163 lines
18 KiB
XML
163 lines
18 KiB
XML
<document id="52ECB9F7B9D492F42FE79CE699F4B644" ID-DOI="http://dx.doi.org/10.3897/subtbiol.19.9513" ID-GBIF-Dataset="43a31cef-65e7-4370-9a9b-d41af3ea3715" ID-Pensoft-Pub="1314-2615-19-23" ID-ZooBank="47000C3AA8C44BC6BB4C063C46D20BD6" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1314-2615-19-23" ModsDocOrigin="Subterranean Biology 19" ModsDocTitle="A new interstitial species of diving beetle from tropical northern Australia provides a scenario for the transition of epigean to stygobitic life (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1469796010600" checkinUser="pensoft" docAuthor="Watts, Chris H. S., Hendrich, Lars & Balke, Michael" docDate="2016" docId="EBBA60E4B0A2E9126384F7118C2A4B3C" docLanguage="en" docName="SubterBiol 19: 23-29" docOrigin="Subterranean Biology 19" docSource="http://dx.doi.org/10.3897/subtbiol.19.9513" docTitle="Exocelina saltusholmesensis Watts, Hendrich & Balke, 2016, sp. n." docType="treatment" docUuid="90C3C890-B056-4AA3-9A3D-82CB91BA2D0B" docUuidSource="ZooBank" docVersion="5" lastPageNumber="26" masterDocId="4D4FA37CFFF9FFE8FFA9FFE93250FF90" masterDocTitle="A new interstitial species of diving beetle from tropical northern Australia provides a scenario for the transition of epigean to stygobitic life (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="29" masterPageNumber="23" pageNumber="24" updateTime="1701317736552" updateUser="plazi">
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<mods:title id="56F1F53C0B58295C05A4B287CC8EDBBF">A new interstitial species of diving beetle from tropical northern Australia provides a scenario for the transition of epigean to stygobitic life (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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<mods:namePart id="5A66CF1CF99E2620F40425767E10A875">Watts, Chris H. S.</mods:namePart>
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<mods:namePart id="D2D6799BFA5E3D5F6E2BF3381325A2E1">Hendrich, Lars</mods:namePart>
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<mods:roleTerm id="7574D40129750433E44B379A1376BCCB">Author</mods:roleTerm>
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<mods:namePart id="58A5B47EE4603255B22A8711C1E1C4EF">Balke, Michael</mods:namePart>
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<mods:title id="B0C7C621124252151A8AA23DBCF58294">Subterranean Biology</mods:title>
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<mods:part id="30ED0F13AF2D78443247D1F4E93A1D4A">
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<mods:date id="6427F1DE19CF3EBDC701E584A859EF93">2016</mods:date>
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<mods:number id="0178D73FF381FD5EAC236A5D92D1D74C">19</mods:number>
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<mods:url id="334C108048E490978E58BB6BD5BE3FCC">http://dx.doi.org/10.3897/subtbiol.19.9513</mods:url>
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<mods:classification id="981E7DC2D3D948FA68852B17062CF19F">journal article</mods:classification>
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<mods:identifier id="C876A519ADBAE7A2F13953B7A5C0B5CA" type="DOI">http://dx.doi.org/10.3897/subtbiol.19.9513</mods:identifier>
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<treatment id="EBBA60E4B0A2E9126384F7118C2A4B3C" ID-GBIF-Taxon="127902061" LSID="urn:lsid:zoobank.org:act:90C3C890-B056-4AA3-9A3D-82CB91BA2D0B" httpUri="http://treatment.plazi.org/id/EBBA60E4B0A2E9126384F7118C2A4B3C" lastPageId="3" lastPageNumber="26" pageId="1" pageNumber="24">
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<subSubSection id="8E408AB8B786C92440B66D007FC8B632" pageId="1" pageNumber="24" type="multiple">
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<paragraph id="954DEC0BD5590456A5248F9B3B4CC454" pageId="1" pageNumber="24">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection id="BFD18B2C74C59E287F3FD3E3642502FB" pageId="1" pageNumber="24" type="nomenclature">
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<paragraph id="837600D9A1C6D65998409C926A859098" pageId="1" pageNumber="24">
