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<document id="15C985B0E26F6BD227828F280770B4FC" ID-DOI="10.11646/zootaxa.4018.1.2" ID-GBIF-Dataset="b3ca8a6f-989a-4e27-905d-21f7db7de681" ID-ISSN="1175-5326" ID-Zenodo-Dep="289448" ID-ZooBank="F378489D-A55C-4027-B02A-2F7CFBE4724A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461251106684" checkinUser="plazi" docAuthor="Li, Xinran &amp; Wang, Zongqing" docDate="2015" docId="ED20879CFF91FF96FF584B7A8504C372" docLanguage="en" docName="zt04018p056.pdf" docOrigin="Zootaxa 4018 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Diploptera maculata Hanitsch 1925" docType="treatment" docVersion="7" lastPageNumber="53" masterDocId="1119FFE4FF83FF84FFCF4A15843BC778" masterDocTitle="A taxonomic study of the beetle cockroaches (Diploptera Saussure) from China, with notes on the genus and species worldwide (Blattodea: Blaberidae: Diplopterinae)" masterLastPageNumber="56" masterPageNumber="35" pageNumber="53" updateTime="1698607473429" updateUser="plazi">
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<mods:title id="8B8B33EBDCFD8E7C2B6E07012DE2FAE9">A taxonomic study of the beetle cockroaches (Diploptera Saussure) from China, with notes on the genus and species worldwide (Blattodea: Blaberidae: Diplopterinae)</mods:title>
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<mods:name id="23D4783AA036843BC963C8BDD93D09ED" type="personal">
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<mods:namePart id="69E8AB25416B19F6B564FE2E288456BB">Li, Xinran</mods:namePart>
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<mods:namePart id="087AE7A426F2179A359D177CC30304D8">Wang, Zongqing</mods:namePart>
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<mods:title id="BCBA75BE86837C396AF42FCDE790ABD8">Zootaxa</mods:title>
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<mods:date id="4AD0CD104DC8EC4417204D066BA3221F">2015</mods:date>
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<mods:number id="643E3618BC3F53B3265D8D7F75C187D4">2015-09-14</mods:number>
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<mods:number id="217F38F13DBF1BD6351C94666D298D00">4018</mods:number>
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<treatment id="ED20879CFF91FF96FF584B7A8504C372" ID-DOI="http://doi.org/10.5281/zenodo.6102069" ID-GBIF-Taxon="127693087" ID-Zenodo-Dep="6102069" LSID="urn:lsid:plazi:treatment:ED20879CFF91FF96FF584B7A8504C372" httpUri="http://treatment.plazi.org/id/ED20879CFF91FF96FF584B7A8504C372" lastPageNumber="53" pageId="18" pageNumber="53">
<subSubSection id="2D936501FF91FF96FF584B7A866BC6F2" box="[151,592,367,394]" pageId="18" pageNumber="53" type="nomenclature">
<paragraph id="6536368AFF91FF96FF584B7A866BC6F2" blockId="18.[151,592,367,427]" box="[151,592,367,394]" pageId="18" pageNumber="53">
<heading id="3E7E81E6FF91FF96FF584B7A866BC6F2" bold="true" box="[151,592,367,394]" fontSize="11" level="1" pageId="18" pageNumber="53" reason="1">
<taxonomicName id="A2894D09FF91FF96FF584B7A866BC6F2" ID-CoL="6CXTN" authority="Hanitsch, 1925" authorityName="Hanitsch" authorityYear="1925" box="[151,592,367,394]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="53" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="57FDEA98FF91FF96FF584B7A866BC6F2" bold="true" box="[151,592,367,394]" pageId="18" pageNumber="53">
<emphasis id="57FDEA98FF91FF96FF584B7A85B4C6F1" bold="true" box="[151,399,367,393]" italics="true" pageId="18" pageNumber="53">Diploptera maculata</emphasis>
<bibRefCitation id="01184B7BFF91FF96FE594B65866BC6F2" author="Hanitsch, R." box="[406,592,368,394]" firstAuthor="Hanitsch" journalOrPublisher="Sarawak Museum Journal" pageId="18" pageNumber="53" pagination="75 - 106" part="8" refId="ref12545" refString="Hanitsch, R. (1925) On a collection of Blattidae from Northern Sarawak, chiefly Mt. Murud and Mt. Dulit. Sarawak Museum Journal, 8, 75 - 106." title="On a collection of Blattidae from Northern Sarawak, chiefly Mt. Murud and Mt. Dulit" type="journal article" year="1925">Hanitsch, 1925</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2D936501FF91FF96FF584B878563C6D3" box="[151,344,402,427]" pageId="18" pageNumber="53" type="description">
