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<mods:title id="0ABBCFE6AC451C7DDE5B495B4C0A48F2">A taxonomic study of the beetle cockroaches (Diploptera Saussure) from China, with notes on the genus and species worldwide (Blattodea: Blaberidae: Diplopterinae)</mods:title>
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<mods:namePart id="E0A7447AFFC625A38B1358A2A9C6549F">Li, Xinran</mods:namePart>
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<paragraph id="6536368AFF87FF80FF584F518645C227" blockId="4.[151,638,1348,1408]" box="[151,638,1348,1375]" pageId="4" pageNumber="39">
<heading id="3E7E81E6FF87FF80FF584F518645C227" bold="true" box="[151,638,1348,1375]" fontSize="11" level="1" pageId="4" pageNumber="39" reason="1">
<taxonomicName id="A2894D09FF87FF80FF584F518645C227" ID-CoL="6CXF3" authority="Eschscholtz, 1822" authorityName="Eschscholtz" authorityYear="1822" box="[151,638,1348,1375]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF87FF80FF584F518645C227" bold="true" box="[151,638,1348,1375]" pageId="4" pageNumber="39">
<emphasis id="57FDEA98FF87FF80FF584F5185B3C226" bold="true" box="[151,392,1348,1374]" italics="true" pageId="4" pageNumber="39">Diploptera punctata</emphasis>
(
<bibRefCitation id="01184B7BFF87FF80FE584F50864DC227" author="Eschscholtz, J. F. von" bookContentInfo="128 pp." box="[407,630,1349,1375]" firstAuthor="Eschscholtz" journalOrPublisher="G. Reimer, Berlin" pageId="4" pageNumber="39" refId="ref12361" refString="Eschscholtz, J. F. von (1822) Entomographien. G. Reimer, Berlin, 128 pp. http: // dx. doi. org / 10.5962 / bhl. title. 5121" title="Entomographien" type="book" year="1822">Eschscholtz, 1822</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
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<subSubSection id="2D936501FF87FF80FF584F728603C2F8" box="[151,568,1383,1408]" pageId="4" pageNumber="39" type="description">
<paragraph id="6536368AFF87FF80FF584F728603C2F8" blockId="4.[151,638,1348,1408]" box="[151,568,1383,1408]" pageId="4" pageNumber="39">
(
<figureCitation id="FDB22A0FFF87FF80FF504F728530C2F8" box="[159,267,1383,1408]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="4" pageNumber="39">Figs. 2–3</figureCitation>
, 
<figureCitation id="FDB22A0FFF87FF80FED94F7D8565C2F8" box="[278,350,1384,1408]" captionStart="FIGURES 17 – 30" captionStartId="7.[151,264,1796,1819]" captionTargetBox="[191,1384,195,1758]" captionTargetId="figure@7.[191,1385,195,1760]" captionTargetPageId="7" captionText="FIGURES 17 – 30. Diploptera spp. &amp; subspp. distributed in China. 17 – 18 D. punctata, female from Yunnan, Menglun; 19 – 21 D. punctata, lectotype (ZMMU) from Hawaii instead of China, photographed by Dr. Andrey L. Ozerov, copyright ZMMU, published with permission; 22 – 24 D. elliptica sp. n.; 22 – 23 male holotype from Yunnan, Cangyuan, Banhong; 24 female paratype from Yunnan, Mengla, Mohan; 25 – 26 D. naevus sp. n., male holotype from Xizang, Motuo, Beibeng; 27 – 28 D. nigrescens guani subsp. n., male holotype from Guangxi, Chongzuo, Daqingshan; 29 – 30 D. nigrescens nigrescens, male syntype (NTU) from Taiwan, photographed by Yi-Ho Liu and downloaded from NTU website, copyright Insect Museum Digital Archives Project, Department of Entomology, National Taiwan University. (Scale bars = 5 mm.)" httpUri="https://zenodo.org/record/289451/files/figure.png" pageId="4" pageNumber="39">17–21</figureCitation>
, 
<figureCitation id="FDB22A0FFF87FF80FEA54F728588C2F8" box="[362,435,1383,1408]" captionStart="FIGURES 31 – 36. D" captionStartId="9.[151,264,1815,1838]" captionTargetBox="[151,1433,198,1789]" captionTargetId="figure@9.[151,1436,195,1791]" captionTargetPageId="9" captionText="FIGURES 31 – 36. D. punctata (Eschscholtz, 1822) from China, Yunnan, Menglun. 31 hind wing, female, with grey coloration indicating sclerotization; 32 – 36 male; 32 terminal segment of dorsal abdomen, in ventral view; 33 subgenital plate in dorsal view; 34 – 36 phallic sclerites in dorsal view; 34 sclerites R’; 35 sclerites L 2 ’; 36 apex of left complex hook, showing sclerite L 3 ’. (ce. = cercus, m. = membranous, p. p. = paraprocts, st. = stylus; other abbreviations as indicated in the text. Scale bars = 10 mm for 31, 1 mm for 32 &amp; 33, 500 µm for 34 – 36.)" httpUri="https://zenodo.org/record/289452/files/figure.png" pageId="4" pageNumber="39">31–36</figureCitation>
, 
<figureCitation id="FDB22A0FFF87FF80FE714F72863CC2F8" box="[446,519,1383,1408]" captionStart="FIGURES 54 – 56. 54 D" captionStartId="13.[151,264,1787,1810]" captionTargetBox="[151,1436,193,1766]" captionTargetId="figure@13.[151,1436,193,1766]" captionTargetPageId="13" captionText="FIGURES 54 – 56. 54 D. punctata eating lemon peel in Hawaii, 2006. XI. 29, image by Forest &amp; Kim Starr (University of Hawaii); 55 Cypress with damage by D. punctata in Hawaii, 2005. VII. 11, image by Forest &amp; Kim Starr (University of Hawaii); 56 D. nigrescens nigrescens Shiraki, 1931 from Taiwan, Chiayi County, Tashan (= Xiaotashan), about 1200 m, photographed by Gaga (Taiwan). The insect was found near a light trap but not on the curtain." httpUri="https://zenodo.org/record/289456/files/figure.png" pageId="4" pageNumber="39">54–55</figureCitation>
, 
<figureCitation id="FDB22A0FFF87FF80FDDD4F7D8614C2F8" box="[530,559,1384,1408]" captionStart="FIGURE 80. A" captionStartId="19.[151,250,1644,1666]" captionTargetBox="[151,1435,658,1622]" captionTargetId="figure@19.[151,1435,658,1622]" captionTargetPageId="19" captionText="FIGURE 80. A male D. punctata in repose near light trap. Photographed by Lu Qiu in Nonggang National Nature Reserve in Guangxi, China. We are very lucky to get this timely photo just before the present paper is published. This is a light-colored individual of the species. Its large abdomen was retracted to be wholly covered by the tegmina when it became a specimen." httpUri="https://zenodo.org/record/289461/files/figure.png" pageId="4" pageNumber="39">80</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="2D936501FF87FF80FF584FA58609C06F" pageId="4" pageNumber="39" type="reference_group">
<paragraph id="6536368AFF87FF80FF584FA58539C17B" blockId="4.[151,1437,1456,1815]" pageId="4" pageNumber="39">
<treatmentCitationGroup id="459911A4FF87FF80FF584FA58539C17B" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584FA5861AC2BE" ID-CoL="M3B5" authority="Eschscholtz, 1822: 86" authorityName="Eschscholtz" authorityPageNumber="86" authorityYear="1822" box="[151,545,1456,1478]" class="Insecta" family="Blaberidae" genus="Blatta" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF87FF80FF584FA5850EC2BE" box="[151,309,1456,1478]" italics="true" pageId="4" pageNumber="39">Blatta punctata</emphasis>
<treatmentCitation id="E428109BFF87FF80FEF24FA5861AC2BE" author="Eschscholtz" box="[317,545,1456,1478]" page="86" pageId="4" pageNumber="39" year="1822">
<bibRefCitation id="01184B7BFF87FF80FEF24FA5861AC2BE" author="Eschscholtz, J. F. von" bookContentInfo="128 pp." box="[317,545,1456,1478]" firstAuthor="Eschscholtz" journalOrPublisher="G. Reimer, Berlin" page="86" pageId="4" pageNumber="39" refId="ref12361" refString="Eschscholtz, J. F. von (1822) Entomographien. G. Reimer, Berlin, 128 pp. http: // dx. doi. org / 10.5962 / bhl. title. 5121" title="Entomographien" type="book" year="1822">Eschscholtz, 1822: 86</bibRefCitation>
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, locality “Sandwichsinseln”, = Sandwich Islands, now Hawaiian Islands; 
<treatmentCitation id="E428109BFF87FF80FAD54FA584D4C29C" author="Kirby" page="151" pageId="4" pageNumber="39" year="1904">
<bibRefCitation id="01184B7BFF87FF80FAD54FA584D4C29C" author="Kirby, W. F." bookContentInfo="501 pp." firstAuthor="Kirby" journalOrPublisher="British Museum, London" page="151" pageId="4" pageNumber="39" refId="ref12618" refString="Kirby, W. F. (1904) A Synonymic Catalogue of Orthoptera. Vol. I. Orthoptera Euplexoptera, Cursoria, et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae.). British Museum, London, 501 pp." title="A Synonymic Catalogue of Orthoptera. Vol. I. Orthoptera Euplexoptera, Cursoria, et Gressoria. (Forficulidae, Hemimeridae, Blattidae, Mantidae, Phasmidae.)" type="book" year="1904">Kirby, 1904: 151</bibRefCitation>
</treatmentCitation>
, syn. of 
<taxonomicName id="A2894D09FF87FF80FE834FDB86BCC29C" authority="(L.)" baseAuthorityName="L." box="[332,647,1486,1509]" class="Insecta" family="Blaberidae" genus="Leucophaea" higherTaxonomySource="CoL" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="surinamensis">
<emphasis id="57FDEA98FF87FF80FE834FDB866EC29D" box="[332,597,1486,1509]" italics="true" pageId="4" pageNumber="39">Leucophaea surinamensis</emphasis>
(L.)
</taxonomicName>
; 
<treatmentCitation id="E428109BFF87FF80FD5B4FDB876FC29C" author="Princis" box="[660,852,1486,1508]" page="162" pageId="4" pageNumber="39" year="1950">
<bibRefCitation id="01184B7BFF87FF80FD5B4FDB876FC29C" author="Princis, K." box="[660,852,1486,1508]" firstAuthor="Princis" journalOrPublisher="Opuscula Entomologica" page="162" pageId="4" pageNumber="39" pagination="161 - 188" part="15" refId="ref12911" refString="Princis, K. (1950) Indomalaiische und australische Blattarien aus dem Entomologischen Museum der Universitat in Lund. Opuscula Entomologica, 15, 161 - 188." title="Indomalaiische und australische Blattarien aus dem Entomologischen Museum der Universitat in Lund" type="journal article" year="1950">Princis, 1950: 162</bibRefCitation>
</treatmentCitation>
, placed in 
<taxonomicName id="A2894D09FF87FF80FC074FDB800CC29C" ID-CoL="8MQVN" authority="Saussure, 1864" authorityName="Saussure" authorityYear="1864" box="[968,1079,1486,1508]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF87FF80FC074FDB800CC29C" box="[968,1079,1486,1508]" italics="true" pageId="4" pageNumber="39">Diploptera</emphasis>
</taxonomicName>
. [not 
<taxonomicName id="A2894D09FF87FF80FBB84FDB84C7C17B" ID-CoL="M3B5" authority="Charpentier, 1825" authorityName="Charpentier" authorityYear="1825" class="Insecta" family="Blaberidae" genus="Blatta" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF87FF80FBB84FDB812CC29D" box="[1143,1303,1486,1509]" italics="true" pageId="4" pageNumber="39">Blatta punctata</emphasis>
Charpentier, 1825
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]
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</paragraph>
<paragraph id="6536368AFF87FF80FF584C19856DC138" blockId="4.[151,1437,1456,1815]" pageId="4" pageNumber="39">
<treatmentCitationGroup id="459911A4FF87FF80FF584C19856DC138" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584C198628C15A" ID-CoL="M353" authority="Serville, 1839: 102" authorityName="Serville" authorityPageNumber="102" authorityYear="1839" box="[151,531,1548,1570]" class="Insecta" family="Blaberidae" genus="Blatta" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="dytiscoides">
<emphasis id="57FDEA98FF87FF80FF584C198570C15A" box="[151,331,1548,1570]" italics="true" pageId="4" pageNumber="39">Blatta dytiscoides</emphasis>
<treatmentCitation id="E428109BFF87FF80FE9E4C198628C15A" author="Serville" box="[337,531,1548,1570]" page="102" pageId="4" pageNumber="39" year="1839">
<bibRefCitation id="01184B7BFF87FF80FE9E4C198628C15A" author="Serville, J. G. A." bookContentInfo="776 pp." box="[337,531,1548,1570]" firstAuthor="Serville" journalOrPublisher="Librairie Encyclopedique de Roret, Paris" page="102" pageId="4" pageNumber="39" refId="ref13217" refString="Serville, J. G. A. (1839) Histoire Naturelle des Insectes. Orthopteres. Librairie Encyclopedique de Roret, Paris, 776 pp. [J. G. A. - Serville = J. G. Audinet-Serville]" title="Histoire Naturelle des Insectes. Orthopteres" type="book" year="1839">Serville, 1839: 102</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, locality “ 
<collectingCountry id="1D9E761AFF87FF80FDB74C19877BC15A" box="[632,832,1548,1570]" name="Australia" pageId="4" pageNumber="39">Nouvelle-Hollande</collectingCountry>
”, = 
<collectingCountry id="1D9E761AFF87FF80FCA84C1887CBC15A" box="[871,1008,1548,1570]" name="Australia" pageId="4" pageNumber="39">New Holland</collectingCountry>
, now 
<collectingCountry id="1D9E761AFF87FF80FBE44C1980B0C15A" box="[1067,1163,1548,1570]" name="Australia" pageId="4" pageNumber="39">Australia</collectingCountry>
; 
<treatmentCitation id="E428109BFF87FF80FB5A4C19816BC15A" author="Walker" box="[1173,1360,1548,1570]" page="125" pageId="4" pageNumber="39" year="1869">
<bibRefCitation id="01184B7BFF87FF80FB5A4C19816BC15A" author="Walker, F." bookContentInfo="224 pp." box="[1173,1360,1548,1570]" firstAuthor="Walker" journalOrPublisher="British Museum, London" page="125" pageId="4" pageNumber="39" refId="ref13338" refString="Walker, F. (1869) Catalogue of the Specimens of Dermaptera Saltatoria and Supplement to the Blattariae in the Collection of the British Museum. British Museum, London, 224 pp." title="Catalogue of the Specimens of Dermaptera Saltatoria and Supplement to the Blattariae in the Collection of the British Museum" type="book" year="1869">Walker, 1869: 125</bibRefCitation>
</treatmentCitation>
, placed in 
<taxonomicName id="A2894D09FF87FF80FF2E4C3F856BC138" ID-CoL="8MQVN" authority="Saussure, 1864" authorityName="Saussure" authorityYear="1864" box="[225,336,1578,1600]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF87FF80FF2E4C3F856BC138" box="[225,336,1578,1600]" italics="true" pageId="4" pageNumber="39">Diploptera</emphasis>
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.
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</paragraph>
<paragraph id="6536368AFF87FF80FF584C5C86FBC127" blockId="4.[151,1437,1456,1815]" box="[151,704,1609,1631]" pageId="4" pageNumber="39">
<treatmentCitationGroup id="459911A4FF87FF80FF584C5C86FBC127" box="[151,704,1609,1631]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584C5C8687C127" authority="Saussure, 1864a: 325" authorityName="Saussure" authorityPageNumber="325" authorityYear="1864" box="[151,700,1609,1631]" class="Insecta" family="Ectobiidae" genus="Prosoplecta" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="silpha" subGenus="Diploptera">
<emphasis id="57FDEA98FF87FF80FF584C5C85E0C127" box="[151,475,1609,1631]" italics="true" pageId="4" pageNumber="39">Prosoplecta (Diploptera) silpha</emphasis>
<treatmentCitation id="E428109BFF87FF80FE2E4C5C8687C127" author="Saussure" box="[481,700,1609,1631]" page="325" pageId="4" pageNumber="39" year="1864">
<bibRefCitation id="01184B7BFF87FF80FE2E4C5C8687C127" author="Saussure, H. de" box="[481,700,1609,1631]" firstAuthor="Saussure" journalOrPublisher="Revue et Magasin de Zoologie Pure et Appliquee" page="325" pageId="4" pageNumber="39" pagination="305 - 326" part="16" refId="ref13151" refString="Saussure, H. de (1864 a) Blattarum novarum species aliquot. Revue et Magasin de Zoologie Pure et Appliquee, 16, 305 - 326." title="Blattarum novarum species aliquot" type="journal article" year="1864" yearSuffix="a">Saussure, 1864a: 325</bibRefCitation>
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.
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</paragraph>
<paragraph id="6536368AFF87FF80FF584C7D8506C1E4" blockId="4.[151,1437,1456,1815]" pageId="4" pageNumber="39">
<treatmentCitationGroup id="459911A4FF87FF80FF584C7D8506C1E4" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584C7D8601C106" ID-CoL="8N9MS" authority="Saussure, 1864b: 178" authorityName="Saussure" authorityPageNumber="178" authorityYear="1864" box="[151,570,1640,1662]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="silpha">
<emphasis id="57FDEA98FF87FF80FF584C7D8577C106" box="[151,332,1640,1662]" italics="true" pageId="4" pageNumber="39">Diploptera silpha</emphasis>
: 
<treatmentCitation id="E428109BFF87FF80FE944C7D8601C106" author="Saussure" box="[347,570,1640,1662]" page="178" pageId="4" pageNumber="39" year="1864">
<bibRefCitation id="01184B7BFF87FF80FE944C7D8601C106" author="Saussure, H. de" bookContentInfo="279 pp." box="[347,570,1640,1662]" firstAuthor="Saussure" journalOrPublisher="Geneva" page="178" pageId="4" pageNumber="39" refId="ref13180" refString="Saussure, H. de (1864 b) Memoires pour Servir a l'Histoire Naturelle du Mexique des Antilles et des Etats-Unis. Vol. 4. Orthopteres de l'Amerique Moyenne. Geneva, 279 pp." title="Memoires pour Servir a l'Histoire Naturelle du Mexique des Antilles et des Etats-Unis. Vol. 4. Orthopteres de l'Amerique Moyenne" type="book" year="1864" yearSuffix="b">Saussure, 1864b: 178</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, fig. 28, locality “ 
<collectingCountry id="1D9E761AFF87FF80FD384C7D8762C106" box="[759,857,1640,1662]" name="Australia" pageId="4" pageNumber="39">Australie</collectingCountry>
”, = 
<collectingCountry id="1D9E761AFF87FF80FC494C7D87DDC106" box="[902,998,1640,1662]" name="Australia" pageId="4" pageNumber="39">Australia</collectingCountry>
; 
<treatmentCitation id="E428109BFF87FF80FC3D4C7D8089C106" author="Walker" box="[1010,1202,1640,1662]" page="125" pageId="4" pageNumber="39" year="1869">
<bibRefCitation id="01184B7BFF87FF80FC3D4C7D8089C106" author="Walker, F." bookContentInfo="224 pp." box="[1010,1202,1640,1662]" firstAuthor="Walker" journalOrPublisher="British Museum, London" page="125" pageId="4" pageNumber="39" refId="ref13338" refString="Walker, F. (1869) Catalogue of the Specimens of Dermaptera Saltatoria and Supplement to the Blattariae in the Collection of the British Museum. British Museum, London, 224 pp." title="Catalogue of the Specimens of Dermaptera Saltatoria and Supplement to the Blattariae in the Collection of the British Museum" type="book" year="1869">Walker, 1869: 125</bibRefCitation>
</treatmentCitation>
, synonymized with 
<taxonomicName id="A2894D09FF87FF80FA4C4C7C8503C1E4" ID-CoL="M353" authority="Serville, 1839: 102" authorityName="Serville" authorityPageNumber="102" authorityYear="1839" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="dytiscoides">
<emphasis id="57FDEA98FF87FF80FA4C4C7C8503C1E4" italics="true" pageId="4" pageNumber="39">D. dytiscoides</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6536368AFF87FF80FF584CB080CCC1C3" blockId="4.[151,1437,1456,1815]" box="[151,1271,1701,1723]" pageId="4" pageNumber="39">
<treatmentCitationGroup id="459911A4FF87FF80FF584CB080CCC1C3" box="[151,1271,1701,1723]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584CB086C5C1C3" authority="Brunner von Wattenwyl, 1865" authorityName="Brunner von Wattenwyl" authorityYear="1865" box="[151,766,1701,1723]" class="Insecta" family="Blaberidae" genus="Eleutheroda" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="dytiscoides">
<emphasis id="57FDEA98FF87FF80FF584CB085B2C1C3" box="[151,393,1701,1723]" italics="true" pageId="4" pageNumber="39">Eleutheroda dytiscoides</emphasis>
: 
<treatmentCitation id="E428109BFF87FF80FE584CB386C5C1C3" author="Brunner von Wattenwyl, C." box="[407,766,1701,1723]" page="265" pageId="4" pageNumber="39" year="1865">
<bibRefCitation id="01184B7BFF87FF80FE584CB386C5C1C3" author="Brunner von Wattenwyl, C." bookContentInfo="426 pp." box="[407,766,1701,1723]" firstAuthor="Brunner von Wattenwyl" journalOrPublisher="Vienna" page="265" pageId="4" pageNumber="39" refId="ref12296" refString="Brunner von Wattenwyl, C. (1865) Nouveau Systeme des Blattaires. Vienna, 426 pp." title="Nouveau Systeme des Blattaires" type="book" year="1865">Brunner von Wattenwyl, 1865: 265</bibRefCitation>
</treatmentCitation>
</taxonomicName>
, fig. 29; 
<treatmentCitation id="E428109BFF87FF80FC974CB0802FC1C3" author="Walker" box="[856,1044,1701,1723]" page="125" pageId="4" pageNumber="39" year="1869">
<bibRefCitation id="01184B7BFF87FF80FC974CB0802FC1C3" author="Walker, F." bookContentInfo="224 pp." box="[856,1044,1701,1723]" firstAuthor="Walker" journalOrPublisher="British Museum, London" page="125" pageId="4" pageNumber="39" refId="ref13338" refString="Walker, F. (1869) Catalogue of the Specimens of Dermaptera Saltatoria and Supplement to the Blattariae in the Collection of the British Museum. British Museum, London, 224 pp." title="Catalogue of the Specimens of Dermaptera Saltatoria and Supplement to the Blattariae in the Collection of the British Museum" type="book" year="1869">Walker, 1869: 125</bibRefCitation>
</treatmentCitation>
, placed in 
<taxonomicName id="A2894D09FF87FF80FB4D4CB080CAC1C3" ID-CoL="8MQVN" authority="Saussure, 1864" authorityName="Saussure" authorityYear="1864" box="[1154,1265,1701,1723]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF87FF80FB4D4CB080CAC1C3" box="[1154,1265,1701,1723]" italics="true" pageId="4" pageNumber="39">Diploptera</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6536368AFF87FF80FF584CD18578C180" blockId="4.[151,1437,1456,1815]" pageId="4" pageNumber="39">
<treatmentCitationGroup id="459911A4FF87FF80FF584CD18578C180" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584CD18673C1A2" ID-CoL="M353" authority="Walker, 1869: 125" authorityName="Walker" authorityPageNumber="125" authorityYear="1869" box="[151,584,1732,1754]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="dytiscoides">
<emphasis id="57FDEA98FF87FF80FF584CD18546C1A2" box="[151,381,1732,1754]" italics="true" pageId="4" pageNumber="39">Diploptera dytiscoides</emphasis>
: 
<treatmentCitation id="E428109BFF87FF80FE444CD18673C1A2" author="Walker" box="[395,584,1732,1754]" page="125" pageId="4" pageNumber="39" year="1869">
<bibRefCitation id="01184B7BFF87FF80FE444CD18673C1A2" author="Walker, F." bookContentInfo="224 pp." box="[395,584,1732,1754]" firstAuthor="Walker" journalOrPublisher="British Museum, London" page="125" pageId="4" pageNumber="39" refId="ref13338" refString="Walker, F. (1869) Catalogue of the Specimens of Dermaptera Saltatoria and Supplement to the Blattariae in the Collection of the British Museum. British Museum, London, 224 pp." title="Catalogue of the Specimens of Dermaptera Saltatoria and Supplement to the Blattariae in the Collection of the British Museum" type="book" year="1869">Walker, 1869: 125</bibRefCitation>
</treatmentCitation>
</taxonomicName>
; 
<treatmentCitation id="E428109BFF87FF80FD9C4CD08795C1A2" author="Brunner von Wattenwyl, C." box="[595,942,1732,1754]" page="41" pageId="4" pageNumber="39" year="1893">
<bibRefCitation id="01184B7BFF87FF80FD9C4CD08795C1A2" author="Brunner von Wattenwyl, C." box="[595,942,1732,1754]" firstAuthor="Brunner von Wattenwyl" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" page="41" pageId="4" pageNumber="39" pagination="5 - 230" part="2" refId="ref12315" refString="Brunner von Wattenwyl, C. (1893) Revision du systeme des Orthopteres et description des especes rapportees par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 13, 5 - 230." title="Revision du systeme des Orthopteres et description des especes rapportees par M. Leonardo Fea de Birmanie" type="journal article" year="1893">Brunner von Wattenwyl, 1893: 41</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="E428109BFF87FF80FC764CD180AAC1A2" author="Bruijning" box="[953,1169,1732,1754]" page="210" pageId="4" pageNumber="39" year="1947">
<bibRefCitation id="01184B7BFF87FF80FC764CD180AAC1A2" author="Bruijning, C. F. A." box="[953,1169,1732,1754]" firstAuthor="Bruijning" journalOrPublisher="Zoologische Mededelingen" page="210" pageId="4" pageNumber="39" pagination="205 - 252" part="27" refId="ref12233" refString="Bruijning, C. F. A. (1947) An account of the Blattidae (Orthoptera) from Celebes, the Moluccas, and New Guinea. Zoologische Mededelingen, 27, 205 - 252." title="An account of the Blattidae (Orthoptera) from Celebes, the Moluccas, and New Guinea" type="journal article" year="1947">Bruijning, 1947: 210</bibRefCitation>
</treatmentCitation>
, 239; 
<treatmentCitation id="E428109BFF87FF80FB014CD181ACC1A2" author="Bruijning" box="[1230,1431,1732,1754]" page="41" pageId="4" pageNumber="39" year="1948">
<bibRefCitation id="01184B7BFF87FF80FB014CD181ACC1A2" author="Bruijning, C. F. A." box="[1230,1431,1732,1754]" firstAuthor="Bruijning" journalOrPublisher="Zoologische Mededeelingen" page="41" pageId="4" pageNumber="39" pagination="1 - 174" part="29" refId="ref12271" refString="Bruijning, C. F. A. (1948) Studies of Malayan Blattidae. Zoologische Mededeelingen, 29, 1 - 174." title="Studies of Malayan Blattidae" type="journal article" year="1948">Bruijning, 1948: 41</bibRefCitation>
</treatmentCitation>
, 152, fig. 22.
</treatmentCitationGroup>
</paragraph>
<paragraph id="6536368AFF87FF80FF584D148609C06F" blockId="4.[151,1437,1456,1815]" box="[151,562,1793,1815]" pageId="4" pageNumber="39">
<treatmentCitationGroup id="459911A4FF87FF80FF584D148609C06F" box="[151,562,1793,1815]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584D148615C06F" ID-CoL="6CXF3" authority="Princis, 1950: 162" authorityName="Princis" authorityPageNumber="162" authorityYear="1950" box="[151,558,1793,1815]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF87FF80FF584D14855EC06F" box="[151,357,1793,1815]" italics="true" pageId="4" pageNumber="39">Diploptera punctata</emphasis>
: 
<treatmentCitation id="E428109BFF87FF80FEBD4D148615C06F" author="Princis" box="[370,558,1793,1815]" page="162" pageId="4" pageNumber="39" year="1950">
<bibRefCitation id="01184B7BFF87FF80FEBD4D148615C06F" author="Princis, K." box="[370,558,1793,1815]" firstAuthor="Princis" journalOrPublisher="Opuscula Entomologica" page="162" pageId="4" pageNumber="39" pagination="161 - 188" part="15" refId="ref12911" refString="Princis, K. (1950) Indomalaiische und australische Blattarien aus dem Entomologischen Museum der Universitat in Lund. Opuscula Entomologica, 15, 161 - 188." title="Indomalaiische und australische Blattarien aus dem Entomologischen Museum der Universitat in Lund" type="journal article" year="1950">Princis, 1950: 162</bibRefCitation>
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="2D936501FF87FF80FF584D518639C0B0" pageId="4" pageNumber="39" type="diagnosis">
<paragraph id="6536368AFF87FF80FF584D518639C0B0" blockId="4.[151,1436,1860,1993]" pageId="4" pageNumber="39">
<emphasis id="57FDEA98FF87FF80FF584D51852AC025" bold="true" box="[151,273,1860,1885]" pageId="4" pageNumber="39">Diagnosis.</emphasis>
<taxonomicName id="A2894D09FF87FF80FED84D5085A0C024" box="[279,411,1861,1884]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF87FF80FED84D5085A0C024" box="[279,411,1861,1884]" italics="true" pageId="4" pageNumber="39">D. punctata</emphasis>
</taxonomicName>
is unspotted and relatively large among the species known in 
<taxonomicName id="A2894D09FF87FF80FB834D5180FCC024" box="[1100,1223,1860,1884]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF87FF80FB834D5180FCC024" box="[1100,1223,1860,1884]" italics="true" pageId="4" pageNumber="39">Diploptera</emphasis>
</taxonomicName>
, and may look like 
<taxonomicName id="A2894D09FF87FF80FF584D7C852EC0F8" box="[151,277,1896,1920]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="elliptica" status="sp. nov.">
<emphasis id="57FDEA98FF87FF80FF584D7C852EC0F8" box="[151,277,1896,1920]" italics="true" pageId="4" pageNumber="39">D. elliptica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF87FF80FED34D728567C0F8" box="[284,348,1895,1920]" pageId="4" pageNumber="39" rank="species">
<emphasis id="57FDEA98FF87FF80FED34D728567C0F8" bold="true" box="[284,348,1895,1920]" pageId="4" pageNumber="39">sp. n.</emphasis>
</taxonomicNameLabel>
at first sight but can be distinguished by the pronotum with posterolateral angles. Also it may be mixed up with 
<taxonomicName id="A2894D09FF87FF80FE844D988589C0DC" box="[331,434,1933,1956]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="57FDEA98FF87FF80FE844D988589C0DC" box="[331,434,1933,1956]" italics="true" pageId="4" pageNumber="39">D. minor</emphasis>
</taxonomicName>
, 
<taxonomicName id="A2894D09FF87FF80FE0B4D9886A8C0DC" box="[452,659,1932,1956]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="erythrocephala">
<emphasis id="57FDEA98FF87FF80FE0B4D9886A8C0DC" box="[452,659,1932,1956]" italics="true" pageId="4" pageNumber="39">D. erythrocephala</emphasis>
</taxonomicName>
or 
<taxonomicName id="A2894D09FF87FF80FD0F4D98871CC0DC" box="[704,807,1933,1956]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="57FDEA98FF87FF80FD0F4D98871CC0DC" box="[704,807,1933,1956]" italics="true" pageId="4" pageNumber="39">D. parva</emphasis>
</taxonomicName>
, which are small-sized and have yellowish to reddish brown legs instead of dark ones.
</paragraph>
</subSubSection>
<caption id="31F66602FF86FF81FF584D1A851AC0D8" httpUri="https://zenodo.org/record/289449/files/figure.png" pageId="5" pageNumber="40" targetBox="[151,1436,195,1782]" targetPageId="5">
<paragraph id="6536368AFF86FF81FF584D1A851AC0D8" blockId="5.[151,1436,1807,1952]" pageId="5" pageNumber="40">
<emphasis id="57FDEA98FF86FF81FF584D1A852CC05C" bold="true" box="[151,279,1807,1829]" pageId="5" pageNumber="40">FIGURE 1.</emphasis>
Distribution of 
<taxonomicName id="A2894D09FF86FF81FE084D1A860DC05D" box="[455,566,1807,1829]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF86FF81FE084D1A860DC05D" box="[455,566,1807,1829]" italics="true" pageId="5" pageNumber="40">Diploptera</emphasis>
</taxonomicName>
in China and Southeast Asia. The precise localities are painted black, whilst the imprecise ones are in white; each type locality is marked with a ring; the (sub)species are indicated by numbers in alphabetical order: 1 
<taxonomicName id="A2894D09FF86FF81FF204D58856CC01A" box="[239,343,1868,1890]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="57FDEA98FF86FF81FF204D58856CC01A" box="[239,343,1868,1890]" italics="true" pageId="5" pageNumber="40">D. bicolor</emphasis>
</taxonomicName>
, 2 
<taxonomicName id="A2894D09FF86FF81FEB74D5885D0C01A" box="[376,491,1868,1890]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="elliptica" status="sp. nov.">
<emphasis id="57FDEA98FF86FF81FEB74D5885D0C01A" box="[376,491,1868,1890]" italics="true" pageId="5" pageNumber="40">D. elliptica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF86FF81FE3C4D588616C01A" box="[499,557,1869,1890]" pageId="5" pageNumber="40" rank="species">
<emphasis id="57FDEA98FF86FF81FE3C4D588616C01A" bold="true" box="[499,557,1869,1890]" pageId="5" pageNumber="40">sp. n.</emphasis>
</taxonomicNameLabel>
, 3 
<taxonomicName id="A2894D09FF86FF81FD814D588733C01A" box="[590,776,1868,1890]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="erythrocephala">
<emphasis id="57FDEA98FF86FF81FD814D588733C01A" box="[590,776,1868,1890]" italics="true" pageId="5" pageNumber="40">D. erythrocephala</emphasis>
</taxonomicName>
, 4 
<taxonomicName id="A2894D09FF86FF81FCE54D588793C01A" box="[810,936,1868,1890]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="57FDEA98FF86FF81FCE54D588793C01A" box="[810,936,1868,1890]" italics="true" pageId="5" pageNumber="40">D. maculata</emphasis>
</taxonomicName>
, 5 
<taxonomicName id="A2894D09FF86FF81FC054D58801DC01A" box="[970,1062,1869,1890]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="57FDEA98FF86FF81FC054D58801DC01A" box="[970,1062,1869,1890]" italics="true" pageId="5" pageNumber="40">D. minor</emphasis>
</taxonomicName>
, 6 
<taxonomicName id="A2894D09FF86FF81FB884D588096C01A" box="[1095,1197,1869,1890]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="naevus" status="sp. nov.">
<emphasis id="57FDEA98FF86FF81FB884D588096C01A" box="[1095,1197,1869,1890]" italics="true" pageId="5" pageNumber="40">D. naevus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF86FF81FB7A4D5880D4C01A" box="[1205,1263,1869,1890]" pageId="5" pageNumber="40" rank="species">
<emphasis id="57FDEA98FF86FF81FB7A4D5880D4C01A" bold="true" box="[1205,1263,1869,1890]" pageId="5" pageNumber="40">sp. n.</emphasis>
</taxonomicNameLabel>
, 7 
<taxonomicName id="A2894D09FF86FF81FADE4D5884EAC0F9" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="subSpecies" species="nigrescens" status="subsp. nov." subSpecies="guani">
<emphasis id="57FDEA98FF86FF81FADE4D5884EAC0F9" italics="true" pageId="5" pageNumber="40">D. nigrescens guani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF86FF81FF184D7E850CC0F8" box="[215,311,1899,1920]" pageId="5" pageNumber="40" rank="subSpecies">
<emphasis id="57FDEA98FF86FF81FF184D7E850CC0F8" bold="true" box="[215,311,1899,1920]" pageId="5" pageNumber="40">subsp. n.</emphasis>
</taxonomicNameLabel>
, 8 
<taxonomicName id="A2894D09FF86FF81FE974D79866DC0F9" box="[344,598,1900,1921]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="subSpecies" species="nigrescens" subSpecies="nigrescens">
<emphasis id="57FDEA98FF86FF81FE974D79866DC0F9" box="[344,598,1900,1921]" italics="true" pageId="5" pageNumber="40">D. nigrescens nigrescens</emphasis>
</taxonomicName>
, 9 
<taxonomicName id="A2894D09FF86FF81FDB84D7986EAC0F9" box="[631,721,1900,1921]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="57FDEA98FF86FF81FDB84D7986EAC0F9" box="[631,721,1900,1921]" italics="true" pageId="5" pageNumber="40">D. parva</emphasis>
</taxonomicName>
, × 
<taxonomicName id="A2894D09FF86FF81FD3B4D798756C0F9" box="[756,877,1900,1921]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF86FF81FD3B4D798756C0F9" box="[756,877,1900,1921]" italics="true" pageId="5" pageNumber="40">D. punctata</emphasis>
</taxonomicName>
; asterisk stands for the habitat of the Thai 
<taxonomicName id="A2894D09FF86FF81FAEB4D7981A7C0F9" box="[1316,1436,1900,1921]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="5" pageNumber="40" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF86FF81FAEB4D7981A7C0F9" box="[1316,1436,1900,1921]" italics="true" pageId="5" pageNumber="40">D. punctata</emphasis>
</taxonomicName>
(see the text).
</paragraph>
</caption>
<caption id="31F66602FF85FF82FF5849AF8760C333" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="6" pageNumber="41" targetBox="[160,1418,196,938]" targetPageId="6">
<paragraph id="6536368AFF85FF82FF5849AF8760C333" blockId="6.[151,1437,954,1099]" pageId="6" pageNumber="41">
<emphasis id="57FDEA98FF85FF82FF5849AF8571C4B7" bold="true" box="[151,330,954,977]" pageId="6" pageNumber="41">FIGURES 2–16.</emphasis>
Pronota of 
<taxonomicName id="A2894D09FF85FF82FE0449AF8601C4A8" box="[459,570,954,976]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF85FF82FE0449AF8601C4A8" box="[459,570,954,976]" italics="true" pageId="6" pageNumber="41">Diploptera</emphasis>
</taxonomicName>
spp. &amp; subspp. presented in monochrome, 
<emphasis id="57FDEA98FF85FF82FBC749AE801CC4A8" box="[1032,1063,955,976]" italics="true" pageId="6" pageNumber="41">i.e.</emphasis>
, white indicates unicolor or lighter coloration, whilst black is for darker portion. 
<emphasis id="57FDEA98FF85FF82FD4F49CC869EC496" bold="true" box="[640,677,985,1006]" pageId="6" pageNumber="41">2–3</emphasis>
<taxonomicName id="A2894D09FF85FF82FD7F49CF8717C497" box="[688,812,986,1007]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF85FF82FD7F49CF8717C497" box="[688,812,986,1007]" italics="true" pageId="6" pageNumber="41">D. punctata</emphasis>
</taxonomicName>
; 
<emphasis id="57FDEA98FF85FF82FCF249CC8772C496" bold="true" box="[829,841,985,1006]" pageId="6" pageNumber="41">2</emphasis>
male; 
<emphasis id="57FDEA98FF85FF82FC5849CC879FC496" bold="true" box="[919,932,985,1006]" pageId="6" pageNumber="41">3</emphasis>
female; 
<emphasis id="57FDEA98FF85FF82FBCB49CC8011C496" bold="true" box="[1028,1066,985,1006]" pageId="6" pageNumber="41">4–5</emphasis>
<taxonomicName id="A2894D09FF85FF82FBFB49CF80A8C497" box="[1076,1171,986,1007]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="57FDEA98FF85FF82FBFB49CF80A8C497" box="[1076,1171,986,1007]" italics="true" pageId="6" pageNumber="41">D. minor</emphasis>
</taxonomicName>
; 
<emphasis id="57FDEA98FF85FF82FB6A49CC8089C496" bold="true" box="[1189,1202,985,1006]" pageId="6" pageNumber="41">4</emphasis>
female; 
<emphasis id="57FDEA98FF85FF82FADC49CC811BC496" bold="true" box="[1299,1312,985,1006]" pageId="6" pageNumber="41">5</emphasis>
male; 
<emphasis id="57FDEA98FF85FF82FAA349CC8142C496" bold="true" box="[1388,1401,985,1006]" pageId="6" pageNumber="41">6</emphasis>
<taxonomicName id="A2894D09FF85FF82FA4C49CF850AC376" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="erythrocephala">
<emphasis id="57FDEA98FF85FF82FA4C49CF850AC376" italics="true" pageId="6" pageNumber="41">D. erythrocephala</emphasis>
</taxonomicName>
, male; 
<emphasis id="57FDEA98FF85FF82FEB249ED8598C375" bold="true" box="[381,419,1016,1037]" pageId="6" pageNumber="41">7–8</emphasis>
<taxonomicName id="A2894D09FF85FF82FE6549EC863EC376" box="[426,517,1017,1038]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="57FDEA98FF85FF82FE6549EC863EC376" box="[426,517,1017,1038]" italics="true" pageId="6" pageNumber="41">D. parva</emphasis>
</taxonomicName>
; 
<emphasis id="57FDEA98FF85FF82FDDC49ED861BC375" bold="true" box="[531,544,1016,1037]" pageId="6" pageNumber="41">7</emphasis>
male; 
<emphasis id="57FDEA98FF85FF82FDA949ED8648C375" bold="true" box="[614,627,1016,1037]" pageId="6" pageNumber="41">8</emphasis>
female; 
<emphasis id="57FDEA98FF85FF82FD0249ED86E1C375" bold="true" box="[717,730,1016,1037]" pageId="6" pageNumber="41">9</emphasis>
<taxonomicName id="A2894D09FF85FF82FD2F49EC877DC376" box="[736,838,1017,1038]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="naevus" status="sp. nov.">
<emphasis id="57FDEA98FF85FF82FD2F49EC877DC376" box="[736,838,1017,1038]" italics="true" pageId="6" pageNumber="41">D. naevus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF85FF82FC8249ED87BCC375" box="[845,903,1016,1037]" pageId="6" pageNumber="41" rank="species">
<emphasis id="57FDEA98FF85FF82FC8249ED87BCC375" bold="true" box="[845,903,1016,1037]" pageId="6" pageNumber="41">sp. n.</emphasis>
</taxonomicNameLabel>
, male; 
<emphasis id="57FDEA98FF85FF82FC1B49ED87D5C375" bold="true" box="[980,1006,1016,1037]" pageId="6" pageNumber="41">10</emphasis>
<taxonomicName id="A2894D09FF85FF82FC3B49EC80CAC376" box="[1012,1265,1017,1038]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="nigrescens" subSpecies="nigrescens">
<emphasis id="57FDEA98FF85FF82FC3B49EC80CAC376" box="[1012,1265,1017,1038]" italics="true" pageId="6" pageNumber="41">D. nigrescens nigrescens</emphasis>
</taxonomicName>
, male; 
<emphasis id="57FDEA98FF85FF82FAF149ED8140C375" bold="true" box="[1342,1403,1016,1037]" pageId="6" pageNumber="41">11–12</emphasis>
<taxonomicName id="A2894D09FF85FF82FA4C49EC857EC354" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="subSpecies" species="nigrescens" status="subsp. nov." subSpecies="guani">
<emphasis id="57FDEA98FF85FF82FA4C49EC857EC354" italics="true" pageId="6" pageNumber="41">D. nigrescens guani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF85FF82FE824E028595C354" box="[333,430,1047,1068]" pageId="6" pageNumber="41" rank="subSpecies">
<emphasis id="57FDEA98FF85FF82FE824E028595C354" bold="true" box="[333,430,1047,1068]" pageId="6" pageNumber="41">subsp. n.</emphasis>
</taxonomicNameLabel>
; 
<emphasis id="57FDEA98FF85FF82FE724E0385E8C353" bold="true" box="[445,467,1046,1067]" pageId="6" pageNumber="41">11</emphasis>
male; 
<emphasis id="57FDEA98FF85FF82FDD14E038603C353" bold="true" box="[542,568,1046,1067]" pageId="6" pageNumber="41">12</emphasis>
female; 
<emphasis id="57FDEA98FF85FF82FD5A4E038695C353" bold="true" box="[661,686,1046,1067]" pageId="6" pageNumber="41">13</emphasis>
<taxonomicName id="A2894D09FF85FF82FD794E02871BC354" box="[694,800,1046,1068]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="57FDEA98FF85FF82FD794E02871BC354" box="[694,800,1046,1068]" italics="true" pageId="6" pageNumber="41">D. bicolor</emphasis>
</taxonomicName>
, male; 
<emphasis id="57FDEA98FF85FF82FCA04E0387B3C353" bold="true" box="[879,904,1046,1067]" pageId="6" pageNumber="41">14</emphasis>
<taxonomicName id="A2894D09FF85FF82FC5E4E02802BC354" box="[913,1040,1046,1068]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="57FDEA98FF85FF82FC5E4E02802BC354" box="[913,1040,1046,1068]" italics="true" pageId="6" pageNumber="41">D. maculata</emphasis>
</taxonomicName>
, female; 
<emphasis id="57FDEA98FF85FF82FBBB4E038088C353" bold="true" box="[1140,1203,1046,1067]" pageId="6" pageNumber="41">15–16</emphasis>
<taxonomicName id="A2894D09FF85FF82FB744E028114C354" box="[1211,1327,1046,1068]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="elliptica" status="sp. nov.">
<emphasis id="57FDEA98FF85FF82FB744E028114C354" box="[1211,1327,1046,1068]" italics="true" pageId="6" pageNumber="41">D. elliptica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF85FF82FAF74E028148C354" box="[1336,1395,1047,1068]" pageId="6" pageNumber="41" rank="species">
<emphasis id="57FDEA98FF85FF82FAF74E028148C354" bold="true" box="[1336,1395,1047,1068]" pageId="6" pageNumber="41">sp. n.</emphasis>
</taxonomicNameLabel>
; 
<emphasis id="57FDEA98FF85FF82FA4C4E0381A7C353" bold="true" box="[1411,1436,1046,1067]" pageId="6" pageNumber="41">15</emphasis>
male; 
<emphasis id="57FDEA98FF85FF82FF1A4E2084D5C332" bold="true" box="[213,238,1077,1098]" pageId="6" pageNumber="41">16</emphasis>
female. (Dashed lines indicate depression. Scale bar = 2mm)
</paragraph>
</caption>
<subSubSection id="2D936501FF85FF82FF084E60878BC1D5" pageId="6" pageNumber="41" type="description">
<paragraph id="6536368AFF85FF82FF084E60802FC382" blockId="6.[151,1436,1141,2033]" pageId="6" pageNumber="41">
<emphasis id="57FDEA98FF85FF82FF084E608562C3F6" bold="true" box="[199,345,1141,1166]" pageId="6" pageNumber="41">Description.</emphasis>
General. Size medium. 
<typeStatus id="BA328828FF85FF82FDA84E6086E6C3F6" box="[615,733,1141,1166]" pageId="6" pageNumber="41" type="lectotype">
<emphasis id="57FDEA98FF85FF82FDA84E6086E6C3F6" bold="true" box="[615,733,1141,1166]" pageId="6" pageNumber="41">Lectotype</emphasis>
</typeStatus>
, sex unknown, left and right tegmen length × width 
<quantity id="A2719B6FFF85FF82FAE54E6381BEC3F6" box="[1322,1413,1142,1166]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="6" pageNumber="41" unit="mm" value="11.8">11.8mm</quantity>
× 
<quantity id="A2719B6FFF85FF82FF584E8C84D2C3C9" box="[151,233,1177,1201]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="41" unit="mm" value="4.4">4.4mm</quantity>
. 
<collectingCountry id="1D9E761AFF85FF82FF3B4E8C850AC3CA" box="[244,305,1177,1202]" name="Malta" pageId="6" pageNumber="41">
<emphasis id="57FDEA98FF85FF82FF3B4E8C850AC3CA" bold="true" box="[244,305,1177,1202]" pageId="6" pageNumber="41">Male</emphasis>
</collectingCountry>
, overall length (= body length) 17.4–19.0mm; pronotum length × width 
<quantity id="A2719B6FFF85FF82FBA64E8C808CC3C9" box="[1129,1207,1177,1201]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="6" pageNumber="41" unit="mm" value="4.1">4.1mm</quantity>
× 
<quantity id="A2719B6FFF85FF82FB184E8C811CC3C9" box="[1239,1319,1177,1202]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="41" unit="mm" value="7.5">7.5mm</quantity>
− 
<quantity id="A2719B6FFF85FF82FAF94E8C81BEC3C9" box="[1334,1413,1177,1201]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="6" pageNumber="41" unit="mm" value="4.3">4.3mm</quantity>
× 
<quantity id="A2719B6FFF85FF82FF584EAB84D2C3AE" box="[151,233,1214,1238]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="6" pageNumber="41" unit="mm" value="7.9">7.9mm</quantity>
; tegmen length 12.0–13.0mm. 
<emphasis id="57FDEA98FF85FF82FD8B4EA886A0C3AE" bold="true" box="[580,667,1213,1238]" pageId="6" pageNumber="41">Female</emphasis>
, overall length 
<quantity id="A2719B6FFF85FF82FC844EA887D7C3AE" box="[843,1004,1213,1238]" metricMagnitude="-2" metricUnit="m" metricValue="2.1149999999999998" metricValueMax="2.28" metricValueMin="1.95" pageId="6" pageNumber="41" unit="mm" value="21.15" valueMax="22.8" valueMin="19.5">19.5−22.8mm</quantity>
; body length 
<quantity id="A2719B6FFF85FF82FB4C4EA8811FC3AE" box="[1155,1316,1213,1238]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.25" metricValueMin="1.95" pageId="6" pageNumber="41" unit="mm" value="21.0" valueMax="22.5" valueMin="19.5">19.5−22.5mm</quantity>
; pronotum length × width 
<quantity id="A2719B6FFF85FF82FE8E4EF485ABC381" box="[321,400,1249,1273]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="6" pageNumber="41" unit="mm" value="4.4">4.4mm</quantity>
× 
<quantity id="A2719B6FFF85FF82FE614EF485C5C382" box="[430,510,1249,1274]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="41" unit="mm" value="7.5">7.5mm</quantity>
− 
<quantity id="A2719B6FFF85FF82FDC24EF48665C382" box="[525,606,1249,1274]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="6" pageNumber="41" unit="mm" value="4.9">4.9mm</quantity>
× 
<quantity id="A2719B6FFF85FF82FDA34EF48686C382" box="[620,701,1249,1274]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="6" pageNumber="41" unit="mm" value="8.2">8.2mm</quantity>
; tegmen length 
<quantity id="A2719B6FFF85FF82FCA14EF4802BC382" box="[878,1040,1249,1274]" metricMagnitude="-2" metricUnit="m" metricValue="1.655" metricValueMax="1.75" metricValueMin="1.56" pageId="6" pageNumber="41" unit="mm" value="16.55" valueMax="17.5" valueMin="15.6">15.6−17.5mm</quantity>
.
</paragraph>
<paragraph id="6536368AFF85FF82FF084F108051C23A" blockId="6.[151,1436,1141,2033]" pageId="6" pageNumber="41">Body blackish brown to black. Antenna brownish; apical part of clypeus and basal part of labrum yellowish brown. Tegmina brownish, always lighter than pronotum. Tarsi brown. Cerci brownish.</paragraph>
<paragraph id="6536368AFF85FF82FF084F5B8515C281" blockId="6.[151,1436,1141,2033]" pageId="6" pageNumber="41">Pronotum and tegmina setose. Pronotum subtrapezoidal, with posterior margin emarginate near posterolateral angles, lateral borders subsided. Tegmina with moderately dense small punctations throughout and larger and distinct ones at basal area, especially basocostal area, reaching nearly, exactly, or exceeding the end of abdomen, sometimes depending on whether the abdomen is retracted or stretched. Hindwing with AP1 and AP2 reaching the outer margin.</paragraph>
<paragraph id="6536368AFF85FF82FF084C1187F1C139" blockId="6.[151,1436,1141,2033]" pageId="6" pageNumber="41">
<collectingCountry id="1D9E761AFF85FF82FF084C11853FC165" box="[199,260,1540,1565]" name="Malta" pageId="6" pageNumber="41">Male</collectingCountry>
. Sclerite L3’ with basal part broad and suddenly narrowing to the curved part, resulting in an obtuse angle at the inner margin; the apex of hook curved and pointing outward.
</paragraph>
<paragraph id="6536368AFF85FF82FF084C59878BC1D5" blockId="6.[151,1436,1141,2033]" pageId="6" pageNumber="41">Variation of male genitalia. Main variations occur in sclerites L2’. Main sclerite of L2’ varies in degree of curvature, whilst the apical sclerite is more or less irregular, with the margin near the main sclerite uneven and sometimes irregularly serrated, since it is diversiform in sclerotization.</paragraph>
</subSubSection>
<subSubSection id="2D936501FF85FF82FF084CAD803FC18D" pageId="6" pageNumber="41" type="distribution">
<paragraph id="6536368AFF85FF82FF084CAD803FC18D" blockId="6.[151,1436,1141,2033]" pageId="6" pageNumber="41">
<emphasis id="57FDEA98FF85FF82FF084CAD855BC1A9" bold="true" box="[199,352,1720,1745]" pageId="6" pageNumber="41">Distribution.</emphasis>
Hawaii, 
<collectingCountry id="1D9E761AFF85FF82FE134CAD867DC1A9" box="[476,582,1720,1745]" name="Australia" pageId="6" pageNumber="41">Australia</collectingCountry>
, 
<collectingCountry id="1D9E761AFF85FF82FD944CAD8691C1A8" box="[603,682,1720,1744]" name="Samoa" pageId="6" pageNumber="41">Samoa</collectingCountry>
, 
<collectingCountry id="1D9E761AFF85FF82FD0F4CAD870BC1A9" box="[704,816,1720,1745]" name="Indonesia" pageId="6" pageNumber="41">Indonesia</collectingCountry>
, 
<collectingCountry id="1D9E761AFF85FF82FC894CAD8794C1A9" box="[838,943,1720,1745]" name="Malaysia" pageId="6" pageNumber="41">Malaysia</collectingCountry>
, 
<collectingCountry id="1D9E761AFF85FF82FC094CAD807CC1A9" box="[966,1095,1720,1745]" name="Philippines" pageId="6" pageNumber="41">Philippines</collectingCountry>
, 
<collectingCountry id="1D9E761AFF85FF82FB924CAD809BC1A9" box="[1117,1184,1720,1745]" name="China" pageId="6" pageNumber="41">China</collectingCountry>
(Hainan 
<collectingCountry id="1D9E761AFF85FF82FAD24CAD815AC1A9" box="[1309,1377,1720,1745]" name="Iceland" pageId="6" pageNumber="41">Island</collectingCountry>
and southwest of the mainland), 
<collectingCountry id="1D9E761AFF85FF82FE1A4CC9860CC18D" box="[469,567,1756,1781]" name="Vietnam" pageId="6" pageNumber="41">Vietnam</collectingCountry>
, 
<collectingCountry id="1D9E761AFF85FF82FD8D4CC98683C18D" box="[578,696,1756,1781]" name="Cambodia" pageId="6" pageNumber="41">Cambodia</collectingCountry>
, 
<collectingCountry id="1D9E761AFF85FF82FD0C4CC98712C18D" box="[707,809,1756,1781]" name="Thailand" pageId="6" pageNumber="41">Thailand</collectingCountry>
, 
<collectingCountry id="1D9E761AFF85FF82FCFB4CC887B8C18C" box="[820,899,1757,1780]" name="Myanmar" pageId="6" pageNumber="41">Burma</collectingCountry>
, 
<collectingCountry id="1D9E761AFF85FF82FC404CC9803BC18D" box="[911,1024,1756,1781]" name="Sri Lanka" pageId="6" pageNumber="41">Sri Lanka</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="2D936501FF85FF82FF084D1585F3C088" pageId="6" pageNumber="41" type="discussion">
<paragraph id="6536368AFF85FF82FF084D1585F3C088" blockId="6.[151,1436,1141,2033]" pageId="6" pageNumber="41">
<emphasis id="57FDEA98FF85FF82FF084D1585ADC061" bold="true" box="[199,406,1792,1817]" pageId="6" pageNumber="41">Taxonomic notes.</emphasis>
In the original description of 
<taxonomicName id="A2894D09FF85FF82FD294D148777C060" box="[742,844,1792,1816]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="silpha">
<emphasis id="57FDEA98FF85FF82FD294D148777C060" box="[742,844,1792,1816]" italics="true" pageId="6" pageNumber="41">D. silpha</emphasis>
</taxonomicName>
by 
<bibRefCitation id="01184B7BFF85FF82FCB84D158000C061" author="Saussure, H. de" box="[887,1083,1792,1817]" firstAuthor="Saussure" journalOrPublisher="Revue et Magasin de Zoologie Pure et Appliquee" pageId="6" pageNumber="41" pagination="305 - 326" part="16" refId="ref13151" refString="Saussure, H. de (1864 a) Blattarum novarum species aliquot. Revue et Magasin de Zoologie Pure et Appliquee, 16, 305 - 326." title="Blattarum novarum species aliquot" type="journal article" year="1864" yearSuffix="a">Saussure (1864a)</bibRefCitation>
, basal 12 segments of antennae are darker than the rest, but the character was not found or not distinct among examined individuals of 
<taxonomicName id="A2894D09FF85FF82FAD74D3081A7C044" box="[1304,1436,1829,1852]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF85FF82FAD74D3081A7C044" box="[1304,1436,1829,1852]" italics="true" pageId="6" pageNumber="41">D. punctata</emphasis>
</taxonomicName>
except for some from Guangxi, 
<collectingCountry id="1D9E761AFF85FF82FDD84D5D8667C019" box="[535,604,1864,1889]" name="China" pageId="6" pageNumber="41">China</collectingCountry>
, which may represent high contrast in this species. Unfortunately, the antennae are lost in the 
<typeStatus id="BA328828FF85FF82FE624D78863FC0FD" box="[429,516,1901,1925]" pageId="6" pageNumber="41" type="syntype">syntype</typeStatus>
of 
<taxonomicName id="A2894D09FF85FF82FDE04D7886A3C0FC" box="[559,664,1900,1924]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="silpha">
<emphasis id="57FDEA98FF85FF82FDE04D7886A3C0FC" box="[559,664,1900,1924]" italics="true" pageId="6" pageNumber="41">D. silpha</emphasis>
</taxonomicName>
; the body structures left in the 
<typeStatus id="BA328828FF85FF82FBC74D788064C0FD" box="[1032,1119,1901,1925]" pageId="6" pageNumber="41" type="syntype">syntype</typeStatus>
are only the pronotum, the tegmina, one hindwing and one leg and a half, which direct us to a 
<taxonomicName id="A2894D09FF85FF82FC4A4D848032C0D0" box="[901,1033,1937,1960]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF85FF82FC4A4D848032C0D0" box="[901,1033,1937,1960]" italics="true" pageId="6" pageNumber="41">D. punctata</emphasis>
</taxonomicName>
. Since the color pattern of antennae appears to be of intraspecific uniformity among other species, it remains uncertain whether 
<taxonomicName id="A2894D09FF85FF82FB274DA0816FC0B4" box="[1256,1364,1972,1996]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="41" phylum="Arthropoda" rank="species" species="silpha">
<emphasis id="57FDEA98FF85FF82FB274DA0816FC0B4" box="[1256,1364,1972,1996]" italics="true" pageId="6" pageNumber="41">D. silpha</emphasis>
</taxonomicName>
is an independent species or not.
</paragraph>
</subSubSection>
<caption id="31F66602FF84FF83FF584D118090C0AA" httpUri="https://zenodo.org/record/289451/files/figure.png" pageId="7" pageNumber="42" targetBox="[191,1384,195,1758]" targetPageId="7">
<paragraph id="6536368AFF84FF83FF584D118090C0AA" blockId="7.[151,1437,1796,2002]" pageId="7" pageNumber="42">
<emphasis id="57FDEA98FF84FF83FF584D11856EC061" bold="true" box="[151,341,1796,1819]" pageId="7" pageNumber="42">FIGURES 17–30.</emphasis>
<taxonomicName id="A2894D09FF84FF83FE944D1185F1C062" box="[347,458,1796,1818]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF84FF83FE944D1185F1C062" box="[347,458,1796,1818]" italics="true" pageId="7" pageNumber="42">Diploptera</emphasis>
</taxonomicName>
spp. &amp; subspp. distributed in China. 
<emphasis id="57FDEA98FF84FF83FC874D1187BCC061" bold="true" box="[840,903,1796,1817]" pageId="7" pageNumber="42">17–18</emphasis>
<taxonomicName id="A2894D09FF84FF83FC414D10803CC062" box="[910,1031,1797,1818]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF84FF83FC414D10803CC062" box="[910,1031,1797,1818]" italics="true" pageId="7" pageNumber="42">D. punctata</emphasis>
</taxonomicName>
, female from Yunnan, Menglun; 
<emphasis id="57FDEA98FF84FF83FA934D1181A7C061" bold="true" box="[1372,1436,1796,1817]" pageId="7" pageNumber="42">19–21</emphasis>
<taxonomicName id="A2894D09FF84FF83FF584D31852AC041" box="[151,273,1828,1849]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF84FF83FF584D31852AC041" box="[151,273,1828,1849]" italics="true" pageId="7" pageNumber="42">D. punctata</emphasis>
</taxonomicName>
, lectotype (ZMMU) from Hawaii instead of China, photographed by Dr. Andrey L. Ozerov, copyright ZMMU, published with permission; 
<emphasis id="57FDEA98FF84FF83FE754D5785C1C02F" bold="true" box="[442,506,1858,1879]" pageId="7" pageNumber="42">22–24</emphasis>
<taxonomicName id="A2894D09FF84FF83FDCB4D568642C020" box="[516,633,1858,1880]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="elliptica" status="sp. nov.">
<emphasis id="57FDEA98FF84FF83FDCB4D568642C020" box="[516,633,1858,1880]" italics="true" pageId="7" pageNumber="42">D. elliptica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF84FF83FD4C4D5786FBC02F" box="[643,704,1858,1879]" pageId="7" pageNumber="42" rank="species">
<emphasis id="57FDEA98FF84FF83FD4C4D5786FBC02F" bold="true" box="[643,704,1858,1879]" pageId="7" pageNumber="42">sp. n.</emphasis>
</taxonomicNameLabel>
; 
<emphasis id="57FDEA98FF84FF83FD1E4D57872BC02F" bold="true" box="[721,784,1858,1879]" pageId="7" pageNumber="42">22–23</emphasis>
male holotype from Yunnan, Cangyuan, Banhong; 
<emphasis id="57FDEA98FF84FF83FAFB4D578176C02F" bold="true" box="[1332,1357,1858,1879]" pageId="7" pageNumber="42">24</emphasis>
female paratype from Yunnan, Mengla, Mohan; 
<emphasis id="57FDEA98FF84FF83FD894D7586BEC00D" bold="true" box="[582,645,1888,1909]" pageId="7" pageNumber="42">25–26</emphasis>
<taxonomicName id="A2894D09FF84FF83FD5F4D7486C3C00E" box="[656,760,1889,1910]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="naevus" status="sp. nov.">
<emphasis id="57FDEA98FF84FF83FD5F4D7486C3C00E" box="[656,760,1889,1910]" italics="true" pageId="7" pageNumber="42">D. naevus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF84FF83FCCC4D758704C00D" box="[771,831,1888,1909]" pageId="7" pageNumber="42" rank="species">
<emphasis id="57FDEA98FF84FF83FCCC4D758704C00D" bold="true" box="[771,831,1888,1909]" pageId="7" pageNumber="42">sp. n.</emphasis>
</taxonomicNameLabel>
, male holotype from Xizang, Motuo, Beibeng; 
<emphasis id="57FDEA98FF84FF83FAF54D758142C00D" bold="true" box="[1338,1401,1888,1909]" pageId="7" pageNumber="42">27–28</emphasis>
<taxonomicName id="A2894D09FF84FF83FA4C4D74857DC0ED" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="nigrescens" status="subsp. nov." subSpecies="guani">
<emphasis id="57FDEA98FF84FF83FA4C4D74857DC0ED" italics="true" pageId="7" pageNumber="42">D. nigrescens guani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF84FF83FE9E4D6A858FC0EC" box="[337,436,1919,1940]" pageId="7" pageNumber="42" rank="subSpecies">
<emphasis id="57FDEA98FF84FF83FE9E4D6A858FC0EC" bold="true" box="[337,436,1919,1940]" pageId="7" pageNumber="42">subsp. n.</emphasis>
</taxonomicNameLabel>
, male holotype from Guangxi, Chongzuo, Daqingshan; 
<emphasis id="57FDEA98FF84FF83FBC34D6A8070C0EC" bold="true" box="[1036,1099,1919,1940]" pageId="7" pageNumber="42">29–30</emphasis>
<taxonomicName id="A2894D09FF84FF83FB9A4D958162C0ED" box="[1109,1369,1920,1941]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="42" phylum="Arthropoda" rank="subSpecies" species="nigrescens" subSpecies="nigrescens">
<emphasis id="57FDEA98FF84FF83FB9A4D958162C0ED" box="[1109,1369,1920,1941]" italics="true" pageId="7" pageNumber="42">D. nigrescens nigrescens</emphasis>
</taxonomicName>
, male syntype (NTU) from Taiwan, photographed by Yi-Ho Liu and downloaded from NTU website, copyright Insect Museum Digital Archives Project, Department of Entomology, National Taiwan University. (Scale bars = 5mm.)
</paragraph>
</caption>
<subSubSection id="2D936501FF8BFF8CFF084A8281BEC3D8" pageId="8" pageNumber="43" type="discussion">
<paragraph id="6536368AFF8BFF8CFF084A82877BC61D" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="43">
<emphasis id="57FDEA98FF8BFF8CFF084A828500C7C8" bold="true" box="[199,315,151,176]" pageId="8" pageNumber="43">Remarks.</emphasis>
<taxonomicName id="A2894D09FF8BFF8CFE894A8C85F4C7C8" box="[326,463,153,176]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF8BFF8CFE894A8C85F4C7C8" box="[326,463,153,176]" italics="true" pageId="8" pageNumber="43">D. punctata</emphasis>
</taxonomicName>
has become commonly recognized by insect physiologists and pet-keepers, and is called Pacific beetle cockroach, but its natural history in the wild remains mysterious to our knowledge, aside from its harm to Pinales in Hawaii (
<figureCitation id="FDB22A0FFF8BFF8CFE224ACA8605C780" box="[493,574,223,248]" captionStart="FIGURES 54 – 56. 54 D" captionStartId="13.[151,264,1787,1810]" captionTargetBox="[151,1436,193,1766]" captionTargetId="figure@13.[151,1436,193,1766]" captionTargetPageId="13" captionText="FIGURES 54 – 56. 54 D. punctata eating lemon peel in Hawaii, 2006. XI. 29, image by Forest &amp; Kim Starr (University of Hawaii); 55 Cypress with damage by D. punctata in Hawaii, 2005. VII. 11, image by Forest &amp; Kim Starr (University of Hawaii); 56 D. nigrescens nigrescens Shiraki, 1931 from Taiwan, Chiayi County, Tashan (= Xiaotashan), about 1200 m, photographed by Gaga (Taiwan). The insect was found near a light trap but not on the curtain." httpUri="https://zenodo.org/record/289456/files/figure.png" pageId="8" pageNumber="43">
Fig. 
<emphasis id="57FDEA98FF8BFF8CFDED4ACA8605C780" bold="true" box="[546,574,223,248]" pageId="8" pageNumber="43">55</emphasis>
</figureCitation>
). They gnaw the bark of branches, often completely girdling them, resulting in yellow or dead foliage (
<bibRefCitation id="01184B7BFF8BFF8CFE004B118652C664" author="Hebard, M." box="[463,617,260,285]" firstAuthor="Hebard" journalOrPublisher="Occasional Papers of the Bernice Pauahi Bishop Museum of Polynesian Ethnology and Natural History" pageId="8" pageNumber="43" pagination="305 - 379" part="7" refId="ref12580" refString="Hebard, M. (1922) The Dermaptera and Orthoptera of Hawaii. Occasional Papers of the Bernice Pauahi Bishop Museum of Polynesian Ethnology and Natural History, 7 (14), 305 - 379." title="The Dermaptera and Orthoptera of Hawaii" type="journal article" year="1922">Hebard 1922</bibRefCitation>
; 
<bibRefCitation id="01184B7BFF8BFF8CFDB64B118756C664" author="Fullaway, D. T. &amp; Tagawa, T. K. &amp; Trujillo, E. &amp; Davis, C. J. &amp; Pung, E." bookContentInfo="16 pp." box="[633,877,260,285]" etAl="et al." firstAuthor="Fullaway" journalOrPublisher="University of Hawaii, Honolulu" pageId="8" pageNumber="43" refId="ref12398" refString="Fullaway, D. T., Tagawa, T. K., Trujillo, E., Davis, C. J. &amp; Pung, E. (1972) Norfolk Island Pine Culture. University of Hawaii, Honolulu, 16 pp." title="Norfolk Island Pine Culture" type="book" year="1972">
Fullaway 
<emphasis id="57FDEA98FF8BFF8CFD244B10871CC664" box="[747,807,260,284]" italics="true" pageId="8" pageNumber="43">et al.</emphasis>
1972
</bibRefCitation>
). Among the materials from 
<collectingCountry id="1D9E761AFF8BFF8CFB0D4B118133C665" box="[1218,1288,260,285]" name="China" pageId="8" pageNumber="43">China</collectingCountry>
, most of the specimens from Yunnan were collected via canopy fogging. This may indicate that the Yunnan population of 
<taxonomicName id="A2894D09FF8BFF8CFA4E4B3C84C2C61C" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF8BFF8CFA4E4B3C84C2C61C" italics="true" pageId="8" pageNumber="43">D. punctata</emphasis>
</taxonomicName>
is a canopy dweller, or at least active in the canopy.
</paragraph>
<paragraph id="6536368AFF8BFF8CFF084B7A8686C6D5" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="43">
This species also exhibits possible positive phototaxis since they can be found at light in the evening (
<bibRefCitation id="01184B7BFF8BFF8CFA8D4B6584E8C6D4" author="Browne, A. C." firstAuthor="Browne" journalOrPublisher="Proceedings of the Hawaiian Entomological Society" pageId="8" pageNumber="43" pagination="151 - 152" part="11" refId="ref12198" refString="Browne, A. C. (1942) Insects taken at light at Kalawahine Place, Honolulu. Proceedings of the Hawaiian Entomological Society, 11 (02), 151 - 152." title="Insects taken at light at Kalawahine Place, Honolulu" type="journal article" year="1942">Browne 1942</bibRefCitation>
; Jianyue Qiu’s personal observation, 2015).
</paragraph>
<paragraph id="6536368AFF8BFF8CFF084BAD8754C4DD" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="43">
In 
<date id="1137104AFF8BFF8CFF254BAD85A7C6A8" box="[234,412,439,464]" pageId="8" pageNumber="43" value="2003" valueMax="2004">2003 and 2004</date>
, some 
<taxonomicName id="A2894D09FF8BFF8CFE3C4BAD8655C6A8" box="[499,622,440,464]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF8BFF8CFE3C4BAD8655C6A8" box="[499,622,440,464]" italics="true" pageId="8" pageNumber="43">Diploptera</emphasis>
</taxonomicName>
sp. which look almost the same as 
<taxonomicName id="A2894D09FF8BFF8CFBD14BAC809CC6A8" box="[1054,1191,441,464]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF8BFF8CFBD14BAC809CC6A8" box="[1054,1191,441,464]" italics="true" pageId="8" pageNumber="43">D. punctata</emphasis>
</taxonomicName>
were collected from Nakornpathom Province in 
<collectingCountry id="1D9E761AFF8BFF8CFE1F4BC9860FC68D" box="[464,564,476,501]" name="Thailand" pageId="8" pageNumber="43">Thailand</collectingCountry>
(see asterisk in 
<figureCitation id="FDB22A0FFF8BFF8CFD224BC98714C68D" box="[749,815,476,501]" captionStart="FIGURE 1" captionStartId="5.[151,250,1807,1829]" captionTargetBox="[151,1436,195,1782]" captionTargetId="figure@5.[151,1436,195,1783]" captionTargetPageId="5" captionText="FIGURE 1. Distribution of Diploptera in China and Southeast Asia. The precise localities are painted black, whilst the imprecise ones are in white; each type locality is marked with a ring; the (sub) species are indicated by numbers in alphabetical order: 1 D. bicolor, 2 D. elliptica sp. n., 3 D. erythrocephala, 4 D. maculata, 5 D. minor, 6 D. naevus sp. n., 7 D. nigrescens guani subsp. n., 8 D. nigrescens nigrescens, 9 D. parva, × D. punctata; asterisk stands for the habitat of the Thai D. punctata (see the text)." httpUri="https://zenodo.org/record/289449/files/figure.png" pageId="8" pageNumber="43">
Fig. 
<emphasis id="57FDEA98FF8BFF8CFCEE4BC98714C68D" bold="true" box="[801,815,476,501]" pageId="8" pageNumber="43">1</emphasis>
</figureCitation>
). These insects were found mainly on trees, especially coconut trees, in the grooves between the stalk of the leaves and the trunk; some were also on the wall in the bright light of fluorescence lamp. Subsequently Dr. Stephen S. Tobe’s lab (University of Toronto) discovered that they seemed to be reproductively isolated from the lab population of 
<taxonomicName id="A2894D09FF8BFF8CFC53485C801DC518" box="[924,1062,585,608]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF8BFF8CFC53485C801DC518" box="[924,1062,585,608]" italics="true" pageId="8" pageNumber="43">D. punctata</emphasis>
</taxonomicName>
, which originated from Hawaii (personal communications with Witoon Wattananit (Silpakorn University) and Dr. Stephen S. Tobe, 2014). Dr. Tobe’s lab went on the studies on both the Hawaiian and 
<collectingCountry id="1D9E761AFF8BFF8CFCF1489A874AC5D0" box="[830,881,655,680]" name="Thailand" pageId="8" pageNumber="43">Thai</collectingCountry>
populations of 
<taxonomicName id="A2894D09FF8BFF8CFBE44885809DC5D0" box="[1067,1190,656,680]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF8BFF8CFBE44885809DC5D0" box="[1067,1190,656,680]" italics="true" pageId="8" pageNumber="43">Diploptera</emphasis>
</taxonomicName>
and performed cross experiments to probe into their relationship; 
<bibRefCitation id="01184B7BFF8BFF8CFD6A48A187ABC5B4" author="Lenkic, L. E. &amp; Wolfe, J. M. &amp; Chang, B. S. W. &amp; Tobe, S. S." box="[677,912,692,717]" etAl="et al." firstAuthor="Lenkic" journalOrPublisher="Journal of Insect Physiology" pageId="8" pageNumber="43" pagination="931 - 938" part="54" refId="ref12752" refString="Lenkic, L. E., Wolfe, J. M., Chang, B. S. W. &amp; Tobe, S. S. (2008) Endocrine and reproductive differences and genetic divergence in two populations of the cockroach Diploptera punctata. Journal of Insect Physiology, 54, 931 - 938. http: // dx. doi. org / 10.1016 / j. jinsphys. 2008.02.009" title="Endocrine and reproductive differences and genetic divergence in two populations of the cockroach Diploptera punctata" type="journal article" year="2008">
Lenkic 
<emphasis id="57FDEA98FF8BFF8CFD3248A08701C5B4" box="[765,826,692,716]" italics="true" pageId="8" pageNumber="43">et al.</emphasis>
(2008)
</bibRefCitation>
found that the two populations, which were recognized as conspecific in their paper, showed a capacity of two-way breeding but a hybrid breakdown had arisen. However, reproductive compatibility and isolation must be affected by the laboratory condition and thus a long-term lab population cannot fully represent the species in the wild. Considering that the 
<collectingCountry id="1D9E761AFF8BFF8CFB0D490A80CDC440" box="[1218,1270,799,824]" name="Thailand" pageId="8" pageNumber="43">Thai</collectingCountry>
and Hawaiian populations are interfertile and morphologically almost identical, we propose a continuance of the views by previous taxonomists to regard these two populations, as well as those from other localities including 
<collectingCountry id="1D9E761AFF8BFF8CFAC14972816FC4F8" box="[1294,1364,871,896]" name="China" pageId="8" pageNumber="43">China</collectingCountry>
, Indo- 
<collectingCountry id="1D9E761AFF8BFF8CFF58499984E6C4DD" box="[151,221,908,933]" name="China" pageId="8" pageNumber="43">China</collectingCountry>
, Malay Archipelago and Pacific Islands, to be conspecific.
</paragraph>
<paragraph id="6536368AFF8BFF8CFF0849A581BEC3D8" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="43">
On the other hand, 
<bibRefCitation id="01184B7BFF8BFF8CFE6449BA86A8C4B0" author="Lenkic, L. E. &amp; Wolfe, J. M. &amp; Chang, B. S. W. &amp; Tobe, S. S." box="[427,659,943,968]" etAl="et al." firstAuthor="Lenkic" journalOrPublisher="Journal of Insect Physiology" pageId="8" pageNumber="43" pagination="931 - 938" part="54" refId="ref12752" refString="Lenkic, L. E., Wolfe, J. M., Chang, B. S. W. &amp; Tobe, S. S. (2008) Endocrine and reproductive differences and genetic divergence in two populations of the cockroach Diploptera punctata. Journal of Insect Physiology, 54, 931 - 938. http: // dx. doi. org / 10.1016 / j. jinsphys. 2008.02.009" title="Endocrine and reproductive differences and genetic divergence in two populations of the cockroach Diploptera punctata" type="journal article" year="2008">
Lenkic 
<emphasis id="57FDEA98FF8BFF8CFDCC49A48604C4B0" box="[515,575,944,968]" italics="true" pageId="8" pageNumber="43">et al.</emphasis>
(2008)
</bibRefCitation>
also found the genetic divergence between the 
<collectingCountry id="1D9E761AFF8BFF8CFB0F49BA80CFC4B0" box="[1216,1268,943,968]" name="Thailand" pageId="8" pageNumber="43">Thai</collectingCountry>
and Hawaiian populations in a fragment of the mitochondrial 16S rRNA gene to be as surprisingly large as 8.6% (or 11.7%, when corrected by PAUP* under GTR+G model), which could even indicate a distinct interspecific divergence rather than that between conspecific populations. Even if the two populations represent naturally alien species to each other, the divergence is too large between sibling species. 
<bibRefCitation id="01184B7BFF8BFF8CFCD04E2A87C7C320" author="Lenkic, L. E. &amp; Wolfe, J. M. &amp; Chang, B. S. W. &amp; Tobe, S. S." box="[799,1020,1087,1112]" etAl="et al." firstAuthor="Lenkic" journalOrPublisher="Journal of Insect Physiology" pageId="8" pageNumber="43" pagination="931 - 938" part="54" refId="ref12752" refString="Lenkic, L. E., Wolfe, J. M., Chang, B. S. W. &amp; Tobe, S. S. (2008) Endocrine and reproductive differences and genetic divergence in two populations of the cockroach Diploptera punctata. Journal of Insect Physiology, 54, 931 - 938. http: // dx. doi. org / 10.1016 / j. jinsphys. 2008.02.009" title="Endocrine and reproductive differences and genetic divergence in two populations of the cockroach Diploptera punctata" type="journal article" year="2008">
Lenkic 
<emphasis id="57FDEA98FF8BFF8CFCBC4E548790C320" box="[883,939,1088,1112]" italics="true" pageId="8" pageNumber="43">et al.</emphasis>
(2008)
</bibRefCitation>
speculated that the populations of 
<taxonomicName id="A2894D09FF8BFF8CFA4E4E5484C2C304" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF8BFF8CFA4E4E5484C2C304" italics="true" pageId="8" pageNumber="43">D. punctata</emphasis>
</taxonomicName>
are in the process of speciation. We just agree with them and, it must be interesting and significant to perform further research on 
<taxonomicName id="A2894D09FF8BFF8CFE1D4E9C866DC3D8" box="[466,598,1161,1184]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF8BFF8CFE1D4E9C866DC3D8" box="[466,598,1161,1184]" italics="true" pageId="8" pageNumber="43">D. punctata</emphasis>
</taxonomicName>
, which is a magic model species in insect physiology and biogeography.
</paragraph>
</subSubSection>
<subSubSection id="2D936501FF8BFF8CFF084EB9806AC0C1" pageId="8" pageNumber="43" type="materials_examined">
<paragraph id="6536368AFF8BFF8CFF084EB98785C200" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="43">
<emphasis id="57FDEA98FF8BFF8CFF084EB98629C3BD" bold="true" box="[199,530,1196,1221]" pageId="8" pageNumber="43">
<typeStatus id="BA328828FF8BFF8CFF084EB98539C3BD" box="[199,258,1196,1221]" pageId="8" pageNumber="43">Type</typeStatus>
materials examined.
</emphasis>
<materialsCitation id="D5E13CD7FF8BFF8CFDEA4EB98799C22D" ID-GBIF-Occurrence="3923978301" pageId="8" pageNumber="43" typeStatus="lectotype">
<typeStatus id="BA328828FF8BFF8CFDEA4EB986FEC3BC" box="[549,709,1196,1220]" pageId="8" pageNumber="43" type="lectotype">LECTOTYPE</typeStatus>
(
<collectionCode id="0398AE4FFF8BFF8CFD104EB98702C3BC" box="[735,825,1196,1220]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="8" pageNumber="43">ZMMU</collectionCode>
) of 
<taxonomicName id="A2894D09FF8BFF8CFCB04EB98003C3BC" box="[895,1080,1196,1220]" class="Insecta" family="Blattidae" genus="Blatta" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF8BFF8CFCB04EB98003C3BC" box="[895,1080,1196,1220]" italics="true" pageId="8" pageNumber="43">Blatta punctata</emphasis>
</taxonomicName>
designated here, with label “ 
<typeStatus id="BA328828FF8BFF8CFF6F4EC58579C390" box="[160,322,1232,1256]" pageId="8" pageNumber="43" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="A2894D09FF8BFF8CFE804EC586E6C390" authority="Eschscholtz, 1822" authorityName="Eschscholtz" authorityYear="1822" box="[335,733,1231,1256]" class="Insecta" family="Blattidae" genus="Blatta" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF8BFF8CFE804EC5863AC390" box="[335,513,1232,1256]" italics="true" pageId="8" pageNumber="43">Blatta punctata</emphasis>
<bibRefCitation id="01184B7BFF8BFF8CFDC34EDA86E6C390" author="Eschscholtz, J. F. von" bookContentInfo="128 pp." box="[524,733,1231,1256]" firstAuthor="Eschscholtz" journalOrPublisher="G. Reimer, Berlin" pageId="8" pageNumber="43" refId="ref12361" refString="Eschscholtz, J. F. von (1822) Entomographien. G. Reimer, Berlin, 128 pp. http: // dx. doi. org / 10.5962 / bhl. title. 5121" title="Entomographien" type="book" year="1822">Eschscholtz, 1822</bibRefCitation>
</taxonomicName>
des. Conlin McCat 2014”, sex unknown, “I. Sandwich” (= Inseln Sandwich = Sandwich Islands, now Hawaiian Islands), also a determination label “ 
<taxonomicName id="A2894D09FF8BFF8CFB1D4EE1816DC274" box="[1234,1366,1268,1292]" class="Insecta" family="Blattidae" genus="Periplaneta" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF8BFF8CFB1D4EE1816DC274" box="[1234,1366,1268,1292]" italics="true" pageId="8" pageNumber="43">Periplaneta</emphasis>
</taxonomicName>
sp.?” [
<emphasis id="57FDEA98FF8BFF8CFF6F4F0D8512C248" box="[160,297,1304,1328]" italics="true" pageId="8" pageNumber="43">special note</emphasis>
: Conlin McCat is identical with the author Xinran Li. The author uses the former in daily life and communications but his institute forces the use of the latter on him];
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFC644F298785C200" ID-GBIF-Occurrence="3923978322" collectionCode="MHNG" pageId="8" pageNumber="43" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
<typeStatus id="BA328828FF8BFF8CFC644F298012C22C" box="[939,1065,1340,1364]" pageId="8" pageNumber="43" type="syntype">SYNTYPE</typeStatus>
(
<collectionCode id="0398AE4FFF8BFF8CFBF64F2980A9C22C" box="[1081,1170,1340,1364]" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="8" pageNumber="43">MHNG</collectionCode>
) of 
<taxonomicName id="A2894D09FF8BFF8CFB0D4F2981AFC22C" box="[1218,1428,1340,1364]" class="Insecta" family="Ectobiidae" genus="Prosoplecta" kingdom="Animalia" order="Blattodea" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="silpha">
<emphasis id="57FDEA98FF8BFF8CFB0D4F2981AFC22C" box="[1218,1428,1340,1364]" italics="true" pageId="8" pageNumber="43">
<originalDetermination id="46978E76FF8BFF8CFB0D4F2981AFC22C" box="[1218,1428,1340,1364]" pageId="8" pageNumber="43">Prosoplecta silpha</originalDetermination>
</emphasis>
</taxonomicName>
, 
<specimenCount id="738FFD03FF8BFF8CFF584F4A84EBC200" box="[151,208,1375,1400]" pageId="8" pageNumber="43" type="male">male</specimenCount>
, “N. Holl.” (= Nouvelle-Hollande = New Holland, now Australia).
</materialsCitation>
</paragraph>
<paragraph id="6536368AFF8BFF8CFF084F91806AC0C1" blockId="8.[151,1437,151,1977]" pageId="8" pageNumber="43">
<emphasis id="57FDEA98FF8BFF8CFF084F918634C2E5" bold="true" box="[199,527,1412,1437]" pageId="8" pageNumber="43">Other materials examined.</emphasis>
<materialsCitation id="D5E13CD7FF8BFF8CFDD44F918630C2B9" ID-GBIF-Occurrence="3923978315" pageId="8" pageNumber="43">
<emphasis id="57FDEA98FF8BFF8CFDD44F9186E6C2E5" bold="true" box="[539,733,1412,1437]" pageId="8" pageNumber="43">
<collectingCountry id="1D9E761AFF8BFF8CFDD44F91865DC2E5" box="[539,614,1412,1437]" name="China" pageId="8" pageNumber="43">China</collectingCountry>
, Yunnan:
</emphasis>
<specimenCount id="738FFD03FF8BFF8CFD264F918775C2E5" box="[745,846,1412,1437]" pageId="8" pageNumber="43" type="female">1 female</specimenCount>
(
<collectionCode id="0398AE4FFF8BFF8CFCAE4F918784C2E4" box="[865,959,1412,1436]" pageId="8" pageNumber="43">IESWU</collectionCode>
), Xishuangbanna, Xiaomengyang, 
<quantity id="A2719B6FFF8BFF8CFA9A4F9181A3C2E5" box="[1365,1432,1412,1437]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="8" pageNumber="43" unit="m" value="850.0">850m</quantity>
, 1957. 
<date id="1137104AFF8BFF8CFF164FB2853AC2B8" box="[217,257,1447,1472]" pageId="8" pageNumber="43">X.8</date>
, coll. Shu-Yong Wang;
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFDDA4FBD8561C29D" ID-GBIF-Occurrence="3923978304" pageId="8" pageNumber="43">
1 sex unknown (
<collectionCode id="0398AE4FFF8BFF8CFD1C4FBD870AC2B8" box="[723,817,1448,1472]" pageId="8" pageNumber="43">IESWU</collectionCode>
), Xishuangbanna, Damenglong, 
<quantity id="A2719B6FFF8BFF8CFB694FB280D2C2B8" box="[1190,1257,1447,1472]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="43" unit="m" value="650.0">650m</quantity>
, 1958. 
<date id="1137104AFF8BFF8CFAF94FB28166C2B8" box="[1334,1373,1447,1472]" pageId="8" pageNumber="43">V.5</date>
, coll. Shu-Yong Wang;
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFEAA4FD981B8C29D" ID-GBIF-Occurrence="3923978333" box="[357,1411,1484,1509]" pageId="8" pageNumber="43">
<specimenCount id="738FFD03FF8BFF8CFEAA4FD985F0C29D" box="[357,459,1484,1509]" pageId="8" pageNumber="43" type="female">1 female</specimenCount>
(IOZCAS), Xishuangbanna, Mengla, 
<quantity id="A2719B6FFF8BFF8CFC484FD98039C29D" box="[903,1026,1484,1509]" metricMagnitude="2" metricUnit="m" metricValue="6.35" metricValueMax="6.5" metricValueMin="6.2" pageId="8" pageNumber="43" unit="m" value="635.0" valueMax="650.0" valueMin="620.0">620–650m</quantity>
, 1959. 
<date id="1137104AFF8BFF8CFB9D4FD980AAC29D" box="[1106,1169,1484,1509]" pageId="8" pageNumber="43" value="2011-06">VI.11</date>
, coll. Fa-Cai Zhang;
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFA414FD98559C170" ID-GBIF-Occurrence="3923978343" pageId="8" pageNumber="43">
<specimenCount id="738FFD03FF8BFF8CFA414FD984D5C170" pageId="8" pageNumber="43" type="female">3 females</specimenCount>
(
<collectionCode id="0398AE4FFF8BFF8CFF324FE5856DC170" box="[253,342,1520,1544]" pageId="8" pageNumber="43">MHBU</collectionCode>
),
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFEA54FFA86D7C154" ID-GBIF-Occurrence="3923978332" pageId="8" pageNumber="43">
Mengla, Wangtianshu, 2007. 
<date id="1137104AFF8BFF8CFD624FFA872AC170" box="[685,785,1519,1544]" pageId="8" pageNumber="43">VIII.6–7</date>
, coll. Guo-Dong Ren, Wen-Jun Hou, Ya-Lin Li; 
<specimenCount id="738FFD03FF8BFF8CFAFB4FE58144C170" box="[1332,1407,1519,1544]" pageId="8" pageNumber="43" type="male">1 male</specimenCount>
&amp; 
<specimenCount id="738FFD03FF8BFF8CFF584C0184C7C155" box="[151,252,1556,1581]" pageId="8" pageNumber="43" type="female">1 female</specimenCount>
(
<figureCitation id="FDB22A0FFF8BFF8CFEDF4C0185A7C154" box="[272,412,1556,1581]" captionStart="FIGURES 17 – 30" captionStartId="7.[151,264,1796,1819]" captionTargetBox="[191,1384,195,1758]" captionTargetId="figure@7.[191,1385,195,1760]" captionTargetPageId="7" captionText="FIGURES 17 – 30. Diploptera spp. &amp; subspp. distributed in China. 17 – 18 D. punctata, female from Yunnan, Menglun; 19 – 21 D. punctata, lectotype (ZMMU) from Hawaii instead of China, photographed by Dr. Andrey L. Ozerov, copyright ZMMU, published with permission; 22 – 24 D. elliptica sp. n.; 22 – 23 male holotype from Yunnan, Cangyuan, Banhong; 24 female paratype from Yunnan, Mengla, Mohan; 25 – 26 D. naevus sp. n., male holotype from Xizang, Motuo, Beibeng; 27 – 28 D. nigrescens guani subsp. n., male holotype from Guangxi, Chongzuo, Daqingshan; 29 – 30 D. nigrescens nigrescens, male syntype (NTU) from Taiwan, photographed by Yi-Ho Liu and downloaded from NTU website, copyright Insect Museum Digital Archives Project, Department of Entomology, National Taiwan University. (Scale bars = 5 mm.)" httpUri="https://zenodo.org/record/289451/files/figure.png" pageId="8" pageNumber="43">Figs. 17–18</figureCitation>
, 
<figureCitation id="FDB22A0FFF8BFF8CFE634C0185F2C154" box="[428,457,1556,1580]" captionStart="FIGURES 31 – 36. D" captionStartId="9.[151,264,1815,1838]" captionTargetBox="[151,1433,198,1789]" captionTargetId="figure@9.[151,1436,195,1791]" captionTargetPageId="9" captionText="FIGURES 31 – 36. D. punctata (Eschscholtz, 1822) from China, Yunnan, Menglun. 31 hind wing, female, with grey coloration indicating sclerotization; 32 – 36 male; 32 terminal segment of dorsal abdomen, in ventral view; 33 subgenital plate in dorsal view; 34 – 36 phallic sclerites in dorsal view; 34 sclerites R’; 35 sclerites L 2 ’; 36 apex of left complex hook, showing sclerite L 3 ’. (ce. = cercus, m. = membranous, p. p. = paraprocts, st. = stylus; other abbreviations as indicated in the text. Scale bars = 10 mm for 31, 1 mm for 32 &amp; 33, 500 µm for 34 – 36.)" httpUri="https://zenodo.org/record/289452/files/figure.png" pageId="8" pageNumber="43">31</figureCitation>
) (
<collectionCode id="0398AE4FFF8BFF8CFE2B4C01867AC154" box="[484,577,1556,1580]" pageId="8" pageNumber="43">IESWU</collectionCode>
, ex IOZCAS),
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFD384C018726C128" ID-GBIF-Occurrence="3923978336" pageId="8" pageNumber="43">
Menglun, 
<taxonomicName id="A2894D09FF8BFF8CFCA14C018055C154" box="[878,1134,1556,1580]" class="Magnoliopsida" family="Combretaceae" genus="Anogeissus" kingdom="Plantae" order="Myrtales" pageId="8" pageNumber="43" phylum="Tracheophyta" rank="species" species="acuminata">
<emphasis id="57FDEA98FF8BFF8CFCA14C018055C154" box="[878,1134,1556,1580]" italics="true" pageId="8" pageNumber="43">Anogeissus acuminata</emphasis>
</taxonomicName>
plantation (about 20 yr.), 21°53.993′, 101°16.810′, 2007. 
<date id="1137104AFF8BFF8CFE354C2D8676C128" box="[506,589,1592,1616]" pageId="8" pageNumber="43" value="1919-08">VIII.19</date>
, coll. Guo Zheng;
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFCE94C2D811CC10C" ID-GBIF-Occurrence="3923978302" pageId="8" pageNumber="43">
<specimenCount id="738FFD03FF8BFF8CFCE94C2D8749C128" box="[806,882,1591,1616]" pageId="8" pageNumber="43" type="male">1 male</specimenCount>
(
<figureCitation id="FDB22A0FFF8BFF8CFC4B4C228037C128" box="[900,1036,1591,1616]" captionStart="FIGURES 31 – 36. D" captionStartId="9.[151,264,1815,1838]" captionTargetBox="[151,1433,198,1789]" captionTargetId="figure@9.[151,1436,195,1791]" captionTargetPageId="9" captionText="FIGURES 31 – 36. D. punctata (Eschscholtz, 1822) from China, Yunnan, Menglun. 31 hind wing, female, with grey coloration indicating sclerotization; 32 – 36 male; 32 terminal segment of dorsal abdomen, in ventral view; 33 subgenital plate in dorsal view; 34 – 36 phallic sclerites in dorsal view; 34 sclerites R’; 35 sclerites L 2 ’; 36 apex of left complex hook, showing sclerite L 3 ’. (ce. = cercus, m. = membranous, p. p. = paraprocts, st. = stylus; other abbreviations as indicated in the text. Scale bars = 10 mm for 31, 1 mm for 32 &amp; 33, 500 µm for 34 – 36.)" httpUri="https://zenodo.org/record/289452/files/figure.png" pageId="8" pageNumber="43">Figs. 32–36</figureCitation>
) (
<collectionCode id="0398AE4FFF8BFF8CFBEB4C2D80BAC128" box="[1060,1153,1592,1616]" pageId="8" pageNumber="43">IESWU</collectionCode>
, ex IOZCAS), Menglun, G213 (national road), 
<quantity id="A2719B6FFF8BFF8CFE6D4C4985DEC10D" box="[418,485,1628,1653]" metricMagnitude="2" metricUnit="m" metricValue="5.9399999999999995" pageId="8" pageNumber="43" unit="m" value="594.0">594m</quantity>
, 21°53.794′, 101°17.152′, 2009. 
<date id="1137104AFF8BFF8CFCAD4C498798C10C" box="[866,931,1628,1652]" pageId="8" pageNumber="43" value="1927-11">XI.27</date>
, coll. Guo Tang, Zhi-Yuan Yao;
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFAFA4C4987C7C1E0" ID-GBIF-Occurrence="3923978325" pageId="8" pageNumber="43">
<specimenCount id="738FFD03FF8BFF8CFAFA4C4981A7C10D" box="[1333,1436,1628,1653]" pageId="8" pageNumber="43" type="female">1 female</specimenCount>
(
<collectionCode id="0398AE4FFF8BFF8CFF504C9584C6C1E0" box="[159,253,1664,1688]" pageId="8" pageNumber="43">IESWU</collectionCode>
), Mengla, Shangyong, Longmencun, 2015. 
<date id="1137104AFF8BFF8CFD214C6A872EC1E0" box="[750,789,1663,1688]" pageId="8" pageNumber="43">V.9</date>
, coll. Jian-Yue Qiu.
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFBCA4C6A8706C1C5" ID-GBIF-Occurrence="3923978306" pageId="8" pageNumber="43">
<emphasis id="57FDEA98FF8BFF8CFBCA4C6A80F4C1E0" bold="true" box="[1029,1231,1663,1688]" pageId="8" pageNumber="43">
<collectingCountry id="1D9E761AFF8BFF8CFBCA4C6A806BC1E0" box="[1029,1104,1663,1688]" name="China" pageId="8" pageNumber="43">China</collectingCountry>
, Guangxi:
</emphasis>
<specimenCount id="738FFD03FF8BFF8CFB164C958106C1E0" box="[1241,1341,1663,1688]" pageId="8" pageNumber="43" type="female">1 female</specimenCount>
(
<collectionCode id="0398AE4FFF8BFF8CFA804C9581B4C1E0" LSID="urn:lsid:biocol.org:col:33074" box="[1359,1423,1664,1688]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="8" pageNumber="43">CAU</collectionCode>
), Longzhou, Nonggang, 
<quantity id="A2719B6FFF8BFF8CFE554CB185E7C1C4" box="[410,476,1700,1724]" metricMagnitude="2" metricUnit="m" metricValue="2.4" pageId="8" pageNumber="43" unit="m" value="240.0">240m</quantity>
, 1982. 
<date id="1137104AFF8BFF8CFDE74CB18666C1C4" box="[552,605,1700,1724]" pageId="8" pageNumber="43" value="1918-05">V.18</date>
, coll. Xin-Li Wang;
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFC8B4CB186BEC198" ID-GBIF-Occurrence="3923978303" pageId="8" pageNumber="43">
<specimenCount id="738FFD03FF8BFF8CFC8B4CB1879EC1C5" box="[836,933,1700,1725]" pageId="8" pageNumber="43" type="female">1 female</specimenCount>
(
<collectionCode id="0398AE4FFF8BFF8CFC7A4CB18037C1C4" box="[949,1036,1700,1724]" pageId="8" pageNumber="43">ZSCIQ</collectionCode>
), Shangsi, Shiwandashan (=Shiwan Mountains), 2014. 
<date id="1137104AFF8BFF8CFEA14CDD85A9C198" box="[366,402,1736,1760]" pageId="8" pageNumber="43">V.1</date>
, coll. Ke-Liang Wu;
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFD5E4CDD8762C07D" ID-GBIF-Occurrence="3923978340" pageId="8" pageNumber="43">
<specimenCount id="738FFD03FF8BFF8CFD5E4CDD86DAC198" box="[657,737,1735,1760]" pageId="8" pageNumber="43" type="male">1 male</specimenCount>
(
<figureCitation id="FDB22A0FFF8BFF8CFD394CD28777C198" box="[758,844,1735,1760]" captionStart="FIGURE 80. A" captionStartId="19.[151,250,1644,1666]" captionTargetBox="[151,1435,658,1622]" captionTargetId="figure@19.[151,1435,658,1622]" captionTargetPageId="19" captionText="FIGURE 80. A male D. punctata in repose near light trap. Photographed by Lu Qiu in Nonggang National Nature Reserve in Guangxi, China. We are very lucky to get this timely photo just before the present paper is published. This is a light-colored individual of the species. Its large abdomen was retracted to be wholly covered by the tegmina when it became a specimen." httpUri="https://zenodo.org/record/289461/files/figure.png" pageId="8" pageNumber="43">Fig. 80</figureCitation>
) (
<collectionCode id="0398AE4FFF8BFF8CFCA74CDD87FDC198" box="[872,966,1736,1760]" pageId="8" pageNumber="43">IESWU</collectionCode>
), Longzhou, Nonggang National Nature Reserve, light trap, 2015. 
<date id="1137104AFF8BFF8CFE074CF98631C07D" box="[456,522,1772,1797]" pageId="8" pageNumber="43" value="1929-06">VI.29</date>
, coll. Lu Qiu, Qi-Kun Bai;
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFCA84CF987FBC050" ID-GBIF-Occurrence="3923978309" pageId="8" pageNumber="43">
<specimenCount id="738FFD03FF8BFF8CFCA84CF98782C07D" box="[871,953,1772,1797]" pageId="8" pageNumber="43" type="male">1 male</specimenCount>
<specimenCount id="738FFD03FF8BFF8CFC074CF98014C07D" box="[968,1071,1772,1797]" pageId="8" pageNumber="43" type="female">1 female</specimenCount>
(
<collectionCode id="0398AE4FFF8BFF8CFB8A4CF98098C07C" box="[1093,1187,1772,1796]" pageId="8" pageNumber="43">IESWU</collectionCode>
), Jingxi, Renzhuang, Tengmaocun, Longlitun, light trap, 2015. 
<date id="1137104AFF8BFF8CFD4B4D1A86F7C050" box="[644,716,1807,1832]" pageId="8" pageNumber="43" value="2011-07">VII.11</date>
, coll. Jian-Yue Qiu.
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFC024D1A8621C035" ID-GBIF-Occurrence="3923978314" pageId="8" pageNumber="43">
<emphasis id="57FDEA98FF8BFF8CFC024D1A80B6C050" bold="true" box="[973,1165,1807,1832]" pageId="8" pageNumber="43">
<collectingCountry id="1D9E761AFF8BFF8CFC024D1A8023C050" box="[973,1048,1807,1832]" name="China" pageId="8" pageNumber="43">China</collectingCountry>
, Hainan:
</emphasis>
<specimenCount id="738FFD03FF8BFF8CFB534D058134C050" box="[1180,1295,1807,1832]" pageId="8" pageNumber="43" type="female">2 females</specimenCount>
(IOZCAS), Tunchang, Xinxing, 1957. 
<date id="1137104AFF8BFF8CFE1B4D21862EC035" box="[468,533,1844,1869]" pageId="8" pageNumber="43" value="2013-06">VI.13</date>
;
</materialsCitation>
<materialsCitation id="D5E13CD7FF8BFF8CFDE44D2180B4C035" ID-GBIF-Occurrence="3923978313" box="[555,1167,1844,1869]" pageId="8" pageNumber="43">
<specimenCount id="738FFD03FF8BFF8CFDE44D2186BBC035" box="[555,640,1844,1869]" pageId="8" pageNumber="43" type="male">1 male</specimenCount>
(
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), 1997. 
<date id="1137104AFF8BFF8CFC9B4D2187B0C035" box="[852,907,1844,1869]" pageId="8" pageNumber="43" value="1925-05">V.25</date>
, coll. Mao-Fa Yang;
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, 2007. 
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, coll. Jian Yao;
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, yellow-disk trap?, 2009. 
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, coll. Ting-Yu Hu;
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, coll. Wen-Xin Lin.
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</paragraph>
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