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<taxonomicName id="8505D6C0A86268A1CD42959701EAE793" ID-CoL="8LXT8" LSID="http://zoobank.org/90C3C890-B056-4AA3-9A3D-82CB91BA2D0B" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina saltusholmesensis" order="Coleoptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="saltusholmesensis">Exocelina saltusholmesensis</taxonomicName>
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<taxonomicNameLabel id="F6558854E89C44F54464531408A20806" pageId="1" pageNumber="24">sp. n.</taxonomicNameLabel>
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Figs 1, 3
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</paragraph>
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</subSubSection>
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<subSubSection id="1BD6088E3789D2BB9C2A8649B56A4BC4" pageId="1" pageNumber="24" type="type locality">
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<paragraph id="F2EA3D0CA81F0A8BDFC367A0EC1CF2AB" pageId="1" pageNumber="24">Type locality.</paragraph>
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<paragraph id="BD5C3A2FEB825CAB106BBC93D3125162" pageId="1" pageNumber="24">Small seasonal creek [-12.400714° 130.931297°], Holmes Jungle Reserve, Darwin, Northern Territory, Australia.</paragraph>
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</subSubSection>
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<subSubSection id="445C796614FE7295E95FD8690CBF9539" pageId="1" pageNumber="24" type="type material">
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<paragraph id="4785B3D4F485D898D083890F1118E9F2" pageId="1" pageNumber="24">Type material.</paragraph>
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<paragraph id="883DDA9FBA1367903A3CE360EE74C3A5" pageId="1" pageNumber="24">
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Holotype, female (South Australian Museum Adelaide, SAMA): "Australia, Northern Territory, Darwin, Holmes Jungle Reserve, -12.400714° 130.931297°, 25. March 2004, C.H.S. Watts (SAMA database no. 25-012926)". The beetle was originally labelled as "
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<taxonomicName id="03179AEFB2BE097AB0325172296354CE" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
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sp.nov." by C.H.S. Watts and has now a red, printed holotype label attached in addition.
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</paragraph>
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<paragraph id="9A37716A377F7566292A6A9594D6BD56" pageId="1" pageNumber="24">European nucleotide archive (genbank) accession number for the 3' end of the mitochondrial cytochrome c oxidase 1 gene: www.ebi.ac.uk/ena/data/view/LN994839</paragraph>
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</subSubSection>
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<subSubSection id="BED1A511B2EB0248CEE6A2377CBE4470" lastPageId="3" lastPageNumber="26" pageId="1" pageNumber="24" type="description">
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<paragraph id="CF9EF17069010D16D38FE5464811E00F" pageId="1" pageNumber="24">Description.</paragraph>
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<paragraph id="E55575EAB7D6B01257D650DE5EAD40B5" pageId="1" pageNumber="24">Size and shape: Beetle small (length of beetle 3.7 mm, greatest width 1.8 mm, length of pronotum 0.6 mm, width of pronotum at base 1.7 mm), comparably parallel-sided (Fig. 1C).</paragraph>
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<paragraph id="DC847E0050B50D64570932224D2E40AC" pageId="1" pageNumber="24">Coloration. Mainly testaceous and slightly translucent with lighter band on elytral base and darker area in middle basal area of pronotum (Fig. 1C).</paragraph>
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<paragraph id="14285FD9D3C1839D3FDA574A6BDBF129" pageId="2" pageNumber="25">
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<pageBreakToken id="34DD4E0DDA7A45BB8643F8716D067772" pageId="2" pageNumber="25" start="start">Surface</pageBreakToken>
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sculpture. Head (frons), pronotum and elytra striolate; with distinct mircoreticulation and coarser punctation. Metacoxal plates and ventrites 1-4 with few strioles, otherwise with fine mircoreticulation and few punctures. The sixth (apical) ventrite broadly rounded.