<paragraph id="6536368AFF91FF96FF584B878563C6D3" blockId="18.[151,592,367,427]" box="[151,344,402,427]" pageId="18" pageNumber="53">
(
<figureCitation id="FDB22A0FFF91FF96FF504B8784C7C6D3" box="[159,252,402,427]" captionStart="FIGURES 2 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 3 D. punctata; 2 male; 3 female; 4 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="18" pageNumber="53">Figs. 14</figureCitation>
,
<figureCitation id="FDB22A0FFF91FF96FEC74B87856BC6D3" box="[264,336,402,427]" captionStart="FIGURES 66 71" captionStartId="15.[151,264,1767,1790]" captionTargetBox="[152,1435,842,1741]" captionTargetId="figure@15.[151,1436,841,1747]" captionTargetPageId="15" captionText="FIGURES 66 71. Diploptera spp. distributed outside China, types and labels. Part 2 / 2. 66 68 D. maculata Hanitsch, 1925, female HOLOTYPE (OUM) from Borneo, photographed by Katherine Child and provided by Amoret Spooner, copyright © Oxford University Museum of Natural History; 69 71 D. bicolor Hanitsch, 1925, male HOLOTYPE (OUM) from Borneo, photographed by Katherine Child and provided by Amoret Spooner, copyright © Oxford University Museum of Natural History. (Scale bars = 5 mm.)" httpUri="https://zenodo.org/record/289458/files/figure.png" pageId="18" pageNumber="53">6668</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="2D936501FF91FF96FF584BCF80CCC688" box="[151,1271,474,496]" pageId="18" pageNumber="53" type="reference_group">
<paragraph id="6536368AFF91FF96FF584BCF80CCC688" blockId="18.[151,1271,474,496]" box="[151,1271,474,496]" pageId="18" pageNumber="53">
<treatmentCitationGroup id="459911A4FF91FF96FF584BCF80CCC688" box="[151,1271,474,496]" pageId="18" pageNumber="53">
<taxonomicName id="A2894D09FF91FF96FF584BCF8604C688" ID-CoL="6CXTN" authority="Hanitsch, 1925: 104" authorityName="Hanitsch" authorityPageNumber="104" authorityYear="1925" box="[151,575,474,496]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="53" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="57FDEA98FF91FF96FF584BCF8551C688" box="[151,362,474,496]" italics="true" pageId="18" pageNumber="53">Diploptera maculata</emphasis>
<treatmentCitation id="E428109BFF91FF96FEBF4BCF8604C688" author="Hanitsch" box="[368,575,474,496]" page="104" pageId="18" pageNumber="53" year="1925">
<bibRefCitation id="01184B7BFF91FF96FEBF4BCF8604C688" author="Hanitsch, R." box="[368,575,474,496]" firstAuthor="Hanitsch" journalOrPublisher="Sarawak Museum Journal" page="104" pageId="18" pageNumber="53" pagination="75 - 106" part="8" refId="ref12545" refString="Hanitsch, R. (1925) On a collection of Blattidae from Northern Sarawak, chiefly Mt. Murud and Mt. Dulit. Sarawak Museum Journal, 8, 75 - 106." title="On a collection of Blattidae from Northern Sarawak, chiefly Mt. Murud and Mt. Dulit" type="journal article" year="1925">Hanitsch, 1925: 104</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, figs. 17, 18, locality Borneo;
<treatmentCitation id="E428109BFF91FF96FCBD4BCF8073C688" author="Bruijning" box="[882,1096,474,496]" page="152" pageId="18" pageNumber="53" year="1948">
<bibRefCitation id="01184B7BFF91FF96FCBD4BCF8073C688" author="Bruijning, C. F. A." box="[882,1096,474,496]" firstAuthor="Bruijning" journalOrPublisher="Zoologische Mededeelingen" page="152" pageId="18" pageNumber="53" pagination="1 - 174" part="29" refId="ref12271" refString="Bruijning, C. F. A. (1948) Studies of Malayan Blattidae. Zoologische Mededeelingen, 29, 1 - 174." title="Studies of Malayan Blattidae" type="journal article" year="1948">Bruijning, 1948: 152</bibRefCitation>
</treatmentCitation>
, locality Borneo.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="2D936501FF91FF96FF58480880A2C401" pageId="18" pageNumber="53" type="discussion">
<paragraph id="6536368AFF91FF96FF584808857CC54E" blockId="18.[151,1437,541,1034]" box="[151,327,541,566]" pageId="18" pageNumber="53">
<collectingCountry id="1D9E761AFF91FF96FF58480884EAC54E" box="[151,209,541,566]" name="Malta" pageId="18" pageNumber="53">Male</collectingCountry>
unknown.
</paragraph>
<paragraph id="6536368AFF91FF96FF08485484D5C476" blockId="18.[151,1437,541,1034]" pageId="18" pageNumber="53">
<emphasis id="57FDEA98FF91FF96FF08485485ADC522" bold="true" box="[199,406,577,602]" pageId="18" pageNumber="53">Taxonomic notes.</emphasis>
<bibRefCitation id="01184B7BFF91FF96FE5248548669C522" author="Hanitsch, R." box="[413,594,577,602]" firstAuthor="Hanitsch" journalOrPublisher="Sarawak Museum Journal" pageId="18" pageNumber="53" pagination="75 - 106" part="8" refId="ref12545" refString="Hanitsch, R. (1925) On a collection of Blattidae from Northern Sarawak, chiefly Mt. Murud and Mt. Dulit. Sarawak Museum Journal, 8, 75 - 106." title="On a collection of Blattidae from Northern Sarawak, chiefly Mt. Murud and Mt. Dulit" type="journal article" year="1925">Hanitsch (1925)</bibRefCitation>
has proposed an interesting likelihood that
<taxonomicName id="A2894D09FF91FF96FBF4485780FEC521" box="[1083,1221,577,601]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="53" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="57FDEA98FF91FF96FBF4485780FEC521" box="[1083,1221,577,601]" italics="true" pageId="18" pageNumber="53">D. maculata</emphasis>
</taxonomicName>
is the female of
<taxonomicName id="A2894D09FF91FF96FA4E485784D3C505" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="53" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="57FDEA98FF91FF96FA4E485784D3C505" italics="true" pageId="18" pageNumber="53">D. bicolor</emphasis>
</taxonomicName>
based on the two
<typeStatus id="BA328828FF91FF96FE7E48708625C506" box="[433,542,613,638]" pageId="18" pageNumber="53" type="holotype">holotypes</typeStatus>
being almost the same except for the gender and the markings on the pronotum; even their different markings share the same contour. A second specimen of
<taxonomicName id="A2894D09FF91FF96FBF5489F80F0C5D9" box="[1082,1227,649,673]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="53" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="57FDEA98FF91FF96FBF5489F80F0C5D9" box="[1082,1227,649,673]" italics="true" pageId="18" pageNumber="53">D. maculata</emphasis>
</taxonomicName>
was recorded by
<bibRefCitation id="01184B7BFF91FF96FF5848B8856FC5BE" author="Bruijning, C. F. A." box="[151,340,685,710]" firstAuthor="Bruijning" journalOrPublisher="Zoologische Mededeelingen" pageId="18" pageNumber="53" pagination="1 - 174" part="29" refId="ref12271" refString="Bruijning, C. F. A. (1948) Studies of Malayan Blattidae. Zoologische Mededeelingen, 29, 1 - 174." title="Studies of Malayan Blattidae" type="journal article" year="1948">Bruijning (1948)</bibRefCitation>
but “unluckily” was the same gender, female, as the
<typeStatus id="BA328828FF91FF96FC6948B88037C5BE" box="[934,1036,685,710]" pageId="18" pageNumber="53" type="holotype">holotype</typeStatus>
. Similarly, a second specimen of
<taxonomicName id="A2894D09FF91FF96FA4E48BB84D3C591" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="53" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="57FDEA98FF91FF96FA4E48BB84D3C591" italics="true" pageId="18" pageNumber="53">D. bicolor</emphasis>
</taxonomicName>
(see below for details) is a male. So the hypothesis that they are the same species has not yet been proven or refuted.
</paragraph>
<paragraph id="6536368AFF91FF96FF08490F80A2C401" blockId="18.[151,1437,541,1034]" pageId="18" pageNumber="53">
In addition, it is noteworthy that Hanitschs illustration was drawn by Mr. V. Knight instead of Hanitsch himself. This is probably why the image has something discrepant with the true insect,
<emphasis id="57FDEA98FF91FF96FBB1492B809DC42D" box="[1150,1190,830,853]" italics="true" pageId="18" pageNumber="53">e.g.</emphasis>
, the size of head, the shape of pronotum, and the seam between tegmina. A similar situation occurs in
<taxonomicName id="A2894D09FF91FF96FBEE497780A9C401" box="[1057,1170,865,889]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="18" pageNumber="53" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="57FDEA98FF91FF96FBEE497780A9C401" box="[1057,1170,865,889]" italics="true" pageId="18" pageNumber="53">D. bicolor</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="2D936501FF91FF96FF0849908504C372" pageId="18" pageNumber="53" type="materials_examined">
<paragraph id="6536368AFF91FF96FF084990865EC4BA" blockId="18.[151,1437,541,1034]" pageId="18" pageNumber="53">
<emphasis id="57FDEA98FF91FF96FF08499085CDC4E6" bold="true" box="[199,502,901,926]" pageId="18" pageNumber="53">
<typeStatus id="BA328828FF91FF96FF0849908539C4E6" box="[199,258,901,926]" pageId="18" pageNumber="53">Type</typeStatus>
material examined.
</emphasis>
<materialsCitation id="D5E13CD7FF91FF96FDCE4993865EC4BA" ID-GBIF-Occurrence="3923978320" collectingDate="1922-10" collectingDateMax="1923-01" collectingDateMin="1922-10" collectionCode="OUM" collectorName="Dr. E. Mjoberg" country="Malaysia" location="Pah Trap" pageId="18" pageNumber="53" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="BA328828FF91FF96FDCE499386AEC4E6" box="[513,661,902,926]" pageId="18" pageNumber="53" type="holotype">HOLOTYPE</typeStatus>
(
<collectionCode id="0398AE4FFF91FF96FD68499386D0C4E6" box="[679,747,902,926]" httpUri="http://grbio.org/cool/g6yf-wft0" name="Oxford University Museum of Natural History" pageId="18" pageNumber="53">OUM</collectionCode>
) fixed by monotypy,
<specimenCount id="738FFD03FF91FF96FC2449908001C4E6" box="[1003,1082,901,926]" pageId="18" pageNumber="53" type="female">female</specimenCount>
, “
<location id="60566051FF91FF96FB9B499080F9C4E5" LSID="urn:lsid:plazi:treatment:ED20879CFF91FF96FF584B7A8504C372:60566051FF91FF96FB9B499080F9C4E5" box="[1108,1218,901,926]" country="Malaysia" name="Pah Trap" pageId="18" pageNumber="53">Pah Trap</location>
,
<location id="60566051FF91FF96FB1F4990811EC4E6" LSID="urn:lsid:plazi:treatment:ED20879CFF91FF96FF584B7A8504C372:60566051FF91FF96FB1F4990811EC4E6" box="[1232,1317,901,926]" country="Malaysia" name="Kalabit" pageId="18" pageNumber="53">Kalabit</location>
,
<collectingCountry id="1D9E761AFF91FF96FAFA499081A3C4E6" box="[1333,1432,901,926]" name="Malaysia" pageId="18" pageNumber="53">Sarawak</collectingCountry>
,
<date id="1137104AFF91FF96FF5849BC84DEC4BA" box="[151,229,937,962]" pageId="18" pageNumber="53" value="1922-10">
<collectingDate id="0173E9A2FF91FF96FF5849BC84DEC4BA" box="[151,229,937,962]" pageId="18" pageNumber="53" value="1922-10">x.1922</collectingDate>
</date>
<date id="1137104AFF91FF96FF3A49BC8507C4BA" box="[245,316,937,962]" pageId="18" pageNumber="53" value="1923-01">
<collectingDate id="0173E9A2FF91FF96FF3A49BC8507C4BA" box="[245,316,937,962]" pageId="18" pageNumber="53" value="1923-01">i.1923</collectingDate>
</date>
, Coll. by
<collectorName id="C87C535CFF91FF96FE6749BF866CC4BA" box="[424,599,937,962]" pageId="18" pageNumber="53">Dr. E. Mjöberg</collectorName>
”.
</materialsCitation>
</paragraph>
<paragraph id="6536368AFF91FF96FF0849D88504C372" blockId="18.[151,1437,541,1034]" pageId="18" pageNumber="53">
<emphasis id="57FDEA98FF91FF96FF0849D8862AC49E" bold="true" box="[199,529,973,998]" pageId="18" pageNumber="53">Other material examined.</emphasis>
<materialsCitation id="D5E13CD7FF91FF96FDED49DB8504C372" ID-GBIF-Occurrence="3923978328" pageId="18" pageNumber="53">
<specimenCount id="738FFD03FF91FF96FDED49DB86B5C49E" box="[546,654,973,998]" pageId="18" pageNumber="53" type="female">1 female</specimenCount>
(
<collectionCode id="0398AE4FFF91FF96FD6849DB873AC49E" LSID="urn:lsid:biocol.org:col:34992" box="[679,769,974,998]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="18" pageNumber="53">RMNH</collectionCode>
), “Z. Kenepai, 1894, Jan”, coll. “Borneo Exped. Dr. Nieuwenhuis”.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>