224 lines
23 KiB
XML
224 lines
23 KiB
XML
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<mods:title id="D8911B6964DA9CEA9918E44CE003060D">Status of the enigmatic Oriental genus Rhithrogeniella Ulmer, 1939 (Ephemeroptera, Heptageniidae)</mods:title>
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<mods:namePart id="99726A6893904E4F74980AF7AB228A62">Sartori, Michel</mods:namePart>
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<mods:date id="D2DBDBE1AABEF221AF1146AF22FE0013">2014</mods:date>
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<mods:classification id="996F9819A9B34E98E12B0F68C8542FAC">journal article</mods:classification>
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<mods:identifier id="5D2DEA5775C21597A2E7C380CC225759" type="DOI">http://dx.doi.org/10.3897/zookeys.429.8116</mods:identifier>
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<treatment id="EC87D9636F53EF803F3E86DAD2946C3C" ID-GBIF-Taxon="152054708" LSID="urn:lsid:plazi:treatment:EC87D9636F53EF803F3E86DAD2946C3C" httpUri="http://treatment.plazi.org/id/EC87D9636F53EF803F3E86DAD2946C3C" lastPageId="8" lastPageNumber="55" pageId="2" pageNumber="49" scope_family="Heptageniidae" scope_order="Ephemeroptera">
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<subSubSection id="25530C62C77BD8EE45639B213CF9286A" pageId="2" pageNumber="49" type="multiple">
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<paragraph id="9F7C7296AE7077342C090858258B0A11" pageId="2" pageNumber="49">Taxon classification Animalia Ephemeroptera Heptageniidae</paragraph>
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</subSubSection>
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<subSubSection id="513D5C6B8412E7718CCCD601462C0B3C" pageId="2" pageNumber="49" type="nomenclature">
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<paragraph id="DEFFE5D4D0BBE9A15B6E745E6AAE1305" pageId="2" pageNumber="49">
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<taxonomicName id="8BBB2498E413475EC550C4E887674868" ID-CoL="47NDC" authority="Ulmer, 1939" authorityName="Ulmer" authorityYear="1939" genus="Rhithrogeniella" lsidName="Rhithrogeniella ornata" pageId="2" pageNumber="49" rank="species" species="ornata">Rhithrogeniella ornata Ulmer, 1939</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="D972527EE5C477857042A149DAE241DF" pageId="2" pageNumber="49" type="reference_group">
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<paragraph id="5278598FB339FCB04907F5C2F4AEE5EE" pageId="2" pageNumber="49">
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<taxonomicName id="54DA6949468270152CD06C4FEEA0B044" genus="Rhithrogeniella" lsidName="Rhithrogeniella ornata" pageId="2" pageNumber="49" rank="species" species="ornata">Rhithrogeniella ornata</taxonomicName>
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Ulmer, 1939, male, female imagos and subimagos
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</paragraph>
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<paragraph id="E7AA88B84FA4310DEA42C2A57A7C0FE8" pageId="2" pageNumber="49">
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<taxonomicName id="3188E81499DD509CB71CAC7DD8DEBEF6" class="Insecta" family="Heptageniidae" genus="Rhithrogena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhithrogena ornata" order="Ephemeroptera" pageId="2" pageNumber="49" phylum="Arthropoda" rank="species" species="ornata">Rhithrogena ornata</taxonomicName>
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Wang & McCafferty, 2004
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</paragraph>
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</subSubSection>
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<subSubSection id="FE85B68769808CADD053AF247AF0164C" pageId="2" pageNumber="49" type="material">
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<paragraph id="C7B932C442C4DF6E9E643E5CB78FE3CA" pageId="2" pageNumber="49">Material.</paragraph>
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<paragraph id="119653783E7FCFB5D2DD179658E79028" pageId="2" pageNumber="49">One male holotype, one female allotype: Indonesia, Java, Buitenzorg, VII 1932, Dr. Lieftinck leg. [ZMH]</paragraph>
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<paragraph id="35E0A48551E3EF7E3415187A0783A15A" pageId="2" pageNumber="49">Paratypes: 4 female subimagos, 1 male subimago: Indonesia, Java, Buitenzorg, Bellevue, caught at light, VII.1929, Prof. Thienemann leg. [ZMH]; 4 female imagos, 2 male subimagos: Indonesia, Sumatra, Padang, VII 1925, Prof. Fulmek leg. [ZMH]; 1 male subimago: Indonesia, Sumatra, Pangkalang, Kota baru, X 1925, Prof. Fulmek leg. [ZMH]</paragraph>
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<paragraph id="EE68361C2E2035800C1C0DC5D7CFE68D" pageId="2" pageNumber="49">All specimens in ethanol, except fore- and hind legs, fore- and hind wings of the male subimago from Buitenzorg mounted on slide in Canada balsam.</paragraph>
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<paragraph id="3571BAA29D091ED06FDA37817F11DAB9" pageId="2" pageNumber="49">
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Other material: 5 nymphs: Indonesia, Sumatra Barat, Sawahlunto, stream, 275m,
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<geoCoordinate id="ECC7B11E5A9165B27C8E5433C07D95D2" direction="south" orientation="latitude" precision="9" value="-0.68883336">00°41.33'S</geoCoordinate>
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,
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<geoCoordinate id="D41298FE6AFAFEE5DF060F7C92A54280" direction="east" orientation="longitude" precision="9" value="100.77866">100°46.72'E</geoCoordinate>
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, (UN5), 10.XI.2011, M. Balke leg. [ZML]; 26 nymphs, of which two entirely mounted on microscopic slides: Indonesia, Sumatra Barat, Talawi, Ombilin River, 277m,
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<geoCoordinate id="36FA026B7AB00055AC7297162EEC9564" direction="south" orientation="latitude" precision="9" value="-0.5691667">00°34.15'S</geoCoordinate>
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,
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<geoCoordinate id="5BA1A3E9710B82392E99BD8DC959DB98" direction="east" orientation="longitude" precision="9" value="100.72567">100°43.54'E</geoCoordinate>
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, (UN4), 8.XI.2011, M. Balke leg. [ZMH, MZL, LIPI]
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</paragraph>
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</subSubSection>
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<subSubSection id="EB245A3E64D5A9C9C4D3B2569D03D187" pageId="2" pageNumber="49" type="complementary description of the male imago (holotype)">
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<paragraph id="8D6627E40FDFA4A06AB3F139C8FA6906" pageId="2" pageNumber="49">Complementary description of the male imago (holotype).</paragraph>
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<paragraph id="5BAB9069BA3108E6558F93F0C0827342" pageId="2" pageNumber="49">
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Specimen completely faded; for color patterns see
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<bibRefCitation id="50D50BEE659FA6E292A116C86DA2DE15" author="Ulmer, G" journalOrPublisher="Archiv fuer Hydrobiologie" pageId="11" pageNumber="58" pagination="443 - 692" title="Eintagsfliegen (Ephemeropteren) von den Sunda-Inseln." volume="16" year="1939">Ulmer (1939)</bibRefCitation>
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.
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</paragraph>
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<paragraph id="8EC3A9570395E6F7759A5FEB42ABB144" pageId="2" pageNumber="49">Mesonotum with transverse suture; medial depression of furcasternum sub parallel anteriorly.</paragraph>
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<paragraph id="E057B6D4ED5C7FE3156C82325ED2F9AB" pageId="2" pageNumber="49">Foreleg with tarsi sub equal in length to the tibia, which is 1.25x longer than the femur. Tarsal composition: 2>3>4>5>1.</paragraph>
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<paragraph id="0E38DE3E26458A1CBE381A50473C2001" pageId="2" pageNumber="49">
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Genitalia (Fig. 1): margin of the styliger fig straight to slightly convex, with two small sub-lateral rounded processes; last gonopod segment ca 0.7
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<normalizedToken id="0919CD6B33DB9B3CAA1EFF3AE136C360" originalValue="×">x</normalizedToken>
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the length of the previous, both together ca 0.75
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<normalizedToken id="0845E8C42D2F74F50960EBFB251A899D" originalValue="×">x</normalizedToken>
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the length of the antepenultimate. Penis constituted of two kidney-shape lobes, separated by a
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<normalizedToken id="9D4D914AA4298040868C3B2883BC740C" originalValue="“U”">"U"</normalizedToken>
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incision, i.e. the inner margin of each lobe is concave and slightly hooked near the apex. No lateral or median titillators, no apical spines visible.