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</paragraph>
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<paragraph id="71BB75B9F22C3FB5BF97B12573C4C30E" lastPageId="3" lastPageNumber="26" pageId="2" pageNumber="25">
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Structures. Eyes small in comparison to epigean
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<taxonomicName id="F3D501CF473B708619AC43BE941165EC" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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, length of eye c. 0.16 mm, distance between eyes c. 0.86 mm (Figs 1C, 3B). Pronotum with faint lateral
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<pageBreakToken id="B15749109895ACA0F338E2665044AC9B" pageId="3" pageNumber="26" start="start">bead</pageBreakToken>
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that does not reach anterior corner. Prosternal process lanceolate, gently rounded ventrally; lateral wings of metaventrite narrow. Flight wings vestigial.
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</paragraph>
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<paragraph id="BD2CAB088F64A0AC09A74FE261428A56" pageId="3" pageNumber="26">Male. Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="408E1A86757397CB6B247529E7F9E1A8" pageId="3" pageNumber="26" type="etymology">
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<paragraph id="394097F31173FDE957E4F46C9D8037FB" pageId="3" pageNumber="26">Etymology.</paragraph>
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<paragraph id="D766FC1E07B88A1E25B6AFC089CD0898" pageId="3" pageNumber="26">The name is an adjective is the nominative singular derived from the locality name - Holmes Jungle. Latin: saltus = woodland.</paragraph>
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</subSubSection>
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<subSubSection id="6F81FC6E77CC4B5411304CD5019286F4" pageId="3" pageNumber="26" type="differential diagnosis">
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<paragraph id="1F0DDC5452D97279F2004A6A1582CAFD" pageId="3" pageNumber="26">Differential diagnosis.</paragraph>
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<paragraph id="14C1A5076BFE432BE35EA08993F2F06D" pageId="3" pageNumber="26">
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The species is sister to a clade containing
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<taxonomicName id="AD78FC7F8384D3F431A32483FDE4F225" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina punctipennis" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="punctipennis">Exocelina punctipennis</taxonomicName>
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(Lea, 1899) and
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<taxonomicName id="6306D8A8E94FBEA8D7EBF162DFA9138F" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ferruginea" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="ferruginea">Exocelina ferruginea</taxonomicName>
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(Sharp, 1882) established based on DNA sequence data (
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<bibRefCitation id="301959F5641F03F48A1D6578400F9523" author="Toussaint, EF" journalOrPublisher="Scientific Reports" pageId="4" pageNumber="27" pagination=": e 16016" title="Mosaic patterns of diversification dynamics following the colonization of Melanesian islands." url="10.1038/srep16016" volume="5" year="2015">Toussaint et al. 2015</bibRefCitation>
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), but both these species are larger (> 5mm), and have larger eyes (Figs 1
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<normalizedToken id="53FC71C66BDE83DD336EE8800C1AEDFB" originalValue="A–C">A-C</normalizedToken>
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, 3A, B). Both are epigean species, distributed along the eastcoast of Queensland (
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<taxonomicName id="D9C0331EE169A8D8E5E129DA3357FE95" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina punctipennis" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="punctipennis">Exocelina punctipennis</taxonomicName>
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) and the Adelaide Hills in South Australia (
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<taxonomicName id="C96F3AE7BEBB18DF11338EFE8B35092C" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ferruginea" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="ferruginea">Exocelina ferruginea</taxonomicName>
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) (Hendrich et al. in prep.). They are also fully winged.
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</paragraph>
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</subSubSection>
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<subSubSection id="C71FA8C7F0EB94D1AE6F4DEB0F14950E" pageId="3" pageNumber="26" type="habitat">
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<paragraph id="029983AFDE79D9EE89DAA84BA4C15DBF" pageId="3" pageNumber="26">Habitat.</paragraph>
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<paragraph id="197275065EC85E57FE241573895E1BA2" pageId="3" pageNumber="26">
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A small pool in the bed of a small ephemeral creek through eucalypt woodland. A small eyed, wingless
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<taxonomicName id="53A251BED4DCBC54695F633D0847D32E" family="Hydraenidae" lsidName="" pageId="3" pageNumber="26" rank="family">Hydraenidae</taxonomicName>
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(
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<taxonomicName id="A784A2BFCC1A6B7B34DA9E9217B9F475" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hydraena wattsi" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="wattsi">Hydraena wattsi</taxonomicName>
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Perkins, 2007) was collected at the same time (
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<bibRefCitation id="4BDFFF8BCE9B4AF167D19A99C49677B4" author="Perkins, PD" journalOrPublisher="Zootaxa" pageId="4" pageNumber="27" pagination="1 - 139" title="A revision of the Australian species of the water beetle genus Hydraena Kugelmann (Coleoptera: Hydraenidae)." volume="1489" year="2007">Perkins 2007</bibRefCitation>
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). During the dry season (
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<normalizedToken id="4E5B3FDE6091F919186667DBA1DF9FC2" originalValue="June–September">June-September</normalizedToken>
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) this creek dries out. However, we assume that subsurface water remains in the watercourse during this season providing an interstitial habitat for these flightless, low vagility species.
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</paragraph>
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<caption id="41905A8606F69DA470F69ACAC6C676F5" pageId="3" pageNumber="26">
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<paragraph id="AD5559D072F0FB7B7D715ADC270A92E2" pageId="3" pageNumber="26">
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Figure 1. Habitus of
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<taxonomicName id="EC7FAE105D11D23506B879E50AD3C312" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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species:
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<taxonomicName id="17D65BB74941EA8A8333A3F6F8735D33" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ferruginea" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="ferruginea">Exocelina ferruginea</taxonomicName>
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(A),
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<taxonomicName id="E883B87CA23E279A4A5A8F6C3C5DD9FF" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina punctipennis" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="punctipennis">Exocelina punctipennis</taxonomicName>
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(B),
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<taxonomicName id="8D6C6B38F439A98C1AE499A40E5C64C1" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina saltusholmesensis" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="saltusholmesensis">Exocelina saltusholmesensis</taxonomicName>
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sp. n. (C).
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</paragraph>
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</caption>
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<caption id="D10D7FCC0F5AC7B57C516F63A27738F8" pageId="3" pageNumber="26">
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<paragraph id="6349B1182D8CB4536D0AB5BBE8022D1A" pageId="3" pageNumber="26">
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Figure 2. Habitus of
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<taxonomicName id="5E7839B412854318FE23D4D09C2B7F92" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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species:
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<taxonomicName id="99A47D70A81BBE5822C494D07D0BA5FF" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina australis" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="australis">Exocelina australis</taxonomicName>
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(A),
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<taxonomicName id="56FA5D2C229F90CCD62CAF69B23CCCFC" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina abdita" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="abdita">Exocelina abdita</taxonomicName>
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(B).
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</paragraph>
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</caption>
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<caption id="F591F050CEC8A8FB9A2876BA02E3C3D4" pageId="3" pageNumber="26">
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<paragraph id="06346810A870CF7BDE3E9B0E14A31718" pageId="3" pageNumber="26">
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Figure 3. Right lateral aspect of head of
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<taxonomicName id="1F01277F8BF9DE0F2FD013FDEF181949" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">Exocelina</taxonomicName>
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species:
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<taxonomicName id="DEB42060D0088A2BBFBAF50BD7177EBF" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina ferruginea" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="ferruginea">Exocelina ferruginea</taxonomicName>
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(A), E.
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<taxonomicName id="66C6BA020528C463A66D43ED2C0A3F63" lsidName="saltusholmesensis" pageId="3" pageNumber="26" rank="species" species="saltusholmesensis">saltusholmesensis</taxonomicName>
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sp. n. (B),
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<taxonomicName id="F0F135C574DF53C456068D91CE866F6F" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina abdita" order="Coleoptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="abdita">Exocelina abdita</taxonomicName>
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(C).
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |