs. 45-54</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="274" type="materials_examined">
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<paragraph blockId="10.[835,1444,1326,1920]" pageId="10" pageNumber="274">
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<emphasis bold="true" box="[835,1084,1326,1350]" pageId="10" pageNumber="274">Material examined.</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3111458303" collectingDate="1996-03-19" collectingDateMax="1996-03-19" collectingDateMin="1996-02" collectionCode="PZ" country="Philippines" elevation="300" location="Salakot Falls Road" municipality="Salakot Falls Road" pageId="10" pageNumber="274" specimenCount="16" specimenCount-female="1" specimenCount-male="15" stateProvince="Palawan" typeStatus="holotype, paratype">
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<typeStatus box="[1103,1215,1326,1350]" pageId="10" pageNumber="274">Holotype</typeStatus>
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<specimenCount box="[1234,1254,1326,1350]" pageId="10" pageNumber="274" type="male">♂</specimenCount>
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and
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<specimenCount box="[1337,1384,1326,1350]" pageId="10" pageNumber="274" type="male">14♂</specimenCount>
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,
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<specimenCount box="[1410,1443,1326,1350]" pageId="10" pageNumber="274" type="female">1♀</specimenCount>
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<typeStatus box="[835,950,1363,1387]" pageId="10" pageNumber="274">paratypes</typeStatus>
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,
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<emphasis bold="true" box="[960,1239,1362,1386]" pageId="10" pageNumber="274">
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<collectingCountry box="[960,1239,1362,1386]" name="Philippines" pageId="10" pageNumber="274">PHILIPPINE ISLANDS</collectingCountry>
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</emphasis>
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,
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<collectingRegion box="[1250,1351,1362,1386]" country="Philippines" name="Palawan" pageId="10" pageNumber="274">Palawan</collectingRegion>
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,
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<collectingMunicipality pageId="10" pageNumber="274">Salakot Falls Road</collectingMunicipality>
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,
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<quantity box="[974,1048,1398,1422]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="274" unit="m" value="300.0">
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<elevation box="[974,1048,1398,1422]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="274" unit="m" value="300.0">300 m</elevation>
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</quantity>
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,
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<date box="[1061,1235,1398,1422]" pageId="10" pageNumber="274" value="1996-03-19">
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<collectingDate box="[1061,1235,1398,1422]" pageId="10" pageNumber="274" value="1996-03-19">19 March 1996</collectingDate>
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</date>
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, (
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<collectionCode box="[1256,1294,1398,1422]" pageId="10" pageNumber="274">PZ</collectionCode>
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).
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</materialsCitation>
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<materialsCitation ID-GBIF-Occurrence="3111458310" collectingDate="1996-02" collectingDateMax="1996-03-19" collectingDateMin="1996-02" collectionCode="PZ, PMSL" country="Philippines" elevation="300" location="Salakot Falls" municipality="Salakot Falls Road" pageId="10" pageNumber="274" specimenCount="24" specimenCount-female="2" specimenCount-male="22" stateProvince="Palawan" typeStatus="holotype">
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Additional
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<typeStatus box="[835,954,1434,1458]" pageId="10" pageNumber="274">paratypes</typeStatus>
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:
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<collectingRegion box="[968,1071,1433,1457]" country="Philippines" name="Palawan" pageId="10" pageNumber="274">Palawan</collectingRegion>
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, San Rafael, Ulanguan, end of
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<date box="[835,1006,1469,1493]" pageId="10" pageNumber="274" value="1996-02">
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<collectingDate box="[835,1006,1469,1493]" pageId="10" pageNumber="274" value="1996-02">February 1996</collectingDate>
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</date>
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,
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<specimenCount box="[1018,1052,1469,1493]" pageId="10" pageNumber="274" type="male">6♂</specimenCount>
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(
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<collectionCode box="[1067,1104,1469,1493]" pageId="10" pageNumber="274">PZ</collectionCode>
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),
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<specimenCount box="[1125,1158,1469,1493]" pageId="10" pageNumber="274" type="male">1♂</specimenCount>
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,
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<specimenCount box="[1173,1206,1469,1493]" pageId="10" pageNumber="274" type="female">1♀</specimenCount>
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(
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<collectionCode box="[1221,1298,1469,1493]" collectionName="Slovenia, Ljubljana, Slovenian Museum of Natural History" country="Slovenia" httpUri="http://grbio.org/cool/4mab-159w" name="Slovenian Museum of Natural History (Prirodosloveni Muzej Slovenije)" pageId="10" pageNumber="274">PMSL</collectionCode>
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).
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</materialsCitation>
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Specimens deposited in the PZ collection, or the
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<collectionCode box="[1332,1404,1505,1529]" country="Slovenia" httpUri="http://grbio.org/cool/4mab-159w" name="Slovenian Museum of Natural History (Prirodosloveni Muzej Slovenije)" pageId="10" pageNumber="274">PMSL</collectionCode>
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as indicated.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="275" pageId="10" pageNumber="274" type="description">
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<paragraph blockId="10.[835,1444,1326,1920]" pageId="10" pageNumber="274">
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<emphasis bold="true" box="[835,1017,1575,1599]" pageId="10" pageNumber="274">Adult habitus.</emphasis>
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General color pale yellow-brown, patterned with pale brown. Head pale brown, slightly darker anterior to M-line (
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<figureCitation box="[1231,1310,1647,1671]" captionStart="Figs" captionStartId="12.[189,241,829,853]" captionTargetBox="[209,1426,197,826]" captionTargetId="figure-112@12.[631,1001,204,742]" captionTargetPageId="12" captionText="Figs. 45-50. Phanoperla circumspina. 45. Head and pronotum. 46. Male terminalia. 47-50. Penis tube and everted sac. 47. Ventral aspect. 48. Dorsal aspect. 49. Lateral aspect. 50. Apical aspect." figureDoi="http://doi.org/10.5281/zenodo.4760587" httpUri="https://zenodo.org/record/4760587/files/figure.png" pageId="10" pageNumber="274">Fig. 45</figureCitation>
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). Pronotum pale with scattered darker rugosities and narrow dark band along median suture. Ocelli almost touching.
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</paragraph>
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<paragraph blockId="10.[835,1444,1326,1920]" lastBlockId="11.[189,797,1361,1919]" lastPageId="11" lastPageNumber="275" pageId="10" pageNumber="274">
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<emphasis bold="true" box="[835,904,1789,1813]" pageId="10" pageNumber="274">Male.</emphasis>
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Forewing length
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<quantity box="[1107,1176,1789,1813]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="274" unit="mm" value="9.0">9 mm</quantity>
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. Tergum 9 with a small median and two larger lateral patches of sensilla (
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<figureCitation captionStart="Figs" captionStartId="12.[189,241,829,853]" captionTargetBox="[209,1426,197,826]" captionTargetId="figure-112@12.[631,1001,204,742]" captionTargetPageId="12" captionText="Figs. 45-50. Phanoperla circumspina. 45. Head and pronotum. 46. Male terminalia. 47-50. Penis tube and everted sac. 47. Ventral aspect. 48. Dorsal aspect. 49. Lateral aspect. 50. Apical aspect." figureDoi="http://doi.org/10.5281/zenodo.4760587" httpUri="https://zenodo.org/record/4760587/files/figure.png" pageId="10" pageNumber="274">Fig. 46</figureCitation>
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). Hemiterga typical, without special modifications, hemitergal lobes short, wide basally, pointed at the apex and triangular in outline. Penis tube subequal to sac in length, without lobes or armature (
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<figureCitation box="[311,438,1432,1457]" captionStart="Figs" captionStartId="12.[189,241,829,853]" captionTargetBox="[209,1426,197,826]" captionTargetId="figure-112@12.[631,1001,204,742]" captionTargetPageId="12" captionText="Figs. 45-50. Phanoperla circumspina. 45. Head and pronotum. 46. Male terminalia. 47-50. Penis tube and everted sac. 47. Ventral aspect. 48. Dorsal aspect. 49. Lateral aspect. 50. Apical aspect." figureDoi="http://doi.org/10.5281/zenodo.4760587" httpUri="https://zenodo.org/record/4760587/files/figure.png" pageId="11" pageNumber="275">Figs. 47-50</figureCitation>
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); dorsal margin of tube weakly sclerotized, ventral margin membranous. Sac armed near apex with double ring of widely spaced, large black spines, mostly clustered in groups of three; a few small spines located near sac base, and intermingled with large black spines; apex a small cylinder armed over most of surface with small black spines.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.4760585" ID-Zenodo-Dep="4760585" box="[189,1370,1255,1279]" httpUri="https://zenodo.org/record/4760585/files/figure.png" pageId="11" pageNumber="275" startId="11.[189,241,1255,1279]" targetBox="[249,1383,271,1192]" targetPageId="11">
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<paragraph blockId="11.[189,1370,1255,1279]" box="[189,1370,1255,1279]" pageId="11" pageNumber="275">
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Figs. 41-44.
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<taxonomicName authorityName="Sivec & Stark" authorityYear="2011" box="[327,511,1255,1279]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="11" pageNumber="275" phylum="Arthropoda" rank="species" species="batac">
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<emphasis box="[327,511,1255,1279]" italics="true" pageId="11" pageNumber="275">Phanoperla batac</emphasis>
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</taxonomicName>
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egg. 41. Entire egg. 42. Chorionic detail. 43. Anterior end. 44. Collar end.
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</paragraph>
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</caption>
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<paragraph blockId="11.[189,797,1361,1919]" pageId="11" pageNumber="275">
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<emphasis bold="true" box="[189,283,1717,1741]" pageId="11" pageNumber="275">Female.</emphasis>
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Forewing length
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<quantity box="[487,567,1717,1741]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="275" unit="mm" value="11.0">11 mm</quantity>
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. Genitalic structures damaged.
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</paragraph>
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<paragraph blockId="11.[189,797,1361,1919]" lastBlockId="11.[835,1444,1361,1920]" pageId="11" pageNumber="275">
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<emphasis bold="true" box="[189,241,1788,1812]" pageId="11" pageNumber="275">Egg.</emphasis>
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Outline oval with short, wide collar (
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<figureCitation box="[701,784,1788,1812]" captionStart="Figs" captionStartId="12.[189,241,1894,1918]" captionTargetBox="[249,1383,925,1847]" captionTargetId="figure-137@12.[249,1383,925,1847]" captionTargetPageId="12" captionText="Figs. 51-54. Phanoperla circumspina egg. 51. Entire egg. 52. Collar end. 53. Chorionic detail. 54. Anterior end." figureDoi="http://doi.org/10.5281/zenodo.4760589" httpUri="https://zenodo.org/record/4760589/files/figure.png" pageId="11" pageNumber="275">Fig. 51</figureCitation>
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). Total length ca. 227 μm, equatorial width ca. 176 μm, collar length ca. 23 μm. Chorion covered throughout with coarse, circular punctations, becoming less abundant on lid (
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<figureCitation box="[1053,1188,1361,1385]" captionStart="Figs" captionStartId="12.[189,241,1894,1918]" captionTargetBox="[249,1383,925,1847]" captionTargetId="figure-137@12.[249,1383,925,1847]" captionTargetPageId="12" captionText="Figs. 51-54. Phanoperla circumspina egg. 51. Entire egg. 52. Collar end. 53. Chorionic detail. 54. Anterior end." figureDoi="http://doi.org/10.5281/zenodo.4760589" httpUri="https://zenodo.org/record/4760589/files/figure.png" pageId="11" pageNumber="275">Figs. 52-54</figureCitation>
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). Collar rim scarcely flanged, sides bearing a series of vertical ridges.
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</paragraph>
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<paragraph blockId="11.[835,1444,1361,1920]" box="[835,1042,1433,1457]" pageId="11" pageNumber="275">
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<emphasis bold="true" box="[835,910,1433,1457]" pageId="11" pageNumber="275">Larva.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="275" type="etymology">
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<paragraph blockId="11.[835,1444,1361,1920]" pageId="11" pageNumber="275">
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<emphasis bold="true" box="[835,973,1469,1493]" pageId="11" pageNumber="275">Etymology.</emphasis>
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The species name refers to the circular arrangement of large spines near the penial sac apex.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="13" lastPageNumber="277" pageId="11" pageNumber="275" type="diagnosis">
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<paragraph blockId="11.[835,1444,1361,1920]" lastBlockId="13.[189,798,225,391]" lastPageId="13" lastPageNumber="277" pageId="11" pageNumber="275">
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<emphasis bold="true" box="[835,964,1540,1564]" pageId="11" pageNumber="275">Diagnosis.</emphasis>
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This species is a member of the
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<taxonomicName authority="(Klapalek, 1910)" baseAuthorityName="Klapalek" baseAuthorityYear="1910" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="11" pageNumber="275" phylum="Arthropoda" rank="species" species="flaveola">
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<emphasis box="[1333,1443,1540,1564]" italics="true" pageId="11" pageNumber="275">P. flaveola</emphasis>
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(
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<bibRefCitation author="Klapalek, F." box="[843,1022,1576,1600]" pageId="11" pageNumber="275" pagination="219 - 222" refId="ref6144" refString="Klapalek, F. 1910. Uber die Neoperla- Arten aus Java. Notes Leyden Museum, 33: 219 - 222." type="journal article" year="1910">Klapálek, 1910</bibRefCitation>
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)
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</taxonomicName>
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species group and appears to be closely related to
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<taxonomicName baseAuthorityName="Klapalek" baseAuthorityYear="1910" box="[1042,1153,1611,1635]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="11" pageNumber="275" phylum="Arthropoda" rank="species" species="flaveola">
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<emphasis box="[1042,1153,1611,1635]" italics="true" pageId="11" pageNumber="275">P. flaveola</emphasis>
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</taxonomicName>
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, a species reported from several of the
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<collectingCountry box="[997,1211,1647,1671]" name="Philippines" pageId="11" pageNumber="275">Philippine islands</collectingCountry>
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, including
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<collectingRegion box="[1342,1443,1647,1671]" country="Philippines" name="Palawan" pageId="11" pageNumber="275">Palawan</collectingRegion>
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(
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<bibRefCitation author="Zwick, P." box="[843,981,1682,1706]" pageId="11" pageNumber="275" pagination="87 - 126" refId="ref6194" refString="Zwick, P. 1982. A revision of the Oriental stonefly genus Phanoperla (Plecoptera: Perlidae). Systematic Entomology, 7: 87 - 126." type="journal article" year="1982">Zwick 1982</bibRefCitation>
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) and to
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<taxonomicName authority="Sivec & Stark, 2011" authorityName="Sivec & Stark" authorityYear="2011" box="[1081,1242,1683,1706]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="11" pageNumber="275" phylum="Arthropoda" rank="species" species="magnaspina" status="sp. n.">
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<emphasis box="[1081,1242,1683,1706]" italics="true" pageId="11" pageNumber="275">P. magnaspina</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1251,1315,1683,1706]" pageId="11" pageNumber="275" rank="species">sp. n.</taxonomicNameLabel>
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, described below. Externally,
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<taxonomicName baseAuthorityName="Klapalek" baseAuthorityYear="1910" box="[1063,1178,1718,1742]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="11" pageNumber="275" phylum="Arthropoda" rank="species" species="flaveola">
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<emphasis box="[1063,1178,1718,1742]" italics="true" pageId="11" pageNumber="275">P. flaveola</emphasis>
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</taxonomicName>
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differs from the new species in the absence of a median patch of sensilla and small sclerite on tergum 9, and the two differ subtly in details of the penial armature. The distal ring of spines in the new species has three, much larger spines ventrally and small gaps adjacent to the larger spines, whereas the distal ring for
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<taxonomicName baseAuthorityName="Klapalek" baseAuthorityYear="1910" box="[684,797,225,249]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="277" phylum="Arthropoda" rank="species" species="flaveola">
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<emphasis box="[684,797,225,249]" italics="true" pageId="13" pageNumber="277">P. flaveola</emphasis>
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</taxonomicName>
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lacks gaps adjacent to the larger spines. In addition, the proximal spine groupings in
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<taxonomicName baseAuthorityName="Klapalek" baseAuthorityYear="1910" box="[571,681,296,320]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="277" phylum="Arthropoda" rank="species" species="flaveola">
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|
<emphasis box="[571,681,296,320]" italics="true" pageId="13" pageNumber="277">P. flaveola</emphasis>
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</taxonomicName>
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consist of five spines and in
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<emphasis box="[397,564,332,355]" italics="true" pageId="13" pageNumber="277">
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|
<taxonomicName authorityName="Sivec & Stark" authorityYear="2011" box="[397,560,332,355]" class="Insecta" family="Perlidae" genus="Phanoperla" kingdom="Animalia" order="Plecoptera" pageId="13" pageNumber="277" phylum="Arthropoda" rank="species" species="circumspina">P. circumspina</taxonomicName>
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,
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</emphasis>
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they consist of three spines.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |