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<mods:title id="663917D29CCB78657CB5795D946FBE1B">A taxonomic study of the beetle cockroaches (Diploptera Saussure) from China, with notes on the genus and species worldwide (Blattodea: Blaberidae: Diplopterinae)</mods:title>
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<heading id="3E7E81E6FF81FF86FF584E158614C362" bold="true" box="[151,559,1023,1050]" fontSize="11" level="1" pageId="2" pageNumber="37" reason="1">
<emphasis id="57FDEA98FF81FF86FF584E158614C362" bold="true" box="[151,559,1023,1050]" pageId="2" pageNumber="37">
Genus 
<taxonomicName id="A2894D09FF81FF86FF2149EA8614C362" ID-CoL="8MQVN" authority="Saussure, 1864" authorityName="Saussure" authorityYear="1864" box="[238,559,1023,1050]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF81FF86FF2149EA8554C361" bold="true" box="[238,367,1023,1049]" italics="true" pageId="2" pageNumber="37">Diploptera</emphasis>
<bibRefCitation id="01184B7BFF81FF86FEB94E158614C362" author="Saussure, H. de" box="[374,559,1024,1050]" firstAuthor="Saussure" journalOrPublisher="Revue et Magasin de Zoologie Pure et Appliquee" pageId="2" pageNumber="37" pagination="305 - 326" part="16" refId="ref13151" refString="Saussure, H. de (1864 a) Blattarum novarum species aliquot. Revue et Magasin de Zoologie Pure et Appliquee, 16, 305 - 326." title="Blattarum novarum species aliquot" type="journal article" year="1864">Saussure, 1864</bibRefCitation>
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</emphasis>
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<paragraph id="6536368AFF81FF86FF584E5387FAC324" blockId="2.[151,1436,1094,1270]" box="[151,961,1094,1116]" pageId="2" pageNumber="37">
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<taxonomicName id="A2894D09FF81FF86FF584E538644C324" authority="Saussure, 1864a: 325" authorityName="Saussure" authorityPageNumber="325" authorityYear="1864" box="[151,639,1094,1116]" class="Insecta" family="Ectobiidae" genus="Prosoplecta" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="37" phylum="Arthropoda" rank="subGenus" subGenus="Diploptera">
<emphasis id="57FDEA98FF81FF86FF584E5385A2C324" box="[151,409,1094,1116]" italics="true" pageId="2" pageNumber="37">Prosoplecta (Diploptera)</emphasis>
<treatmentCitation id="E428109BFF81FF86FE6F4E538644C324" author="Saussure" box="[416,639,1094,1116]" page="325" pageId="2" pageNumber="37" year="1864">
<bibRefCitation id="01184B7BFF81FF86FE6F4E538644C324" author="Saussure, H. de" box="[416,639,1094,1116]" firstAuthor="Saussure" journalOrPublisher="Revue et Magasin de Zoologie Pure et Appliquee" page="325" pageId="2" pageNumber="37" pagination="305 - 326" part="16" refId="ref13151" refString="Saussure, H. de (1864 a) Blattarum novarum species aliquot. Revue et Magasin de Zoologie Pure et Appliquee, 16, 305 - 326." title="Blattarum novarum species aliquot" type="journal article" year="1864" yearSuffix="a">Saussure, 1864a: 325</bibRefCitation>
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; 
<treatmentCitation id="E428109BFF81FF86FD444E538751C324" author="Saussure" box="[651,874,1094,1116]" page="177" pageId="2" pageNumber="37" year="1864">
<bibRefCitation id="01184B7BFF81FF86FD444E538751C324" author="Saussure, H. de" bookContentInfo="279 pp." box="[651,874,1094,1116]" firstAuthor="Saussure" journalOrPublisher="Geneva" page="177" pageId="2" pageNumber="37" refId="ref13180" refString="Saussure, H. de (1864 b) Memoires pour Servir a l'Histoire Naturelle du Mexique des Antilles et des Etats-Unis. Vol. 4. Orthopteres de l'Amerique Moyenne. Geneva, 279 pp." title="Memoires pour Servir a l'Histoire Naturelle du Mexique des Antilles et des Etats-Unis. Vol. 4. Orthopteres de l'Amerique Moyenne" type="book" year="1864" yearSuffix="b">Saussure, 1864b: 177</bibRefCitation>
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, fig. 28.
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<typeStatus id="BA328828FF81FF86FC064E5287C0C324" box="[969,1019,1095,1116]" pageId="2" pageNumber="37">Type</typeStatus>
species 
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<emphasis id="57FDEA98FF81FF86FB9B4E5381A7C324" box="[1108,1436,1094,1116]" italics="true" pageId="2" pageNumber="37">Prosoplecta (Diploptera) silpha</emphasis>
<bibRefCitation id="01184B7BFF81FF86FF084E708554C303" author="Saussure, H. de" box="[199,367,1125,1147]" firstAuthor="Saussure" journalOrPublisher="Revue et Magasin de Zoologie Pure et Appliquee" pageId="2" pageNumber="37" pagination="305 - 326" part="16" refId="ref13151" refString="Saussure, H. de (1864 a) Blattarum novarum species aliquot. Revue et Magasin de Zoologie Pure et Appliquee, 16, 305 - 326." title="Blattarum novarum species aliquot" type="journal article" year="1864" yearSuffix="a">Saussure, 1864a</bibRefCitation>
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.
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<taxonomicName id="A2894D09FF81FF86FF584E91863EC3E2" ID-CoL="8MQVN" authority="Saussure, 1864b: 166" authorityName="Saussure" authorityPageNumber="166" authorityYear="1864" box="[151,517,1156,1178]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF81FF86FF584E91853DC3E2" box="[151,262,1156,1178]" italics="true" pageId="2" pageNumber="37">Diploptera</emphasis>
: 
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<bibRefCitation id="01184B7BFF81FF86FED44E91863EC3E2" author="Saussure, H. de" bookContentInfo="279 pp." box="[283,517,1156,1178]" firstAuthor="Saussure" journalOrPublisher="Geneva" page="166" pageId="2" pageNumber="37" refId="ref13180" refString="Saussure, H. de (1864 b) Memoires pour Servir a l'Histoire Naturelle du Mexique des Antilles et des Etats-Unis. Vol. 4. Orthopteres de l'Amerique Moyenne. Geneva, 279 pp." title="Memoires pour Servir a l'Histoire Naturelle du Mexique des Antilles et des Etats-Unis. Vol. 4. Orthopteres de l'Amerique Moyenne" type="book" year="1864" yearSuffix="b">Saussure, 1864b: 166</bibRefCitation>
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, 177, elevated from subgenus; 
<treatmentCitation id="E428109BFF81FF86FCAA4E91801EC3E2" author="Walker" box="[869,1061,1156,1178]" page="57" pageId="2" pageNumber="37" year="1868">
<bibRefCitation id="01184B7BFF81FF86FCAA4E91801EC3E2" author="Walker, F." bookContentInfo="239 pp." box="[869,1061,1156,1178]" firstAuthor="Walker" journalOrPublisher="British Museum, London" page="57" pageId="2" pageNumber="37" refId="ref13309" refString="Walker, F. (1868) Catalogue of the Specimens of Blattariae in the Collection of the British Museum. British Museum, London, 239 pp." title="Catalogue of the Specimens of Blattariae in the Collection of the British Museum" type="book" year="1868">Walker, 1868: 57</bibRefCitation>
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; 
<treatmentCitation id="E428109BFF81FF86FBF84E918138C3E2" author="Walker" box="[1079,1283,1156,1178]" page="125" pageId="2" pageNumber="37" year="1869">
<bibRefCitation id="01184B7BFF81FF86FBF84E918138C3E2" author="Walker, F." bookContentInfo="224 pp." box="[1079,1283,1156,1178]" firstAuthor="Walker" journalOrPublisher="British Museum, London" page="125" pageId="2" pageNumber="37" refId="ref13338" refString="Walker, F. (1869) Catalogue of the Specimens of Dermaptera Saltatoria and Supplement to the Blattariae in the Collection of the British Museum. British Museum, London, 224 pp." title="Catalogue of the Specimens of Dermaptera Saltatoria and Supplement to the Blattariae in the Collection of the British Museum" type="book" year="1869">Walker, 1869: 125</bibRefCitation>
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; 
<treatmentCitation id="E428109BFF81FF86FADA4E9085A0C3C1" author="Brunner von Wattenwyl, C." page="40" pageId="2" pageNumber="37" year="1863">
<bibRefCitation id="01184B7BFF81FF86FADA4E9085ADC3C1" author="Brunner von Wattenwyl, C." firstAuthor="Brunner von Wattenwyl" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" page="40" pageId="2" pageNumber="37" pagination="5 - 230" part="2" refId="ref12315" refString="Brunner von Wattenwyl, C. (1893) Revision du systeme des Orthopteres et description des especes rapportees par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 13, 5 - 230." title="Revision du systeme des Orthopteres et description des especes rapportees par M. Leonardo Fea de Birmanie" type="journal article" year="1893">Brunner von Wattenwyl,1893: 40</bibRefCitation>
.
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</paragraph>
<paragraph id="6536368AFF81FF86FF584ED48648C38E" blockId="2.[151,1436,1094,1270]" pageId="2" pageNumber="37">
<treatmentCitationGroup id="459911A4FF81FF86FF584ED48648C38E" pageId="2" pageNumber="37">
<taxonomicName id="A2894D09FF81FF86FF584ED486B0C3AF" ID-CoL="8N6ZQ" authority="Brunner von Wattenwyl, 1865" authorityName="Brunner von Wattenwyl" authorityYear="1865" box="[151,651,1217,1239]" class="Insecta" family="Blaberidae" genus="Eleutheroda" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF81FF86FF584ED48528C3AF" box="[151,275,1217,1239]" italics="true" pageId="2" pageNumber="37">Eleutheroda</emphasis>
<treatmentCitation id="E428109BFF81FF86FED44ED786B0C3AF" author="Brunner von Wattenwyl, C." box="[283,651,1217,1239]" page="264" pageId="2" pageNumber="37" year="1865">
<bibRefCitation id="01184B7BFF81FF86FED44ED786B0C3AF" author="Brunner von Wattenwyl, C." bookContentInfo="426 pp." box="[283,651,1217,1239]" firstAuthor="Brunner von Wattenwyl" journalOrPublisher="Vienna" page="264" pageId="2" pageNumber="37" refId="ref12296" refString="Brunner von Wattenwyl, C. (1865) Nouveau Systeme des Blattaires. Vienna, 426 pp." title="Nouveau Systeme des Blattaires" type="book" year="1865">Brunner von Wattenwyl, 1865: 264</bibRefCitation>
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</taxonomicName>
; 
<treatmentCitation id="E428109BFF81FF86FD584ED4876CC3AF" author="Walker" box="[663,855,1217,1239]" page="125" pageId="2" pageNumber="37" year="1869">
<bibRefCitation id="01184B7BFF81FF86FD584ED4876CC3AF" author="Walker, F." bookContentInfo="224 pp." box="[663,855,1217,1239]" firstAuthor="Walker" journalOrPublisher="British Museum, London" page="125" pageId="2" pageNumber="37" refId="ref13338" refString="Walker, F. (1869) Catalogue of the Specimens of Dermaptera Saltatoria and Supplement to the Blattariae in the Collection of the British Museum. British Museum, London, 224 pp." title="Catalogue of the Specimens of Dermaptera Saltatoria and Supplement to the Blattariae in the Collection of the British Museum" type="book" year="1869">Walker, 1869: 125</bibRefCitation>
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, synonymized with 
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<emphasis id="57FDEA98FF81FF86FBE84ED480ADC3AF" box="[1063,1174,1217,1239]" italics="true" pageId="2" pageNumber="37">Diploptera</emphasis>
</taxonomicName>
; 
<treatmentCitation id="E428109BFF81FF86FB694ED78518C38E" author="Brunner von Wattenwyl, C." page="40" pageId="2" pageNumber="37" year="1893">
<bibRefCitation id="01184B7BFF81FF86FB694ED78518C38E" author="Brunner von Wattenwyl, C." firstAuthor="Brunner von Wattenwyl" journalOrPublisher="Annali del Museo Civico di Storia Naturale di Genova" page="40" pageId="2" pageNumber="37" pagination="5 - 230" part="2" refId="ref12315" refString="Brunner von Wattenwyl, C. (1893) Revision du systeme des Orthopteres et description des especes rapportees par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale di Genova, Series 2, 13, 5 - 230." title="Revision du systeme des Orthopteres et description des especes rapportees par M. Leonardo Fea de Birmanie" type="journal article" year="1893">Brunner von Wattenwyl 1893: 40</bibRefCitation>
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, Walker’s revision was accepted.
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</paragraph>
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<subSubSection id="2D936501FF81FF86FF584F378772C140" pageId="2" pageNumber="37" type="diagnosis">
<paragraph id="6536368AFF81FF86FF584F378772C140" blockId="2.[151,1437,1314,2023]" pageId="2" pageNumber="37">
<emphasis id="57FDEA98FF81FF86FF584F37852AC243" bold="true" box="[151,273,1314,1339]" pageId="2" pageNumber="37">Diagnosis.</emphasis>
<taxonomicName id="A2894D09FF81FF86FED24F3685A3C243" box="[285,408,1315,1339]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF81FF86FED24F3685A3C243" box="[285,408,1315,1339]" italics="true" pageId="2" pageNumber="37">Diploptera</emphasis>
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is distinguished from other blaberid cockroaches mainly by its wings: tegmen heavily sclerotized, resembling the elytron of beetles; hindwing folds following the similar pattern of beetle’s, with abundant cross-veins and a transverse fold, along which the hindwing reflexes and which runs transversely and nearly halves the hindwing, even with an uncommon anal fold ending at the apical angle. Compared to other cockroaches with transversely reflexed hindwings, 
<taxonomicName id="A2894D09FF81FF86FD2D4FA68766C2B3" box="[738,861,1459,1483]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF81FF86FD2D4FA68766C2B3" box="[738,861,1459,1483]" italics="true" pageId="2" pageNumber="37">Diploptera</emphasis>
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possesses a different appendicular field on which cross-veins and several longitudinal veins are present, whilst other groups have one or two anal veins only, such as 
<taxonomicName id="A2894D09FF81FF86FF584FEE8532C16B" box="[151,265,1531,1555]" class="Insecta" family="Ectobiidae" genus="Anaplecta" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF81FF86FF584FEE8532C16B" box="[151,265,1531,1555]" italics="true" pageId="2" pageNumber="37">Anaplecta</emphasis>
</taxonomicName>
and 
<taxonomicName id="A2894D09FF81FF86FE8E4FEE85FFC16B" box="[321,452,1531,1555]" class="Insecta" family="Ectobiidae" genus="Plectoptera" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF81FF86FE8E4FEE85FFC16B" box="[321,452,1531,1555]" italics="true" pageId="2" pageNumber="37">Plectoptera</emphasis>
</taxonomicName>
. 
<taxonomicName id="A2894D09FF81FF86FE1C4FEE8675C16B" box="[467,590,1531,1555]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="2" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF81FF86FE1C4FEE8675C16B" box="[467,590,1531,1555]" italics="true" pageId="2" pageNumber="37">Diploptera</emphasis>
</taxonomicName>
is also unique by the three-segmented cercus, whilst that of other blaberid cockroaches is multi-segmented or coalesced to one segment.
</paragraph>
</subSubSection>
<subSubSection id="2D936501FF81FF87FF084C578580C465" lastPageId="3" lastPageNumber="38" pageId="2" pageNumber="37" type="description">
<paragraph id="6536368AFF81FF87FF084C578137C631" blockId="2.[151,1437,1314,2023]" lastBlockId="3.[151,1437,151,1877]" lastPageId="3" lastPageNumber="38" pageId="2" pageNumber="37">
<emphasis id="57FDEA98FF81FF86FF084C57854FC123" bold="true" box="[199,372,1602,1627]" pageId="2" pageNumber="37">Redescription.</emphasis>
General. The insects resemble beetles. Body size medium to small, females always obviously larger than males. Head relatively large, exposed; ocular distance accounting for about half of head width, subequal to that between antennal sockets, greater than ocellar distance; ocellus spot oval or crescent, oblique; antennae always with the same length of body or longer. Pronotum and tegmina always setose or pubescent. Mesonotum with an exposed triangular bump bordered by pronotum and tegmina, which resembles a scutellum and was described as so (see 
<bibRefCitation id="01184B7BFF81FF86FE454CE38670C077" author="Anisyutkin, L. N." box="[394,587,1782,1807]" firstAuthor="Anisyutkin" journalOrPublisher="Zoosystematica Rossica" pageId="2" pageNumber="37" pagination="173 - 175" part="16" refId="ref12157" refString="Anisyutkin, L. N. (2007) A new species of the genus Diploptera Saussure, 1864 from Borneo (Dictyoptera: Blaberidae: Diplopterinae). Zoosystematica Rossica, 16 (2), 173 - 175." title="A new species of the genus Diploptera Saussure, 1864 from Borneo (Dictyoptera: Blaberidae: Diplopterinae)" type="journal article" year="2007">Anisyutkin 2007</bibRefCitation>
) but is not that case. Tegmina and hind wings fully developed. Tegmina strongly sclerotized, with veins almost indistinguishable; the base in ventral view with a ridge along the subcosta (Sc); the covered portion of right tegmen translucent. Hindwings large (
<figureCitation id="FDB22A0FFF81FF86FC1E4D2B8014C02F" box="[977,1071,1854,1879]" captionStart="FIGURES 31 – 36. D" captionStartId="9.[151,264,1815,1838]" captionTargetBox="[151,1433,198,1789]" captionTargetId="figure@9.[151,1436,195,1791]" captionTargetPageId="9" captionText="FIGURES 31 – 36. D. punctata (Eschscholtz, 1822) from China, Yunnan, Menglun. 31 hind wing, female, with grey coloration indicating sclerotization; 32 – 36 male; 32 terminal segment of dorsal abdomen, in ventral view; 33 subgenital plate in dorsal view; 34 – 36 phallic sclerites in dorsal view; 34 sclerites R’; 35 sclerites L 2 ’; 36 apex of left complex hook, showing sclerite L 3 ’. (ce. = cercus, m. = membranous, p. p. = paraprocts, st. = stylus; other abbreviations as indicated in the text. Scale bars = 10 mm for 31, 1 mm for 32 &amp; 33, 500 µm for 34 – 36.)" httpUri="https://zenodo.org/record/289452/files/figure.png" pageId="2" pageNumber="37">
Figs. 
<emphasis id="57FDEA98FF81FF86FBDC4D2B8014C02F" bold="true" box="[1043,1071,1854,1879]" pageId="2" pageNumber="37">31</emphasis>
</figureCitation>
, 
<figureCitation id="FDB22A0FFF81FF86FBF04D2B8060C02F" box="[1087,1115,1854,1879]" captionStart="FIGURES 42 – 47" captionStartId="11.[151,264,1868,1891]" captionTargetBox="[154,1429,988,1841]" captionTargetId="figure@11.[151,1436,986,1844]" captionTargetPageId="11" captionText="FIGURES 42 – 47. Diploptera naevus sp. n., holotype. 42 hind wing, with grey coloration indicating sclerotization; 43 terminal segment of dorsal abdomen, in ventral view; 44 subgenital plate in dorsal view; 45 – 47 phallic sclerites in dorsal view; 45 sclerites R’; 46 sclerites L 2 ’; 47 apex of left complex hook, showing sclerite L 3 ’. (m. = membranous. Scale bars = 5 mm for 42, 1 mm for 43 &amp; 44, 500 µm for 45 – 47.)" httpUri="https://zenodo.org/record/289454/files/figure.png" pageId="2" pageNumber="37">
<emphasis id="57FDEA98FF81FF86FBF04D2B8060C02F" bold="true" box="[1087,1115,1854,1879]" pageId="2" pageNumber="37">42</emphasis>
</figureCitation>
&amp; 
<figureCitation id="FDB22A0FFF81FF86FB4C4D2B80A4C02F" box="[1155,1183,1854,1879]" captionStart="FIGURES 48 – 53" captionStartId="12.[151,264,1914,1937]" captionTargetBox="[192,1384,1016,1885]" captionTargetId="figure@12.[186,1391,1009,1888]" captionTargetPageId="12" captionText="FIGURES 48 – 53. Diploptera nigrescens guani subsp. n., holotype. 48 hind wing, with grey coloration indicating sclerotization; 49 terminal segment of dorsal abdomen, in ventral view; 50 subgenital plate in dorsal view; 51 – 53 phallic sclerites in dorsal view; 51 sclerites R’; 52 sclerites L 2 ’; 53 apex of left complex hook, showing sclerite L 3 ’. (m. = membranous. Scale bars = 5 mm for 48, 1 mm for 49 &amp; 50, 500 µm for 51 – 53.)" httpUri="https://zenodo.org/record/289455/files/figure.png" pageId="2" pageNumber="37">
<emphasis id="57FDEA98FF81FF86FB4C4D2B80A4C02F" bold="true" box="[1155,1183,1854,1879]" pageId="2" pageNumber="37">48</emphasis>
</figureCitation>
), about two times the length of tegmina when unfolded, the costal border sclerotized (see grey coloration in 
<figureCitation id="FDB22A0FFF81FF86FB904D778082C003" box="[1119,1209,1890,1915]" captionStart="FIGURES 31 – 36. D" captionStartId="9.[151,264,1815,1838]" captionTargetBox="[151,1433,198,1789]" captionTargetId="figure@9.[151,1436,195,1791]" captionTargetPageId="9" captionText="FIGURES 31 – 36. D. punctata (Eschscholtz, 1822) from China, Yunnan, Menglun. 31 hind wing, female, with grey coloration indicating sclerotization; 32 – 36 male; 32 terminal segment of dorsal abdomen, in ventral view; 33 subgenital plate in dorsal view; 34 – 36 phallic sclerites in dorsal view; 34 sclerites R’; 35 sclerites L 2 ’; 36 apex of left complex hook, showing sclerite L 3 ’. (ce. = cercus, m. = membranous, p. p. = paraprocts, st. = stylus; other abbreviations as indicated in the text. Scale bars = 10 mm for 31, 1 mm for 32 &amp; 33, 500 µm for 34 – 36.)" httpUri="https://zenodo.org/record/289452/files/figure.png" pageId="2" pageNumber="37">
Figs. 
<emphasis id="57FDEA98FF81FF86FB524D778082C003" bold="true" box="[1181,1209,1890,1915]" pageId="2" pageNumber="37">31</emphasis>
</figureCitation>
, 
<figureCitation id="FDB22A0FFF81FF86FB084D7780D8C003" box="[1223,1251,1890,1915]" captionStart="FIGURES 42 – 47" captionStartId="11.[151,264,1868,1891]" captionTargetBox="[154,1429,988,1841]" captionTargetId="figure@11.[151,1436,986,1844]" captionTargetPageId="11" captionText="FIGURES 42 – 47. Diploptera naevus sp. n., holotype. 42 hind wing, with grey coloration indicating sclerotization; 43 terminal segment of dorsal abdomen, in ventral view; 44 subgenital plate in dorsal view; 45 – 47 phallic sclerites in dorsal view; 45 sclerites R’; 46 sclerites L 2 ’; 47 apex of left complex hook, showing sclerite L 3 ’. (m. = membranous. Scale bars = 5 mm for 42, 1 mm for 43 &amp; 44, 500 µm for 45 – 47.)" httpUri="https://zenodo.org/record/289454/files/figure.png" pageId="2" pageNumber="37">
<emphasis id="57FDEA98FF81FF86FB084D7780D8C003" bold="true" box="[1223,1251,1890,1915]" pageId="2" pageNumber="37">42</emphasis>
</figureCitation>
&amp; 
<figureCitation id="FDB22A0FFF81FF86FAC84D778118C003" box="[1287,1315,1890,1915]" captionStart="FIGURES 48 – 53" captionStartId="12.[151,264,1914,1937]" captionTargetBox="[192,1384,1016,1885]" captionTargetId="figure@12.[186,1391,1009,1888]" captionTargetPageId="12" captionText="FIGURES 48 – 53. Diploptera nigrescens guani subsp. n., holotype. 48 hind wing, with grey coloration indicating sclerotization; 49 terminal segment of dorsal abdomen, in ventral view; 50 subgenital plate in dorsal view; 51 – 53 phallic sclerites in dorsal view; 51 sclerites R’; 52 sclerites L 2 ’; 53 apex of left complex hook, showing sclerite L 3 ’. (m. = membranous. Scale bars = 5 mm for 48, 1 mm for 49 &amp; 50, 500 µm for 51 – 53.)" httpUri="https://zenodo.org/record/289455/files/figure.png" pageId="2" pageNumber="37">
<emphasis id="57FDEA98FF81FF86FAC84D778118C003" bold="true" box="[1287,1315,1890,1915]" pageId="2" pageNumber="37">48</emphasis>
</figureCitation>
); venation reticular, longitudinal veins curved near the transverse fold (tfd), which runs across the middle of hindwing and results in a large appendicular field occupying nearly half of the hindwing; subcosta (Sc) and media (M) simple, radius (R) bifurcated near arculus (= mp-cua cross-vein); anterior cubitus (CuA) robust, branched at tfd; first three anal veins (AA, AP1 and AP2) curved strongly near tfd, AA towards costal margin (but the curved end may be a splice with a cross-vein), whilst the AP veins towards outer margin; anal fold (afd) running between AA and AP1, ending at the apical angle; branches of AP1 occupying a remarkably large area. Front femur 
<typeStatus id="BA328828FF80FF87FB694AF380EDC786" box="[1190,1238,230,254]" pageId="3" pageNumber="38">type</typeStatus>
C1; tarsal pulvilli large, present on the first to the fourth proximal tarsomeres; claws symmetrical and unspecialized; arolia large. Cerci tapered, with three segments and the basal one accounting for more than half of the total length.
</paragraph>
<paragraph id="6536368AFF80FF87FF084B418745C5C8" blockId="3.[151,1437,151,1877]" pageId="3" pageNumber="38">
<collectingCountry id="1D9E761AFF80FF87FF084B41853FC615" box="[199,260,340,365]" name="Malta" pageId="3" pageNumber="38">Male</collectingCountry>
. Abdomen with T1–T9 specialization and tergal gland absent or invisible; T8 and T9 covered, S8 covered by S7. Supra-anal plate (T10) subtrapezoidal, specialized with a transverse membranous field (m.) in the middle; right paraproct with a hook at the apex. Subgenital plate (S9) asymmetrical (
<figureCitation id="FDB22A0FFF80FF87FBDC4B89804AC6CD" box="[1043,1137,412,437]" captionStart="FIGURES 31 – 36. D" captionStartId="9.[151,264,1815,1838]" captionTargetBox="[151,1433,198,1789]" captionTargetId="figure@9.[151,1436,195,1791]" captionTargetPageId="9" captionText="FIGURES 31 – 36. D. punctata (Eschscholtz, 1822) from China, Yunnan, Menglun. 31 hind wing, female, with grey coloration indicating sclerotization; 32 – 36 male; 32 terminal segment of dorsal abdomen, in ventral view; 33 subgenital plate in dorsal view; 34 – 36 phallic sclerites in dorsal view; 34 sclerites R’; 35 sclerites L 2 ’; 36 apex of left complex hook, showing sclerite L 3 ’. (ce. = cercus, m. = membranous, p. p. = paraprocts, st. = stylus; other abbreviations as indicated in the text. Scale bars = 10 mm for 31, 1 mm for 32 &amp; 33, 500 µm for 34 – 36.)" httpUri="https://zenodo.org/record/289452/files/figure.png" pageId="3" pageNumber="38">
Figs. 
<emphasis id="57FDEA98FF80FF87FB9A4B89804AC6CD" bold="true" box="[1109,1137,412,437]" pageId="3" pageNumber="38">33</emphasis>
</figureCitation>
, 
<figureCitation id="FDB22A0FFF80FF87FB4D4B8980A5C6CD" box="[1154,1182,412,437]" captionStart="FIGURES 37 – 41" captionStartId="10.[151,264,1906,1929]" captionTargetBox="[151,1436,1061,1878]" captionTargetId="figure@10.[151,1436,1058,1883]" captionTargetPageId="10" captionText="FIGURES 37 – 41. Diploptera elliptica sp. n., holotype. 37 terminal segment of dorsal abdomen, in ventral view; 38 subgenital plate in dorsal view; 39 – 41 phallic sclerites in dorsal view; 39 sclerites R’; 40 sclerites L 2 ’; 41 apex of left complex hook, showing sclerite L 3 ’. (m. = membranous. Scale bars = 1 mm for 37 – 38, 500 µm for 39 – 41.)" httpUri="https://zenodo.org/record/289453/files/figure.png" pageId="3" pageNumber="38">
<emphasis id="57FDEA98FF80FF87FB4D4B8980A5C6CD" bold="true" box="[1154,1182,412,437]" pageId="3" pageNumber="38">38</emphasis>
</figureCitation>
, 
<figureCitation id="FDB22A0FFF80FF87FB604B8980F0C6CD" box="[1199,1227,412,437]" captionStart="FIGURES 42 – 47" captionStartId="11.[151,264,1868,1891]" captionTargetBox="[154,1429,988,1841]" captionTargetId="figure@11.[151,1436,986,1844]" captionTargetPageId="11" captionText="FIGURES 42 – 47. Diploptera naevus sp. n., holotype. 42 hind wing, with grey coloration indicating sclerotization; 43 terminal segment of dorsal abdomen, in ventral view; 44 subgenital plate in dorsal view; 45 – 47 phallic sclerites in dorsal view; 45 sclerites R’; 46 sclerites L 2 ’; 47 apex of left complex hook, showing sclerite L 3 ’. (m. = membranous. Scale bars = 5 mm for 42, 1 mm for 43 &amp; 44, 500 µm for 45 – 47.)" httpUri="https://zenodo.org/record/289454/files/figure.png" pageId="3" pageNumber="38">
<emphasis id="57FDEA98FF80FF87FB604B8980F0C6CD" bold="true" box="[1199,1227,412,437]" pageId="3" pageNumber="38">44</emphasis>
</figureCitation>
&amp; 
<figureCitation id="FDB22A0FFF80FF87FB3A4B89812AC6CD" box="[1269,1297,412,437]" captionStart="FIGURES 48 – 53" captionStartId="12.[151,264,1914,1937]" captionTargetBox="[192,1384,1016,1885]" captionTargetId="figure@12.[186,1391,1009,1888]" captionTargetPageId="12" captionText="FIGURES 48 – 53. Diploptera nigrescens guani subsp. n., holotype. 48 hind wing, with grey coloration indicating sclerotization; 49 terminal segment of dorsal abdomen, in ventral view; 50 subgenital plate in dorsal view; 51 – 53 phallic sclerites in dorsal view; 51 sclerites R’; 52 sclerites L 2 ’; 53 apex of left complex hook, showing sclerite L 3 ’. (m. = membranous. Scale bars = 5 mm for 48, 1 mm for 49 &amp; 50, 500 µm for 51 – 53.)" httpUri="https://zenodo.org/record/289455/files/figure.png" pageId="3" pageNumber="38">
<emphasis id="57FDEA98FF80FF87FB3A4B89812AC6CD" bold="true" box="[1269,1297,412,437]" pageId="3" pageNumber="38">50</emphasis>
</figureCitation>
), with right posterolateral margin emarginate and with two dorsal sclerotized lobes (S9d), the left larger than the right; styli similar, fingerlike, of which the insertions on the subgenital plate are membranous, the left membranous area markedly smaller than the right. Genitalia with hook-like sclerite L3’ (= R 
<quantity id="A2719B6FFF80FF87FBEC481C806FC559" box="[1059,1108,520,545]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="38" unit="in" value="2.0">2 in</quantity>
<bibRefCitation id="01184B7BFF80FF87FBAE481D811CC559" author="McKittrick, F. A." box="[1121,1319,520,545]" firstAuthor="McKittrick" journalOrPublisher="Cornell University Agricultural Experiment Station Memoir" pageId="3" pageNumber="38" pagination="1 - 197" part="389" refId="ref12884" refString="McKittrick, F. A. (1964) Evolutionary studies of cockroaches. Cornell University Agricultural Experiment Station Memoir, 389, 1 - 197." title="Evolutionary studies of cockroaches" type="journal article" year="1964">McKittrick 1964</bibRefCitation>
, similarly hereinafter) on the right side, main sclerite of L2’ (= L2vm) rodlike, apical sclerite of L2’ (= L2d) discoid; R’ (= L1) composed of two sclerites, of which the folded one sandwiches part of the other, the latter with a long, arcuate, and spinous structure running parallelly with the anterior margin of the main part of the sclerite (
<figureCitation id="FDB22A0FFF80FF87FB1F48618110C5F5" box="[1232,1323,628,653]" captionStart="FIGURES 31 – 36. D" captionStartId="9.[151,264,1815,1838]" captionTargetBox="[151,1433,198,1789]" captionTargetId="figure@9.[151,1436,195,1791]" captionTargetPageId="9" captionText="FIGURES 31 – 36. D. punctata (Eschscholtz, 1822) from China, Yunnan, Menglun. 31 hind wing, female, with grey coloration indicating sclerotization; 32 – 36 male; 32 terminal segment of dorsal abdomen, in ventral view; 33 subgenital plate in dorsal view; 34 – 36 phallic sclerites in dorsal view; 34 sclerites R’; 35 sclerites L 2 ’; 36 apex of left complex hook, showing sclerite L 3 ’. (ce. = cercus, m. = membranous, p. p. = paraprocts, st. = stylus; other abbreviations as indicated in the text. Scale bars = 10 mm for 31, 1 mm for 32 &amp; 33, 500 µm for 34 – 36.)" httpUri="https://zenodo.org/record/289452/files/figure.png" pageId="3" pageNumber="38">
Figs. 
<emphasis id="57FDEA98FF80FF87FAC048618110C5F5" bold="true" box="[1295,1323,628,653]" pageId="3" pageNumber="38">34</emphasis>
</figureCitation>
, 
<figureCitation id="FDB22A0FFF80FF87FAF64861816EC5F5" box="[1337,1365,628,653]" captionStart="FIGURES 37 – 41" captionStartId="10.[151,264,1906,1929]" captionTargetBox="[151,1436,1061,1878]" captionTargetId="figure@10.[151,1436,1058,1883]" captionTargetPageId="10" captionText="FIGURES 37 – 41. Diploptera elliptica sp. n., holotype. 37 terminal segment of dorsal abdomen, in ventral view; 38 subgenital plate in dorsal view; 39 – 41 phallic sclerites in dorsal view; 39 sclerites R’; 40 sclerites L 2 ’; 41 apex of left complex hook, showing sclerite L 3 ’. (m. = membranous. Scale bars = 1 mm for 37 – 38, 500 µm for 39 – 41.)" httpUri="https://zenodo.org/record/289453/files/figure.png" pageId="3" pageNumber="38">
<emphasis id="57FDEA98FF80FF87FAF64861816EC5F5" bold="true" box="[1337,1365,628,653]" pageId="3" pageNumber="38">39</emphasis>
</figureCitation>
, 
<figureCitation id="FDB22A0FFF80FF87FAAC48618144C5F5" box="[1379,1407,628,653]" captionStart="FIGURES 42 – 47" captionStartId="11.[151,264,1868,1891]" captionTargetBox="[154,1429,988,1841]" captionTargetId="figure@11.[151,1436,986,1844]" captionTargetPageId="11" captionText="FIGURES 42 – 47. Diploptera naevus sp. n., holotype. 42 hind wing, with grey coloration indicating sclerotization; 43 terminal segment of dorsal abdomen, in ventral view; 44 subgenital plate in dorsal view; 45 – 47 phallic sclerites in dorsal view; 45 sclerites R’; 46 sclerites L 2 ’; 47 apex of left complex hook, showing sclerite L 3 ’. (m. = membranous. Scale bars = 5 mm for 42, 1 mm for 43 &amp; 44, 500 µm for 45 – 47.)" httpUri="https://zenodo.org/record/289454/files/figure.png" pageId="3" pageNumber="38">
<emphasis id="57FDEA98FF80FF87FAAC48618144C5F5" bold="true" box="[1379,1407,628,653]" pageId="3" pageNumber="38">45</emphasis>
</figureCitation>
&amp; 
<figureCitation id="FDB22A0FFF80FF87FF58488D8488C5C9" box="[151,179,664,689]" captionStart="FIGURES 48 – 53" captionStartId="12.[151,264,1914,1937]" captionTargetBox="[192,1384,1016,1885]" captionTargetId="figure@12.[186,1391,1009,1888]" captionTargetPageId="12" captionText="FIGURES 48 – 53. Diploptera nigrescens guani subsp. n., holotype. 48 hind wing, with grey coloration indicating sclerotization; 49 terminal segment of dorsal abdomen, in ventral view; 50 subgenital plate in dorsal view; 51 – 53 phallic sclerites in dorsal view; 51 sclerites R’; 52 sclerites L 2 ’; 53 apex of left complex hook, showing sclerite L 3 ’. (m. = membranous. Scale bars = 5 mm for 48, 1 mm for 49 &amp; 50, 500 µm for 51 – 53.)" httpUri="https://zenodo.org/record/289455/files/figure.png" pageId="3" pageNumber="38">
<emphasis id="57FDEA98FF80FF87FF58488D8488C5C9" bold="true" box="[151,179,664,689]" pageId="3" pageNumber="38">51</emphasis>
</figureCitation>
). The spatial arrangement of genitalia is as indicated in 
<figureCitation id="FDB22A0FFF80FF87FCE7488D874CC5C9" box="[808,887,664,689]" captionStart="FIGURES 72 – 75. D" captionStartId="17.[151,264,692,715]" captionTargetBox="[152,1436,195,669]" captionTargetId="figure@17.[151,1436,195,669]" captionTargetPageId="17" captionText="FIGURES 72 – 75. D. minor (Brunner von Wattenwyl, 1865), genitalia slide (NHMW) of the male syntype. 72 terminal two segments of dorsal abdomen, in dorsal view; 73 subgenital plate and phallic sclerites, in dorsal view with a green arrow pointing the apical sclerite of L 2 ’; 74 apical part of sclerite L 3 ’ in detail, showing the curved tip; 75 overall view of the slide. (Excluding 74, photographed by Harald Bruckner, copyright © by Natural History Museum Vienna, NOaS Image Collection, published with permission. Scale bars = 500 µm for 72 &amp; 73, 100 µm for 74.)" httpUri="https://zenodo.org/record/289459/files/figure.png" pageId="3" pageNumber="38">
Fig. 
<emphasis id="57FDEA98FF80FF87FC94488D874CC5C9" bold="true" box="[859,887,664,689]" pageId="3" pageNumber="38">73</emphasis>
</figureCitation>
.
</paragraph>
<paragraph id="6536368AFF80FF87FF0848A98580C465" blockId="3.[151,1437,151,1877]" pageId="3" pageNumber="38">
Female. Abdomen with T8 covered by T7, T 
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anterior-posterior extension extremely short and almost thready. Supra-anal plate (T10) subtrapezoidal, without membranous specialization. Subgenital plate (S7) with posterior margin rounded.
</paragraph>
</subSubSection>
<subSubSection id="2D936501FF80FF87FF08493D84D9C41D" pageId="3" pageNumber="38" type="distribution">
<paragraph id="6536368AFF80FF87FF08493D84D9C41D" blockId="3.[151,1437,151,1877]" pageId="3" pageNumber="38">
<emphasis id="57FDEA98FF80FF87FF08493D8562C439" bold="true" box="[199,345,808,833]" pageId="3" pageNumber="38">Distribution</emphasis>
(
<figureCitation id="FDB22A0FFF80FF87FEA5493D8595C439" box="[362,430,808,833]" captionStart="FIGURE 1" captionStartId="5.[151,250,1807,1829]" captionTargetBox="[151,1436,195,1782]" captionTargetId="figure@5.[151,1436,195,1783]" captionTargetPageId="5" captionText="FIGURE 1. Distribution of Diploptera in China and Southeast Asia. The precise localities are painted black, whilst the imprecise ones are in white; each type locality is marked with a ring; the (sub) species are indicated by numbers in alphabetical order: 1 D. bicolor, 2 D. elliptica sp. n., 3 D. erythrocephala, 4 D. maculata, 5 D. minor, 6 D. naevus sp. n., 7 D. nigrescens guani subsp. n., 8 D. nigrescens nigrescens, 9 D. parva, × D. punctata; asterisk stands for the habitat of the Thai D. punctata (see the text)." httpUri="https://zenodo.org/record/289449/files/figure.png" pageId="3" pageNumber="38">
Fig. 
<emphasis id="57FDEA98FF80FF87FE6F493D8595C439" bold="true" box="[416,430,808,833]" pageId="3" pageNumber="38">1</emphasis>
</figureCitation>
). Oriental Region and the north of Australian Region, mostly Southeast Asia and South 
<collectingCountry id="1D9E761AFF80FF87FF58495984E5C41D" box="[151,222,844,869]" name="China" pageId="3" pageNumber="38">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="2D936501FF80FF87FF08496586C6C389" pageId="3" pageNumber="38" type="discussion">
<paragraph id="6536368AFF80FF87FF0849658149C345" blockId="3.[151,1437,151,1877]" pageId="3" pageNumber="38">
<emphasis id="57FDEA98FF80FF87FF08496585ADC4F1" bold="true" box="[199,406,880,905]" pageId="3" pageNumber="38">Taxonomic notes.</emphasis>
Since early descriptions of species were too simple and were based mainly on body size and color pattern, there would be difficulties in separating one species from each other. It is noteworthy enough that the relative length of tegmen had been used as a diagnostic character; however, as a result of the retractable abdomen, it is not adequate to rely on whether the tegmen exceeds the end of abdomen or not. The relative length of tegmen could be of diagnostic value by comparing it to the size of pronotum, which stands reliably for body size; however, tegmen length may be varied too among conspecific individuals so as to make itself less valuable in taxonomy.
</paragraph>
<paragraph id="6536368AFF80FF87FF084E5D8671C3B5" blockId="3.[151,1437,151,1877]" pageId="3" pageNumber="38">
After examining the 
<typeStatus id="BA328828FF80FF87FE7E4E5C85D7C319" box="[433,492,1097,1121]" pageId="3" pageNumber="38">types</typeStatus>
and other specimens, we suggest establishing a 
<emphasis id="57FDEA98FF80FF87FBC24E5C8125C319" bold="true" box="[1037,1310,1096,1121]" pageId="3" pageNumber="38">
<taxonomicName id="A2894D09FF80FF87FBC24E5C804AC318" box="[1037,1137,1097,1120]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF80FF87FBC24E5C804AC318" bold="true" box="[1037,1137,1097,1120]" italics="true" pageId="3" pageNumber="38">punctata</emphasis>
</taxonomicName>
-species-group
</emphasis>
containing the following nominal species: 
<taxonomicName id="A2894D09FF80FF87FE394E788641C3FC" box="[502,634,1133,1156]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF80FF87FE394E788641C3FC" box="[502,634,1133,1156]" italics="true" pageId="3" pageNumber="38">D. punctata</emphasis>
</taxonomicName>
, 
<taxonomicName id="A2894D09FF80FF87FD474E7886D7C3FC" box="[648,748,1133,1156]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="57FDEA98FF80FF87FD474E7886D7C3FC" box="[648,748,1133,1156]" italics="true" pageId="3" pageNumber="38">D. minor</emphasis>
</taxonomicName>
, 
<taxonomicName id="A2894D09FF80FF87FD354E78805BC3FD" authority="Princis, 1950" authorityName="Princis" authorityYear="1950" box="[762,1120,1132,1157]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="erythrocephala">
<emphasis id="57FDEA98FF80FF87FD354E7887FDC3FC" box="[762,966,1132,1156]" italics="true" pageId="3" pageNumber="38">D. erythrocephala</emphasis>
<bibRefCitation id="01184B7BFF80FF87FC034E79805BC3FD" author="Princis, K." box="[972,1120,1132,1157]" firstAuthor="Princis" journalOrPublisher="Opuscula Entomologica" pageId="3" pageNumber="38" pagination="161 - 188" part="15" refId="ref12911" refString="Princis, K. (1950) Indomalaiische und australische Blattarien aus dem Entomologischen Museum der Universitat in Lund. Opuscula Entomologica, 15, 161 - 188." title="Indomalaiische und australische Blattarien aus dem Entomologischen Museum der Universitat in Lund" type="journal article" year="1950">Princis, 1950</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="A2894D09FF80FF87FB584E7881A3C3FD" authority="Princis, 1953" authorityName="Princis" authorityYear="1953" box="[1175,1432,1132,1157]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="38" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="57FDEA98FF80FF87FB584E7880C0C3FC" box="[1175,1275,1133,1156]" italics="true" pageId="3" pageNumber="38">D. parva</emphasis>
<bibRefCitation id="01184B7BFF80FF87FACE4E7981A3C3FD" author="Princis, K." box="[1281,1432,1132,1157]" firstAuthor="Princis" journalOrPublisher="Entomologisk Tidskrift" pageId="3" pageNumber="38" pagination="203 - 213" part="74" refId="ref12940" refString="Princis, K. (1953) Kleine Beitrage zur Kenntnis der Blattarien und ihrer Verbreitung. VI. Entomologisk Tidskrift, 74 (4), 203 - 213." title="Kleine Beitrage zur Kenntnis der Blattarien und ihrer Verbreitung. VI" type="journal article" year="1953">Princis, 1953</bibRefCitation>
</taxonomicName>
. These are almost indistinguishable from each other, and there is the probability that one or more may become junior synonyms with further research.
</paragraph>
<paragraph id="6536368AFF80FF87FF084ECD86C6C389" blockId="3.[151,1437,151,1877]" box="[199,765,1240,1265]" pageId="3" pageNumber="38">Details are given within the entries for taxa below.</paragraph>
</subSubSection>
<subSubSection id="2D936501FF80FF87FF084EE9810AC02C" pageId="3" pageNumber="38" type="discussion">
<paragraph id="6536368AFF80FF87FF084EE98585C121" blockId="3.[151,1437,151,1877]" pageId="3" pageNumber="38">
<emphasis id="57FDEA98FF80FF87FF084EE98500C26D" bold="true" box="[199,315,1276,1301]" pageId="3" pageNumber="38">Remarks.</emphasis>
<taxonomicName id="A2894D09FF80FF87FE8C4EE98585C26C" box="[323,446,1276,1300]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF80FF87FE8C4EE98585C26C" box="[323,446,1276,1300]" italics="true" pageId="3" pageNumber="38">Diploptera</emphasis>
</taxonomicName>
is unique among genera of cockroaches in that the species vary more in the shape and markings of pronotum (
<figureCitation id="FDB22A0FFF80FF87FE724F358605C241" box="[445,574,1312,1337]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="3" pageNumber="38">
Figs. 
<emphasis id="57FDEA98FF80FF87FDC94F358605C241" bold="true" box="[518,574,1312,1337]" pageId="3" pageNumber="38">2–16</emphasis>
</figureCitation>
) than in the male genitalia and genital segments. Such a significant differentiation in the posterolateral angles of pronotum could indicate two routes of evolution and the pronotum is almost uniformed among species in most of cockroach genera; when pronotal shape is various in one genus, it is often correlative to wings development and the cercus length, 
<emphasis id="57FDEA98FF80FF87FCAB4F9887BDC2DC" box="[868,902,1421,1444]" italics="true" pageId="3" pageNumber="38">i.e.</emphasis>
, shorter wings and cerci may company with a pronotum with sharper posterolateral angles, but 
<taxonomicName id="A2894D09FF80FF87FD0D4FA58706C2B0" box="[706,829,1456,1480]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF80FF87FD0D4FA58706C2B0" box="[706,829,1456,1480]" italics="true" pageId="3" pageNumber="38">Diploptera</emphasis>
</taxonomicName>
is not that case. We could definitely divide the genus into two groups by the shape of pronotum (cp. 
<figureCitation id="FDB22A0FFF80FF87FD744FC18708C295" box="[699,819,1492,1517]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="3" pageNumber="38">
Figs. 
<emphasis id="57FDEA98FF80FF87FD334FC18708C295" bold="true" box="[764,819,1492,1517]" pageId="3" pageNumber="38">2–12</emphasis>
</figureCitation>
with 
<figureCitation id="FDB22A0FFF80FF87FCB74FC187C4C295" box="[888,1023,1492,1517]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="3" pageNumber="38">
Figs. 
<emphasis id="57FDEA98FF80FF87FC764FC187C4C295" bold="true" box="[953,1023,1492,1517]" pageId="3" pageNumber="38">13–16</emphasis>
</figureCitation>
); but aside from male genitalia and genital segments, other characters are also less distinct, such as tergal specialization, front femur 
<typeStatus id="BA328828FF80FF87FB004FEC80C4C169" box="[1231,1279,1529,1553]" pageId="3" pageNumber="38">type</typeStatus>
and venation, therefore the species should remain in one genus rather than be placed in two or more artificial genera based merely on such a single character.
</paragraph>
<paragraph id="6536368AFF80FF87FF084C70810AC02C" blockId="3.[151,1437,151,1877]" pageId="3" pageNumber="38">
Some cockroach genera also have a hindwing with a reflexed area (appendicular field or apical triangle), but they evolve probably in a different way from 
<taxonomicName id="A2894D09FF80FF87FD594C9D872AC1D8" box="[662,785,1672,1696]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF80FF87FD594C9D872AC1D8" box="[662,785,1672,1696]" italics="true" pageId="3" pageNumber="38">Diploptera</emphasis>
</taxonomicName>
. Reflexed area of them is bordered anteriorly by CuP and posteriorly by AP, the fact could tell that the area in evolutionary history is enlarged from the narrow space between CuP and AP apically, along which the folding lines run; the area is free of cross-veins as its original. Differently, 
<taxonomicName id="A2894D09FF80FF87FF584CE18529C074" box="[151,274,1780,1804]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="3" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF80FF87FF584CE18529C074" box="[151,274,1780,1804]" italics="true" pageId="3" pageNumber="38">Diploptera</emphasis>
</taxonomicName>
gained the appendicular field by cutting off the main veins at the middle and thus the apical half of these veins as well as the apical half of the hindwing can reflex over their basal half; the area bordered by CuP and AP is full of cross-veins resulting from the presence of abundant branches of AP for supporting this area.
</paragraph>
</subSubSection>
<subSubSection id="2D936501FF87FF80FF584A8D8721C5B4" pageId="4" pageNumber="39" type="discussion">
<paragraph id="6536368AFF87FF80FF584A8D878BC7CA" blockId="4.[151,944,151,178]" box="[151,944,151,178]" pageId="4" pageNumber="39">
<heading id="3E7E81E6FF87FF80FF584A8D878BC7CA" bold="true" box="[151,944,151,178]" fontSize="11" level="1" pageId="4" pageNumber="39" reason="1">
<emphasis id="57FDEA98FF87FF80FF584A8D878BC7CA" bold="true" box="[151,944,151,178]" pageId="4" pageNumber="39">
Checklist of 
<taxonomicName id="A2894D09FF87FF80FEFC4A82858FC7C9" box="[307,436,151,177]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF87FF80FEFC4A82858FC7C9" bold="true" box="[307,436,151,177]" italics="true" pageId="4" pageNumber="39">Diploptera</emphasis>
</taxonomicName>
species (9) and subspecies (2) worldwide
</emphasis>
</heading>
</paragraph>
<paragraph id="6536368AFF87FF80FF584ACA8787C78E" blockId="4.[151,1409,222,607]" box="[151,956,222,247]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584ACA8787C78E" authority="Hanitsch, 1925: 102 Borneo" authorityName="Hanitsch, 1925: 102 Borneo" box="[151,956,222,247]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="57FDEA98FF87FF80FF584ACA8531C78E" box="[151,266,222,246]" italics="true" pageId="4" pageNumber="39">D. bicolor</emphasis>
<bibRefCitation id="01184B7BFF87FF80FEDE4ACB85C8C78F" author="Hanitsch, R." box="[273,499,222,247]" firstAuthor="Hanitsch" journalOrPublisher="Sarawak Museum Journal" page="102" pageId="4" pageNumber="39" pagination="75 - 106" part="8" refId="ref12545" refString="Hanitsch, R. (1925) On a collection of Blattidae from Northern Sarawak, chiefly Mt. Murud and Mt. Dulit. Sarawak Museum Journal, 8, 75 - 106." title="On a collection of Blattidae from Northern Sarawak, chiefly Mt. Murud and Mt. Dulit" type="journal article" year="1925">Hanitsch, 1925: 102</bibRefCitation>
Borneo
</taxonomicName>
</paragraph>
<paragraph id="6536368AFF87FF80FF584B17803BC663" blockId="4.[151,1409,222,607]" box="[151,1024,258,283]" pageId="4" pageNumber="39">
<emphasis id="57FDEA98FF87FF80FF584B17803BC663" bold="true" box="[151,1024,258,283]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584B17854EC663" box="[151,373,258,283]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="elliptica" status="sp. nov.">
<emphasis id="57FDEA98FF87FF80FF584B17854EC663" bold="true" box="[151,373,258,283]" italics="true" pageId="4" pageNumber="39">Diploptera elliptica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF87FF80FEB34B178580C663" box="[380,443,258,283]" pageId="4" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
South 
<collectingCountry id="1D9E761AFF87FF80FC794B17803BC663" box="[950,1024,258,283]" name="China" pageId="4" pageNumber="39">China</collectingCountry>
</emphasis>
</paragraph>
<paragraph id="6536368AFF87FF80FF584B328787C646" blockId="4.[151,1409,222,607]" box="[151,956,294,319]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584B328787C646" authority="Princis, 1950: 163 Borneo" authorityName="Princis, 1950: 163 Borneo" box="[151,956,294,319]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="erythrocephala">
<emphasis id="57FDEA98FF87FF80FF584B328558C646" box="[151,355,294,318]" italics="true" pageId="4" pageNumber="39">D. erythrocephala</emphasis>
<bibRefCitation id="01184B7BFF87FF80FEA54B33860CC647" author="Princis, K." box="[362,567,294,319]" firstAuthor="Princis" journalOrPublisher="Opuscula Entomologica" page="163" pageId="4" pageNumber="39" pagination="161 - 188" part="15" refId="ref12911" refString="Princis, K. (1950) Indomalaiische und australische Blattarien aus dem Entomologischen Museum der Universitat in Lund. Opuscula Entomologica, 15, 161 - 188." title="Indomalaiische und australische Blattarien aus dem Entomologischen Museum der Universitat in Lund" type="journal article" year="1950">Princis, 1950: 163</bibRefCitation>
Borneo
</taxonomicName>
</paragraph>
<paragraph id="6536368AFF87FF80FF584B5E8787C61A" blockId="4.[151,1409,222,607]" box="[151,956,330,355]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584B5E8787C61A" authority="Hanitsch, 1925: 104 Borneo" authorityName="Hanitsch, 1925: 104 Borneo" box="[151,956,330,355]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="57FDEA98FF87FF80FF584B5E851AC61B" box="[151,289,331,355]" italics="true" pageId="4" pageNumber="39">D. maculata</emphasis>
<bibRefCitation id="01184B7BFF87FF80FEE74B5F8630C61B" author="Hanitsch, R." box="[296,523,330,355]" firstAuthor="Hanitsch" journalOrPublisher="Sarawak Museum Journal" page="104" pageId="4" pageNumber="39" pagination="75 - 106" part="8" refId="ref12545" refString="Hanitsch, R. (1925) On a collection of Blattidae from Northern Sarawak, chiefly Mt. Murud and Mt. Dulit. Sarawak Museum Journal, 8, 75 - 106." title="On a collection of Blattidae from Northern Sarawak, chiefly Mt. Murud and Mt. Dulit" type="journal article" year="1925">Hanitsch, 1925: 104</bibRefCitation>
Borneo
</taxonomicName>
</paragraph>
<paragraph id="6536368AFF87FF80FF584B7A87DCC6FF" blockId="4.[151,1409,222,607]" box="[151,999,366,391]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584B7A84C7C6FE" box="[151,252,367,390]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="57FDEA98FF87FF80FF584B7A84C7C6FE" box="[151,252,367,390]" italics="true" pageId="4" pageNumber="39">D. minor</emphasis>
</taxonomicName>
(
<bibRefCitation id="01184B7BFF87FF80FEC54B7A86A0C6FF" author="Brunner von Wattenwyl, C." bookContentInfo="426 pp." box="[266,667,366,391]" firstAuthor="Brunner von Wattenwyl" journalOrPublisher="Vienna" page="295" pageId="4" pageNumber="39" refId="ref12296" refString="Brunner von Wattenwyl, C. (1865) Nouveau Systeme des Blattaires. Vienna, 426 pp." title="Nouveau Systeme des Blattaires" type="book" year="1865">Brunner von Wattenwyl, 1865: 295</bibRefCitation>
, 
<taxonomicName id="A2894D09FF87FF80FD6A4B7B8715C6FE" box="[677,814,366,390]" class="Insecta" family="Blaberidae" genus="Eleutheroda" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF87FF80FD6A4B7B8715C6FE" box="[677,814,366,390]" italics="true" pageId="4" pageNumber="39">Eleutheroda</emphasis>
</taxonomicName>
) 
<collectingCountry id="1D9E761AFF87FF80FCA64B7B87DCC6FF" box="[873,999,366,391]" name="Philippines" pageId="4" pageNumber="39">Philippines</collectingCountry>
</paragraph>
<paragraph id="6536368AFF87FF80FF584B87808DC6D3" blockId="4.[151,1409,222,607]" box="[151,1206,402,427]" pageId="4" pageNumber="39">
<emphasis id="57FDEA98FF87FF80FF584B87808DC6D3" bold="true" box="[151,1206,402,427]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584B878551C6D2" box="[151,362,402,427]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="naevus" status="sp. nov.">
<emphasis id="57FDEA98FF87FF80FF584B878551C6D2" bold="true" box="[151,362,402,427]" italics="true" pageId="4" pageNumber="39">Diploptera naevus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF87FF80FEBE4B87858BC6D3" box="[369,432,402,427]" pageId="4" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
<collectingCountry id="1D9E761AFF87FF80FCA64B878789C6D3" box="[873,946,402,427]" name="China" pageId="4" pageNumber="39">China</collectingCountry>
(southeast of Xizang)
</emphasis>
</paragraph>
<paragraph id="6536368AFF87FF80FF584BA2803BC6B7" blockId="4.[151,1409,222,607]" box="[151,1024,438,463]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF584BA2863CC6B7" authority="Shiraki, 1931: 174" authorityName="Shiraki" authorityPageNumber="174" authorityYear="1931" box="[151,519,438,463]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="nigrescens">
<emphasis id="57FDEA98FF87FF80FF584BA2850BC6B6" box="[151,304,439,462]" italics="true" pageId="4" pageNumber="39">D. nigrescens</emphasis>
<bibRefCitation id="01184B7BFF87FF80FEF84BA3863CC6B7" author="Shiraki, T." box="[311,519,438,463]" firstAuthor="Shiraki" journalOrPublisher="Insecta Matsumurana" page="174" pageId="4" pageNumber="39" pagination="171 - 209" part="5" refId="ref13277" refString="Shiraki, T. (1931) Orthoptera of the Japanese Empire. Part II. (Blattidae). Insecta Matsumurana, 5 (4), 171 - 209." title="Orthoptera of the Japanese Empire. Part II. (Blattidae)" type="journal article" year="1931">Shiraki, 1931: 174</bibRefCitation>
</taxonomicName>
<emphasis id="57FDEA98FF87FF80FCA64BA3803BC6B7" bold="true" box="[873,1024,438,463]" pageId="4" pageNumber="39">
South 
<collectingCountry id="1D9E761AFF87FF80FC794BA3803BC6B7" box="[950,1024,438,463]" name="China" pageId="4" pageNumber="39">China</collectingCountry>
</emphasis>
</paragraph>
<paragraph id="6536368AFF87FF80FF084BCF80B0C68B" blockId="4.[151,1409,222,607]" box="[199,1163,474,499]" pageId="4" pageNumber="39">
<emphasis id="57FDEA98FF87FF80FF084BCF80B0C68B" bold="true" box="[199,1163,474,499]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF084BCF8635C68B" box="[199,526,474,499]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="nigrescens" status="subsp. nov." subSpecies="guani">
<emphasis id="57FDEA98FF87FF80FF084BCF8635C68B" bold="true" box="[199,526,474,499]" italics="true" pageId="4" pageNumber="39">Diploptera nigrescens guani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF87FF80FDDB4BCF8645C68B" box="[532,638,474,499]" pageId="4" pageNumber="39" rank="subSpecies">subsp. n.</taxonomicNameLabel>
South 
<collectingCountry id="1D9E761AFF87FF80FC794BCF87C4C68B" box="[950,1023,474,499]" name="China" pageId="4" pageNumber="39">China</collectingCountry>
(mainland)
</emphasis>
</paragraph>
<paragraph id="6536368AFF87FF80FF084BEA80F8C56F" blockId="4.[151,1409,222,607]" box="[199,1219,510,535]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF084BEA868FC56F" authority="Shiraki, 1931: 174" authorityName="Shiraki" authorityPageNumber="174" authorityYear="1931" box="[199,692,510,535]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="nigrescens" subSpecies="nigrescens">
<emphasis id="57FDEA98FF87FF80FF084BEA85E5C56E" box="[199,478,511,534]" italics="true" pageId="4" pageNumber="39">D. nigrescens nigrescens</emphasis>
<bibRefCitation id="01184B7BFF87FF80FE2B4BEB868FC56F" author="Shiraki, T." box="[484,692,510,535]" firstAuthor="Shiraki" journalOrPublisher="Insecta Matsumurana" page="174" pageId="4" pageNumber="39" pagination="171 - 209" part="5" refId="ref13277" refString="Shiraki, T. (1931) Orthoptera of the Japanese Empire. Part II. (Blattidae). Insecta Matsumurana, 5 (4), 171 - 209." title="Orthoptera of the Japanese Empire. Part II. (Blattidae)" type="journal article" year="1931">Shiraki, 1931: 174</bibRefCitation>
</taxonomicName>
<emphasis id="57FDEA98FF87FF80FCA64BEB80F8C56F" bold="true" box="[873,1219,510,535]" pageId="4" pageNumber="39">
South 
<collectingCountry id="1D9E761AFF87FF80FC794BEB87C4C56F" box="[950,1023,510,535]" name="China" pageId="4" pageNumber="39">China</collectingCountry>
(
<collectingCountry id="1D9E761AFF87FF80FBC14BEB8053C56F" box="[1038,1128,510,535]" name="Taiwan" pageId="4" pageNumber="39">Taiwan</collectingCountry>
<collectingCountry id="1D9E761AFF87FF80FBA14BEB8080C56F" box="[1134,1211,510,535]" name="Iceland" pageId="4" pageNumber="39">Island</collectingCountry>
)
</emphasis>
</paragraph>
<paragraph id="6536368AFF87FF80FF5848368014C543" blockId="4.[151,1409,222,607]" box="[151,1071,546,571]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF58483685F5C543" authority="Princis, 1953: 208" authorityName="Princis" authorityPageNumber="208" authorityYear="1953" box="[151,462,546,571]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="57FDEA98FF87FF80FF58483684C1C542" box="[151,250,547,570]" italics="true" pageId="4" pageNumber="39">D. parva</emphasis>
<bibRefCitation id="01184B7BFF87FF80FECE483785F5C543" author="Princis, K." box="[257,462,546,571]" firstAuthor="Princis" journalOrPublisher="Entomologisk Tidskrift" page="208" pageId="4" pageNumber="39" pagination="203 - 213" part="74" refId="ref12940" refString="Princis, K. (1953) Kleine Beitrage zur Kenntnis der Blattarien und ihrer Verbreitung. VI. Entomologisk Tidskrift, 74 (4), 203 - 213." title="Kleine Beitrage zur Kenntnis der Blattarien und ihrer Verbreitung. VI" type="journal article" year="1953">Princis, 1953: 208</bibRefCitation>
</taxonomicName>
Java and Sumatra
</paragraph>
<paragraph id="6536368AFF87FF80FF58485281BAC527" blockId="4.[151,1409,222,607]" box="[151,1409,582,607]" pageId="4" pageNumber="39">
<taxonomicName id="A2894D09FF87FF80FF5848528520C526" box="[151,283,583,606]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF87FF80FF5848528520C526" box="[151,283,583,606]" italics="true" pageId="4" pageNumber="39">D. punctata</emphasis>
</taxonomicName>
(
<bibRefCitation id="01184B7BFF87FF80FEE54853864FC526" author="Eschscholtz, J. F. von" bookContentInfo="128 pp." box="[298,628,582,607]" journalOrPublisher="G. Reimer, Berlin" pageId="4" pageNumber="39" refId="ref12361" refString="Eschscholtz, J. F. von (1822) Entomographien. G. Reimer, Berlin, 128 pp. http: // dx. doi. org / 10.5962 / bhl. title. 5121" title="Entomographien" type="book" year="1822">
Eschscholtz, 1822: 86, 
<taxonomicName id="A2894D09FF87FF80FDFF4853864FC526" box="[560,628,582,606]" class="Insecta" family="Blattidae" genus="Blatta" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF87FF80FDFF4853864FC526" box="[560,628,582,606]" italics="true" pageId="4" pageNumber="39">Blatta</emphasis>
</taxonomicName>
</bibRefCitation>
) South Pacific, Southeast Asia and 
<emphasis id="57FDEA98FF87FF80FB24485381BAC527" bold="true" box="[1259,1409,582,607]" pageId="4" pageNumber="39">
South 
<collectingCountry id="1D9E761AFF87FF80FAF7485381BAC527" box="[1336,1409,582,607]" name="China" pageId="4" pageNumber="39">China</collectingCountry>
</emphasis>
</paragraph>
<paragraph id="6536368AFF87FF80FF5848A78721C5B4" blockId="4.[151,794,690,716]" box="[151,794,690,716]" pageId="4" pageNumber="39">
<heading id="3E7E81E6FF87FF80FF5848A78721C5B4" bold="true" box="[151,794,690,716]" fontSize="11" level="1" pageId="4" pageNumber="39" reason="1">
<emphasis id="57FDEA98FF87FF80FF5848A78721C5B4" bold="true" box="[151,794,690,716]" pageId="4" pageNumber="39">
Key to 
<taxonomicName id="A2894D09FF87FF80FF3F48A7854AC5B4" box="[240,369,690,716]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="57FDEA98FF87FF80FF3F48A7854AC5B4" bold="true" box="[240,369,690,716]" italics="true" pageId="4" pageNumber="39">Diploptera</emphasis>
</taxonomicName>
species and subspecies worldwide
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="2D936501FF87FF80FF5848EC81A0C279" pageId="4" pageNumber="39" type="key">
<keyStep id="DE7D2E2FFF87FF80FF5848EC81A0C453" pageId="4" pageNumber="39">
<paragraph id="6536368AFF87FF80FF5848EC81A0C476" blockId="4.[151,1436,761,1281]" box="[151,1435,761,782]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF5848EC81A0C476" box="[151,1435,761,782]" pageId="4" pageNumber="39">
1. Pronotum more or less suboval, without sharp posterolateral angles (
<figureCitation id="FDB22A0FFF87FF80FCBC48EC87D2C476" box="[883,1001,761,782]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="4" pageNumber="39">Figs. 13–16</figureCitation>
)................................... 2
</keyLead>
</paragraph>
<paragraph id="6536368AFF87FF80FF58490381A0C453" blockId="4.[151,1436,761,1281]" box="[151,1435,790,811]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF58490381A0C453" box="[151,1435,790,811]" pageId="4" pageNumber="39">
- Pronotum subsemicircular or subtrapezoidal, with sharp posterolateral angles (
<figureCitation id="FDB22A0FFF87FF80FC1D49038002C453" box="[978,1081,790,811]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="4" pageNumber="39">Figs. 2–12</figureCitation>
)............................ 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE7D2E2FFF87FF80FF58492181A0C41E" pageId="4" pageNumber="39">
<paragraph id="6536368AFF87FF80FF58492181A7C431" blockId="4.[151,1436,761,1281]" box="[151,1436,819,841]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF58492181A7C431" box="[151,1436,819,841]" pageId="4" pageNumber="39">
2. Pronotum unicolor in brown (
<figureCitation id="FDB22A0FFF87FF80FE314921864AC431" box="[510,625,820,841]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="4" pageNumber="39">Figs. 15–16</figureCitation>
).............................................. 
<taxonomicName id="A2894D09FF87FF80FB6E49268164C431" ID-CoL="6CXTM" box="[1185,1375,819,841]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="elliptica" status="sp. nov.">
<emphasis id="57FDEA98FF87FF80FB6E49268164C431" box="[1185,1375,819,841]" italics="true" pageId="4" pageNumber="39">Diploptera elliptica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF87FF80FAA9492681A7C431" box="[1382,1436,819,841]" pageId="4" pageNumber="39" rank="species">
<emphasis id="57FDEA98FF87FF80FAA9492681A7C431" bold="true" box="[1382,1436,819,841]" pageId="4" pageNumber="39">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="6536368AFF87FF80FF58494481A0C41E" blockId="4.[151,1436,761,1281]" box="[151,1435,849,870]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF58494481A0C41E" box="[151,1435,849,870]" pageId="4" pageNumber="39">
- Pronotum with markings in blackish and orange-yellowish (
<figureCitation id="FDB22A0FFF87FF80FCEF494487AEC41E" box="[800,917,849,870]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="4" pageNumber="39">Figs. 13–14</figureCitation>
).......................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE7D2E2FFF87FF80FF58497B81A0C4D9" pageId="4" pageNumber="39">
<paragraph id="6536368AFF87FF80FF58497B81A1C4FB" blockId="4.[151,1436,761,1281]" box="[151,1434,877,899]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF58497B81A1C4FB" box="[151,1434,877,899]" pageId="4" pageNumber="39">
3. Pronotum with disc blackish and lateral borders orange-yellowish (
<figureCitation id="FDB22A0FFF87FF80FCAB497B8792C4FB" box="[868,937,878,899]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="4" pageNumber="39">Fig. 13</figureCitation>
)................................. 
<taxonomicName id="A2894D09FF87FF80FAFA497B81A1C4FB" ID-CoL="6D9TL" box="[1333,1434,877,899]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="57FDEA98FF87FF80FAFA497B81A1C4FB" box="[1333,1434,877,899]" italics="true" pageId="4" pageNumber="39">D. bicolor</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="6536368AFF87FF80FF58499981A0C4D9" blockId="4.[151,1436,761,1281]" box="[151,1435,907,929]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF58499981A0C4D9" box="[151,1435,907,929]" pageId="4" pageNumber="39">
- Pronotum orange-yellowish, striped with a pair of vittae and a macula in blackish (
<figureCitation id="FDB22A0FFF87FF80FC3549998004C4D9" box="[1018,1087,908,929]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="4" pageNumber="39">Fig. 14</figureCitation>
).................. 
<taxonomicName id="A2894D09FF87FF80FAEA499981A0C4D9" ID-CoL="6CXTN" box="[1317,1435,907,929]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="maculata">
<emphasis id="57FDEA98FF87FF80FAEA499981A0C4D9" box="[1317,1435,907,929]" italics="true" pageId="4" pageNumber="39">D. maculata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE7D2E2FFF87FF80FF5849BC81A0C4A3" pageId="4" pageNumber="39">
<paragraph id="6536368AFF87FF80FF5849BC81A7C4C6" blockId="4.[151,1436,761,1281]" box="[151,1436,936,958]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF5849BC81A7C4C6" box="[151,1436,936,958]" pageId="4" pageNumber="39">
4. Pronotum with disc blackish and wide lateral borders orangish (
<figureCitation id="FDB22A0FFF87FF80FC8A49BC8745C4C6" box="[837,894,937,958]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="4" pageNumber="39">Fig. 9</figureCitation>
)........................ 
<taxonomicName id="A2894D09FF87FF80FB6149BD815BC4C6" ID-CoL="6D9TN" box="[1198,1376,936,958]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="naevus" status="sp. nov.">
<emphasis id="57FDEA98FF87FF80FB6149BD815BC4C6" box="[1198,1376,936,958]" italics="true" pageId="4" pageNumber="39">Diploptera naevus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF87FF80FAA949BD81A7C4C6" box="[1382,1436,936,958]" pageId="4" pageNumber="39" rank="species">
<emphasis id="57FDEA98FF87FF80FAA949BD81A7C4C6" bold="true" box="[1382,1436,936,958]" pageId="4" pageNumber="39">sp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="6536368AFF87FF80FF5849D381A0C4A3" blockId="4.[151,1436,761,1281]" box="[151,1435,966,987]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF5849D381A0C4A3" box="[151,1435,966,987]" pageId="4" pageNumber="39">
- Pronotum unicolor (
<figureCitation id="FDB22A0FFF87FF80FE6D49D385C4C4A3" box="[418,511,966,987]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="4" pageNumber="39">Figs. 2–8</figureCitation>
), or limbate narrowly (
<figureCitation id="FDB22A0FFF87FF80FD1349D38774C4A3" box="[732,847,966,987]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="4" pageNumber="39">Figs. 10–12</figureCitation>
)............................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE7D2E2FFF87FF80FF5849F181A7C36E" pageId="4" pageNumber="39">
<paragraph id="6536368AFF87FF80FF5849F181A7C481" blockId="4.[151,1436,761,1281]" box="[151,1436,996,1017]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF5849F181A7C481" box="[151,1436,996,1017]" pageId="4" pageNumber="39">
5. Pronotum blackish, limbate in reddish narrowly (
<figureCitation id="FDB22A0FFF87FF80FD7749F18710C481" box="[696,811,996,1017]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="4" pageNumber="39">Figs. 10–12</figureCitation>
)......................................(
<taxonomicName id="A2894D09FF87FF80FACA49F181B3C481" ID-CoL="6CXTP" box="[1285,1416,996,1017]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="nigrescens">
<emphasis id="57FDEA98FF87FF80FACA49F181B3C481" box="[1285,1416,996,1017]" italics="true" pageId="4" pageNumber="39">D. nigrescens</emphasis>
</taxonomicName>
)6
</keyLead>
</paragraph>
<paragraph id="6536368AFF87FF80FF584E1481A7C36E" blockId="4.[151,1436,761,1281]" box="[151,1436,1025,1046]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF584E1481A7C36E" box="[151,1436,1025,1046]" pageId="4" pageNumber="39">
- Pronotum unicolor in brownish (
<figureCitation id="FDB22A0FFF87FF80FDD54E14864EC36E" box="[538,629,1025,1046]" captionStart="FIGURES 2 – 16" captionStartId="6.[151,264,954,977]" captionTargetBox="[160,1418,196,938]" captionTargetId="figure@6.[160,1420,193,940]" captionTargetPageId="6" captionText="FIGURES 2 – 16. Pronota of Diploptera spp. &amp; subspp. presented in monochrome, i. e., white indicates unicolor or lighter coloration, whilst black is for darker portion. 2 – 3 D. punctata; 2 male; 3 female; 4 – 5 D. minor; 4 female; 5 male; 6 D. erythrocephala, male; 7 – 8 D. parva; 7 male; 8 female; 9 D. naevus sp. n., male; 10 D. nigrescens nigrescens, male; 11 – 12 D. nigrescens guani subsp. n.; 11 male; 12 female; 13 D. bicolor, male; 14 D. maculata, female; 15 – 16 D. elliptica sp. n.; 15 male; 16 female. (Dashed lines indicate depression. Scale bar = 2 mm)" httpUri="https://zenodo.org/record/289450/files/figure.png" pageId="4" pageNumber="39">Figs. 2–8</figureCitation>
)............................................ (
<taxonomicName id="A2894D09FF87FF80FB6E4E1480CEC36E" ID-CoL="6CXF3" box="[1185,1269,1025,1046]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF87FF80FB6E4E1480CEC36E" box="[1185,1269,1025,1046]" italics="true" pageId="4" pageNumber="39">punctata</emphasis>
</taxonomicName>
-species-group) 7
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE7D2E2FFF87FF80FF584E0B81A7C329" pageId="4" pageNumber="39">
<paragraph id="6536368AFF87FF80FF584E0B81A1C34B" blockId="4.[151,1436,761,1281]" box="[151,1434,1054,1075]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF584E0B81A1C34B" box="[151,1434,1054,1075]" pageId="4" pageNumber="39">
6. Head blackish (
<figureCitation id="FDB22A0FFF87FF80FEB94E0B85D0C34B" box="[374,491,1054,1075]" captionStart="FIGURES 17 – 30" captionStartId="7.[151,264,1796,1819]" captionTargetBox="[191,1384,195,1758]" captionTargetId="figure@7.[191,1385,195,1760]" captionTargetPageId="7" captionText="FIGURES 17 – 30. Diploptera spp. &amp; subspp. distributed in China. 17 – 18 D. punctata, female from Yunnan, Menglun; 19 – 21 D. punctata, lectotype (ZMMU) from Hawaii instead of China, photographed by Dr. Andrey L. Ozerov, copyright ZMMU, published with permission; 22 – 24 D. elliptica sp. n.; 22 – 23 male holotype from Yunnan, Cangyuan, Banhong; 24 female paratype from Yunnan, Mengla, Mohan; 25 – 26 D. naevus sp. n., male holotype from Xizang, Motuo, Beibeng; 27 – 28 D. nigrescens guani subsp. n., male holotype from Guangxi, Chongzuo, Daqingshan; 29 – 30 D. nigrescens nigrescens, male syntype (NTU) from Taiwan, photographed by Yi-Ho Liu and downloaded from NTU website, copyright Insect Museum Digital Archives Project, Department of Entomology, National Taiwan University. (Scale bars = 5 mm.)" httpUri="https://zenodo.org/record/289451/files/figure.png" pageId="4" pageNumber="39">Figs. 29–30</figureCitation>
, 
<figureCitation id="FDB22A0FFF87FF80FE3B4E0B8634C34B" box="[500,527,1054,1075]" captionStart="FIGURES 54 – 56. 54 D" captionStartId="13.[151,264,1787,1810]" captionTargetBox="[151,1436,193,1766]" captionTargetId="figure@13.[151,1436,193,1766]" captionTargetPageId="13" captionText="FIGURES 54 – 56. 54 D. punctata eating lemon peel in Hawaii, 2006. XI. 29, image by Forest &amp; Kim Starr (University of Hawaii); 55 Cypress with damage by D. punctata in Hawaii, 2005. VII. 11, image by Forest &amp; Kim Starr (University of Hawaii); 56 D. nigrescens nigrescens Shiraki, 1931 from Taiwan, Chiayi County, Tashan (= Xiaotashan), about 1200 m, photographed by Gaga (Taiwan). The insect was found near a light trap but not on the curtain." httpUri="https://zenodo.org/record/289456/files/figure.png" pageId="4" pageNumber="39">56</figureCitation>
), locality 
<collectingCountry id="1D9E761AFF87FF80FDA14E0B868FC34B" box="[622,692,1054,1075]" name="Taiwan" pageId="4" pageNumber="39">Taiwan</collectingCountry>
<collectingCountry id="1D9E761AFF87FF80FD754E0B86CEC34B" box="[698,757,1054,1075]" name="Iceland" pageId="4" pageNumber="39">Island</collectingCountry>
.................................... 
<taxonomicName id="A2894D09FF87FF80FB634E0B81A1C34B" ID-CoL="7GRF4" box="[1196,1434,1054,1075]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="nigrescens" subSpecies="nigrescens">
<emphasis id="57FDEA98FF87FF80FB634E0B81A1C34B" box="[1196,1434,1054,1075]" italics="true" pageId="4" pageNumber="39">D. nigrescens nigrescens</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="6536368AFF87FF80FF584E2981A7C329" blockId="4.[151,1436,761,1281]" box="[151,1436,1083,1105]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF584E2981A7C329" box="[151,1436,1083,1105]" pageId="4" pageNumber="39">
- Head reddish brown (
<figureCitation id="FDB22A0FFF87FF80FE7F4E29861DC329" box="[432,550,1084,1105]" captionStart="FIGURES 17 – 30" captionStartId="7.[151,264,1796,1819]" captionTargetBox="[191,1384,195,1758]" captionTargetId="figure@7.[191,1385,195,1760]" captionTargetPageId="7" captionText="FIGURES 17 – 30. Diploptera spp. &amp; subspp. distributed in China. 17 – 18 D. punctata, female from Yunnan, Menglun; 19 – 21 D. punctata, lectotype (ZMMU) from Hawaii instead of China, photographed by Dr. Andrey L. Ozerov, copyright ZMMU, published with permission; 22 – 24 D. elliptica sp. n.; 22 – 23 male holotype from Yunnan, Cangyuan, Banhong; 24 female paratype from Yunnan, Mengla, Mohan; 25 – 26 D. naevus sp. n., male holotype from Xizang, Motuo, Beibeng; 27 – 28 D. nigrescens guani subsp. n., male holotype from Guangxi, Chongzuo, Daqingshan; 29 – 30 D. nigrescens nigrescens, male syntype (NTU) from Taiwan, photographed by Yi-Ho Liu and downloaded from NTU website, copyright Insect Museum Digital Archives Project, Department of Entomology, National Taiwan University. (Scale bars = 5 mm.)" httpUri="https://zenodo.org/record/289451/files/figure.png" pageId="4" pageNumber="39">Figs. 27–28</figureCitation>
), locality south of mainland 
<collectingCountry id="1D9E761AFF87FF80FCF84E29874BC329" box="[823,880,1084,1105]" name="China" pageId="4" pageNumber="39">China</collectingCountry>
............... 
<taxonomicName id="A2894D09FF87FF80FBE54E2E8107C329" ID-CoL="7H3FC" box="[1066,1340,1083,1105]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="nigrescens" status="subsp. nov." subSpecies="guani">
<emphasis id="57FDEA98FF87FF80FBE54E2E8107C329" box="[1066,1340,1083,1105]" italics="true" pageId="4" pageNumber="39">Diploptera nigrescens guani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4CCE57E3FF87FF80FA8D4E2E81A7C329" box="[1346,1436,1083,1105]" pageId="4" pageNumber="39" rank="subSpecies">
<emphasis id="57FDEA98FF87FF80FA8D4E2E81A7C329" bold="true" box="[1346,1436,1083,1105]" pageId="4" pageNumber="39">subsp. n.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE7D2E2FFF87FF80FF584E4C81A0C3F3" pageId="4" pageNumber="39">
<paragraph id="6536368AFF87FF80FF584E4C81A7C316" blockId="4.[151,1436,761,1281]" box="[151,1436,1113,1134]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF584E4C81A7C316" box="[151,1436,1113,1134]" pageId="4" pageNumber="39">
7. Body relatively large, legs dark brown............................................................ 
<taxonomicName id="A2894D09FF87FF80FAE64E4C81A7C316" ID-CoL="6CXF3" box="[1321,1436,1113,1134]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="punctata">
<emphasis id="57FDEA98FF87FF80FAE64E4C81A7C316" box="[1321,1436,1113,1134]" italics="true" pageId="4" pageNumber="39">D. punctata</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="6536368AFF87FF80FF584E6381A0C3F3" blockId="4.[151,1436,761,1281]" box="[151,1435,1142,1163]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF584E6381A0C3F3" box="[151,1435,1142,1163]" pageId="4" pageNumber="39">- Body relatively small, legs (or except coxae) yellowish to reddish brown.......................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE7D2E2FFF87FF80FF584E8181A7C3BE" pageId="4" pageNumber="39">
<paragraph id="6536368AFF87FF80FF584E8181A0C3D1" blockId="4.[151,1436,761,1281]" box="[151,1435,1172,1193]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF584E8181A0C3D1" box="[151,1435,1172,1193]" pageId="4" pageNumber="39">8. Tegmina with distinct punctations......................................................................... 9</keyLead>
</paragraph>
<paragraph id="6536368AFF87FF80FF584EA481A7C3BE" blockId="4.[151,1436,761,1281]" box="[151,1436,1201,1222]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF584EA481A7C3BE" box="[151,1436,1201,1222]" pageId="4" pageNumber="39">
- Tegmina with very indistinct punctations............................................................. 
<taxonomicName id="A2894D09FF87FF80FA8A4EA481A7C3BE" ID-CoL="6CY5P" box="[1349,1436,1201,1222]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="parva">
<emphasis id="57FDEA98FF87FF80FA8A4EA481A7C3BE" box="[1349,1436,1201,1222]" italics="true" pageId="4" pageNumber="39">D. parva</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="DE7D2E2FFF87FF80FF584EDB81A0C279" pageId="4" pageNumber="39">
<paragraph id="6536368AFF87FF80FF584EDB81A7C39B" blockId="4.[151,1436,761,1281]" box="[151,1436,1229,1251]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF584EDB81A7C39B" box="[151,1436,1229,1251]" pageId="4" pageNumber="39">
9. Head reddish brown..................................................................... 
<taxonomicName id="A2894D09FF87FF80FB224EDB81A7C39B" ID-CoL="6CY5M" box="[1261,1436,1229,1251]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="erythrocephala">
<emphasis id="57FDEA98FF87FF80FB224EDB81A7C39B" box="[1261,1436,1229,1251]" italics="true" pageId="4" pageNumber="39">D. erythrocephala</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="6536368AFF87FF80FF584EF981A0C279" blockId="4.[151,1436,761,1281]" box="[151,1435,1260,1281]" pageId="4" pageNumber="39">
<keyLead id="DE7895BFFF87FF80FF584EF981A0C279" box="[151,1435,1260,1281]" pageId="4" pageNumber="39">
- Head black or blackish brown...................................................................... 
<taxonomicName id="A2894D09FF87FF80FA8E4EF981A0C279" ID-CoL="6CY5N" box="[1345,1435,1260,1281]" class="Insecta" family="Blaberidae" genus="Diploptera" kingdom="Animalia" order="Blattodea" pageId="4" pageNumber="39" phylum="Arthropoda" rank="species" species="minor">
<emphasis id="57FDEA98FF87FF80FA8E4EF981A0C279" box="[1345,1435,1260,1281]" italics="true" pageId="4" pageNumber="39">D. minor</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>