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</paragraph>
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</subSubSection>
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<subSubSection id="1B3E524ACCD45D3C0D818345397CB831" lastPageId="4" lastPageNumber="51" pageId="3" pageNumber="50" type="complementary description of the male subimago">
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<paragraph id="BD94E9895FD2E486AD815F375C851645" pageId="3" pageNumber="50">
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<pageBreakToken id="A9984443BC645779E4C4318A551435E7" pageId="3" pageNumber="50" start="start">Complementary</pageBreakToken>
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description of the male subimago.
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</paragraph>
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<paragraph id="83605A27FA0283796EE80761028AC173" pageId="3" pageNumber="50">Fore leg (Fig. 2) with femur ca 1.15x the length of tibia, which is subequal in length to tarsi. Tarsal composition 4≥2>3≥1>5.</paragraph>
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<paragraph id="8BFD0CC789751391539B6B5F693C45A1" pageId="3" pageNumber="50">
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Hind leg (Fig. 3) with femur ca 1.35x the length of tibia, which is ca 1.45
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<normalizedToken id="787A110155E8B4139F17233DF09300E8" originalValue="×">x</normalizedToken>
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the length of tarsi. Tarsal composition 1=2=5>3≥4.
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</paragraph>
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<paragraph id="5F719BEF3BBA833FB0C721CF0F34DEBE" pageId="4" pageNumber="51">
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<pageBreakToken id="585518CF15E4A961ACF86A11506DEC7E" pageId="4" pageNumber="51" start="start">Genitalia</pageBreakToken>
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(Fig. 4) with penis lobes rounded, ellipsoid, without any spine or titillators; in median position, a pair of membranous processes ending with a spine like sclerotization present in ventral view.
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</paragraph>
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</subSubSection>
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<subSubSection id="6E3534F416EA082606ECC2626F789960" pageId="4" pageNumber="51" type="complementary description of the female imago (allotype)">
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<paragraph id="A3A40A1E546E4F25F7636AF667A142A4" pageId="4" pageNumber="51">Complementary description of the female imago (allotype).</paragraph>
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<paragraph id="58F9B7E01375BC795A1C256CC5800DD9" pageId="4" pageNumber="51">Thoracic structures similar to the male.</paragraph>
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<paragraph id="18D2F41CC21649F65304ED880AFF38FA" pageId="4" pageNumber="51">
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Eggs (Fig. 5): ovoid, ca 130
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<normalizedToken id="934B6B87B2DDEB035604D5A939B1A9DB" originalValue="µm">µm</normalizedToken>
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<normalizedToken id="2BB0EE9A2BBB090118DB3098E013B5A0" originalValue="×">x</normalizedToken>
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90
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<normalizedToken id="50012467E6654C2E17D9B910E2650341" originalValue="µm">µm</normalizedToken>
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; chorion regularly covered with hexagonal mesh ridges, with KCT in-between, not larger at poles; micropyle rounded to slightly oval in equatorial area.
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</paragraph>
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</subSubSection>
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<subSubSection id="B334EB4FF71BD378E4A3A85263D7F225" lastPageId="8" lastPageNumber="55" pageId="4" pageNumber="51" type="first description of the nymph">
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<paragraph id="BC780DB1DC95A33C2CAE067CAF80C3E0" pageId="4" pageNumber="51">First description of the nymph.</paragraph>
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<paragraph id="C4D3A629AA6611553BDC17232592E2F7" pageId="4" pageNumber="51">
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Size: Body length: up to 5.2 mm and 5.6 mm for male and female respectively; cerci and terminal filament subequal and ca
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<normalizedToken id="90A544B7BFCAB5BD46539A68FF385BE4" originalValue="¾">3/4</normalizedToken>
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the length of the body.
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</paragraph>
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<paragraph id="DDA8581A15BE2331BDE647E1199EEAEB" pageId="4" pageNumber="51">Coloration similar to Figs 6 and 7.</paragraph>
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<paragraph id="417FE0FBDA8F582FE6BBCF32A379D5F4" lastPageId="6" lastPageNumber="53" pageId="5" pageNumber="52">
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<pageBreakToken id="33B7BACAC3ED105459EFBE3430FC3D9E" pageId="5" pageNumber="52" start="start">Labrum</pageBreakToken>
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(Fig. 8) moderately expended laterally, ca 2.6
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<normalizedToken id="AC7181EBBFB7B7EF784DA80A33B21836" originalValue="×">x</normalizedToken>
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wider than long; lateral margins regularly rounded; no anteromedian emargination; dorsal face covered with long and thin setae anteriorly; ventral face with shorter and stout setae along the anterior margin. Mandibles covered with numerous long and thin setae on the outer margin; right mandible with outer incisor saw-like, inner one with a trifid apex with 2-3 pectinate setae below it, and 2-3 long and simple setae below the mola; left mandible with outer incisor saw-like, inner one with a bifid apex with 3-4 pectinate setae below it, and 3-4 long and simple setae below the mola. Maxillary palp three-segmented; first segment covered with thin setae on inner and outer margin; second segment with thin setae on the outer margin; third segment slightly pointed, only with long and thin setae. Maxillae with fimbriate scattered setae on the ventral surface (Fig. 13): 13-14 comb-shape setae on the crown of the galea, median ones with 10-11 teeth (Fig. 14); proximal dentiseta bifid, outer margin feathered; distal dentiseta simple, entire and unbranched (Fig. 12). Labium (Fig. 9) with glossae rhomboid, inner margin covered with long and thin setae, apex characteristic with scale-like margin (Fig. 10); paraglos
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<pageBreakToken id="6CEC7C52D86D3A06015636B0463630ED" pageId="6" pageNumber="53" start="start">sae</pageBreakToken>
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moderately expended laterally. Hypopharynx (Fig. 11) with rhomboid lingua bearing a tuft of short and thin setae at apex; superlinguae well developed and expended laterally with rounded apex and setae on the outer margin extended beyond the apex.
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</paragraph>
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<paragraph id="A90914408F6D33E807D7827FF1122D24" pageId="7" pageNumber="54">
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<pageBreakToken id="2C1E3C14345A0FBC6D710944B5760615" pageId="7" pageNumber="54" start="start">Pronotum</pageBreakToken>
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moderately expended laterally. Foreleg with femur ca 2.6
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<normalizedToken id="8DBCF486C2E764FBE84E604AABE4FE71" originalValue="×">x</normalizedToken>
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longer than wide; outer margin covered with long and stout setae, becoming thinner near the apex; inner margin with only few spine-like setae on the distal third. Outer margin of tibia with very few thin and short setae (Fig. 15), inner margin with few spine-like setae in the middle; tarsi with only a few spine-like setae in the middle of the inner margin. Hind leg similar, except the spine-like setae on inner margin of the femur present on the whole margin; outer margin of tibia with a row of long and thin setae (Fig. 16) and inner margin with more numerous spine-like setae. Middle leg similar to hind leg, except spine-like setae on the inner margin of the femur only present on the distal half. Bristles on the upper face of femora variable in length, always with divergent margins and rounded apically (Fig. 17). Tarsal claw moderately hooked, bearing 4-6 teeth (Fig. 18). No supracoxal spurs present.
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</paragraph>
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<paragraph id="52D8BFDFDCE4A36AE97A6171F6D1CB0B" lastPageId="8" lastPageNumber="55" pageId="7" pageNumber="54">
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Abdomen with posterolateral extensions weakly developed, visible only on segments
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<normalizedToken id="EE83B18DF6BA79C4BB968613BA0F4808" originalValue="V–VIII">V-VIII</normalizedToken>
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. Gills present on abdominal segments
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<normalizedToken id="C8AB9B84FBAF0E6326D3B1310F33BBCB" originalValue="I–VII">I-VII</normalizedToken>
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. Gill I banana-shape
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<pageBreakToken id="83B977DB9724150A1007F9AB241CF543" pageId="8" pageNumber="55" start="start">(</pageBreakToken>
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Fig. 20), with fibrillar part well developed, gill IV ca 1.5
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<normalizedToken id="18E3E8F487E6A14D53B4594749F7B455" originalValue="×">x</normalizedToken>
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longer than wide, strongly asymmetrical (Fig. 21), gill VI with well-developed fibrillar part, more elongated and slightly asymmetrical (Fig. 22), gill VII ca 2.5
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<normalizedToken id="802072CF3F401CF9011D0DC54C719044" originalValue="×">x</normalizedToken>
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longer than wide, without fibrillar part and slightly asymmetrical (Fig. 23). Posterior margin of abdominal terga with weakly developed spines of different size and shape (Fig. 19). Cerci and terminal filament with long and stout setae in whorls on the proximal part (Fig. 24), together with long and thin setae in the median and distal part (Fig. 25).
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</paragraph>
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</subSubSection>
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<subSubSection id="15FF10CFB4BDCF0EEF9217782B556CFE" pageId="8" pageNumber="55" type="sequence data">
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<paragraph id="0344004F57ED097C9BD2284BF9673225" pageId="8" pageNumber="55">Sequence data.</paragraph>
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<paragraph id="78C641CEA79155385ABFA1DA0B8AB57D" pageId="8" pageNumber="55">
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One specimen has been used for the study by
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<bibRefCitation id="9D77B97F4F20D66B68D9EED157192F7B" author="Vuataz, L" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="11" pageNumber="58" pagination="979 - 991" title="Endemism and diversification in freshwater insects of Madagascar revealed by coalescent and phylogenetic analysis of museum and field collections." volume="66" year="2013">Vuataz et al. (2013)</bibRefCitation>
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under the name "
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<taxonomicName id="21AA5A112F941AF4899124D10CD78199" class="Insecta" family="Heptageniidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Ephemeroptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="family">Heptageniidae</taxonomicName>
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1" in figures and "
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<taxonomicName id="6D876A3B6652B9F7D8F880CBFCA96F54" class="Insecta" family="Heptageniidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Ephemeroptera" pageId="8" pageNumber="55" phylum="Arthropoda" rank="family">Heptageniidae</taxonomicName>
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sp. 1" in table S1, with one mitochondrial (CO1) and two nuclear genes (H3, wg) sequenced. Access numbers in GenBank are for CO1: HF536605, for wg: HF536598, for H3: HF536591.
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</paragraph>
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<caption id="8326871C0EFA29759391E6A59EC83BAD" pageId="8" pageNumber="55">
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<paragraph id="E96A407065327702C968E13F0763F848" pageId="8" pageNumber="55">
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Figures 1-4.
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<taxonomicName id="34816B9B2B64D088314DDFC249B81319" genus="Rhithrogeniella" lsidName="Rhithrogeniella ornata" pageId="8" pageNumber="55" rank="species" species="ornata">Rhithrogeniella ornata</taxonomicName>
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||
Ulmer, 1939. 1 Genitalia of the male imago (holotype) in ventral view 2 Foreleg of a male subimago (paratype) 3 Hindleg of a male subimago (paratype) 4 Penis lobes of a male subimago (paratype): plain line, cuticular structures of the subimago; dotted line, outline of the imago penis lobes.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="01278A807B6F0A26914EAD490F9CEFAC" pageId="8" pageNumber="55">
|
||
<paragraph id="7793E33B664C8E1671605FD96A7C98FA" pageId="8" pageNumber="55">
|
||
Figure 5.
|
||
<taxonomicName id="AB1C259C07A663B40D3E3BB322D3D8D4" genus="Rhithrogeniella" lsidName="Rhithrogeniella ornata" pageId="8" pageNumber="55" rank="species" species="ornata">Rhithrogeniella ornata</taxonomicName>
|
||
Ulmer, 1939, SEM pictures of egg structures. 5a Egg extracted from a female subimago paratype from Padang, Sumatra 5b Details of the chorionic structure of a female nymph from Ombilin River, Sumatra 5c Details of the chorionic structure and micropyle of a female subimago paratype from Buitenzorg [Bogor], Java 5d chorionic surface of the female allotype from Buitenzorg [Bogor], Java.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="6B92BB88F26A3A95BA7D47F30B0D41E6" pageId="8" pageNumber="55">
|
||
<paragraph id="5D4B211F1B47A6AC37CE918D1AE3F948" pageId="8" pageNumber="55">
|
||
Figures 6-7.
|
||
<taxonomicName id="6CDEE1C3A622FA8C9A7B5990582F4CA6" genus="Rhithrogeniella" lsidName="Rhithrogeniella ornata" pageId="8" pageNumber="55" rank="species" species="ornata">Rhithrogeniella ornata</taxonomicName>
|
||
Ulmer, 1939. 6 Male nymph 7 Female nymph with slight color variations.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F3A016E83B836CB6CF16ABCCD22761FF" pageId="8" pageNumber="55">
|
||
<paragraph id="21C585BFCAEE7503E25A1ABE341E8DEF" pageId="8" pageNumber="55">
|
||
Figures 8-11.
|
||
<taxonomicName id="8340B8B3DF358F1C87419DDEB4A71D9A" genus="Rhithrogeniella" lsidName="Rhithrogeniella ornata" pageId="8" pageNumber="55" rank="species" species="ornata">Rhithrogeniella ornata</taxonomicName>
|
||
Ulmer, 1939, nymphal mouthparts. 8 Labrum in dorsal view 9 Left glossae and paraglossae of the labium 10 Detail of the glossae from 9 11 Hypopharynx, ventral view lingua and left superlingua.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B452B1B8E9C3CF068F51CF346638A11C" pageId="8" pageNumber="55">
|
||
<paragraph id="9ACF127429CCF4C64A16054B2E3F039C" pageId="8" pageNumber="55">
|
||
Figures 12-14.
|
||
<taxonomicName id="B915D7301482EC1700D9066EE7A6FDE8" genus="Rhithrogeniella" lsidName="Rhithrogeniella ornata" pageId="8" pageNumber="55" rank="species" species="ornata">Rhithrogeniella ornata</taxonomicName>
|
||
Ulmer, 1939, SEM pictures of the maxilla. 12 Dentisetae (DP: proximal dentiseta, DD: distal dentiseta) 13 Fimbriate setae on the ventral surface 14 Comb-shape setae on the crown of the galea-lacinia.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="6FBB01AF6333E5330380524675D0B0A8" pageId="8" pageNumber="55">
|
||
<paragraph id="42EFD0C62C5F73B0FBB13EC2CD067329" pageId="8" pageNumber="55">
|
||
Figures 15-19.
|
||
<taxonomicName id="788961D6ACBF60737B196551355F23B6" genus="Rhithrogeniella" lsidName="Rhithrogeniella ornata" pageId="8" pageNumber="55" rank="species" species="ornata">Rhithrogeniella ornata</taxonomicName>
|
||
Ulmer, 1939. 15 Outer margin of the fore tibia 16 Outer margin of the hind tibia 17 Bristles on the dorsal surface of hind femur 18 Tarsal claw 19 Posterior margin of tergite V.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A3D8D913A5471BB5F9F01D7F4D14868A" pageId="8" pageNumber="55">
|
||
<paragraph id="F41A6DA07314FC1FBAB301D08F4B8050" pageId="8" pageNumber="55">
|
||
Figures 20-23.
|
||
<taxonomicName id="5AADF6814651B7F44809BC0564583942" genus="Rhithrogeniella" lsidName="Rhithrogeniella ornata" pageId="8" pageNumber="55" rank="species" species="ornata">Rhithrogeniella ornata</taxonomicName>
|
||
Ulmer, 1939. 20 Gill I 21 Gill IV 22 Gill VI 23 Gill VII.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DDF7D5621D14C55781E495A4006E75CC" pageId="8" pageNumber="55">
|
||
<paragraph id="ADF95C2967258E9C6ED689DE95CAB1D2" pageId="8" pageNumber="55">
|
||
Figures 24-25.
|
||
<taxonomicName id="CF9E0A6C77C5A7B84690D5C79F4ACB83" genus="Rhithrogeniella" lsidName="Rhithrogeniella ornata" pageId="8" pageNumber="55" rank="species" species="ornata">Rhithrogeniella ornata</taxonomicName>
|
||
Ulmer, 1939. 24 Proximal part of the terminal filament 25 Median part of the terminal filament.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |