treatments-xml/data/EB/7A/87/EB7A87EBA76F4C7132BFF99B43686CE8.xml

<document id="F2366AF875B975B7ED34304186047725" ID-CLB-Dataset="46854" ID-DOI="10.11646/zootaxa.4072.3.5" ID-GBIF-Dataset="25f4af80-2b6f-47e3-94ad-49bcbe074c06" ID-ISSN="1175-5326" ID-Zenodo-Dep="255244" ID-ZooBank="8BE57C06-DBD3-40EA-9107-4E6959AD1247" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460358389516" checkinUser="plazi" docAuthor="Li, Yang &amp; Xu, Kuidong" docDate="2016" docId="EB7A87EBA76F4C7132BFF99B43686CE8" docLanguage="en" docName="zootaxa.4072.3.5.pdf" docOrigin="Zootaxa 4072 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphelliactis tangi Li &amp; Xu, 2016, n. sp." docType="treatment" docVersion="8" lastPageNumber="365" masterDocId="1743FF93A76D4C763228F947411C6D60" masterDocTitle="Paraphelliactis tangi n. sp. and Phelliactis yapensis n. sp., two new deep-sea species of Hormathiidae (Cnidaria: Anthozoa: Actiniaria) from a seamount in the tropical Western Pacific" masterLastPageNumber="372" masterPageNumber="358" pageNumber="360" updateTime="1698294167650" updateUser="ExternalLinkService">
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<mods:title id="4F6D4BAFF709F281EA33BC20A428116F">Paraphelliactis tangi n. sp. and Phelliactis yapensis n. sp., two new deep-sea species of Hormathiidae (Cnidaria: Anthozoa: Actiniaria) from a seamount in the tropical Western Pacific</mods:title>
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<mods:namePart id="2B6C3AC37876D31F89A41B16BF9B08E4">Li, Yang</mods:namePart>
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<mods:namePart id="691BA215164960B9D340C06445A99BD8">Xu, Kuidong</mods:namePart>
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<treatment id="EB7A87EBA76F4C7132BFF99B43686CE8" ID-DOI="http://doi.org/10.5281/zenodo.5658344" ID-GBIF-Taxon="119409188" ID-Zenodo-Dep="5658344" LSID="urn:lsid:plazi:treatment:EB7A87EBA76F4C7132BFF99B43686CE8" httpUri="http://treatment.plazi.org/id/EB7A87EBA76F4C7132BFF99B43686CE8" lastPageId="7" lastPageNumber="365" pageId="2" pageNumber="360">
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<heading id="38248191A76F4C7432BFF99B40C06D97" bold="true" box="[151,476,220,247]" fontSize="11" level="1" pageId="2" pageNumber="360" reason="1">
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<taxonomicName id="A4D34D7EA76F4C7432BFF99B408E6D96" ID-CoL="4D7ZF" box="[151,402,220,246]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="2" pageNumber="360" phylum="Cnidaria" rank="species" species="tangi" status="sp. nov.">
<emphasis id="51A7EAEFA76F4C7432BFF99B408E6D96" bold="true" box="[151,402,220,246]" italics="true" pageId="2" pageNumber="360">Paraphelliactis tangi</emphasis>
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<taxonomicNameLabel id="4A945794A76F4C7433B1F99A40C06D97" box="[409,476,221,247]" pageId="2" pageNumber="360" rank="species">n. sp.</taxonomicNameLabel>
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(
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; 
<tableCitation id="2E510346A76F4C74333FF9B840696C78" box="[279,373,255,280]" captionStart="TABLE 1" captionStartId="4.[151,239,832,855]" captionText="TABLE 1. Distribution and size of the cnidae of Paraphelliactis tangi n. sp. Letters after each type of cnida refer to Fig. 4. N is the proportion of examined individuals with a particular type of cnida; n is the number of cnidae measured. *, no acontium was examined in the smaller paratype." pageId="2" pageNumber="360">Tables 1</tableCitation>
, 2)
</paragraph>
</subSubSection>
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<emphasis id="51A7EAEFA76F4C7432BFF800409F6C00" bold="true" box="[151,387,327,352]" pageId="2" pageNumber="360">Material examined.</emphasis>
<typeStatus id="BC68885FA76F4C7433A5F80040E66C00" box="[397,506,327,352]" pageId="2" pageNumber="360" type="holotype">Holotype</typeStatus>
: 
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<collectingCountry id="1BC4766DA76F4C74302FF80043076C00" box="[519,539,327,352]" name="Italy" pageId="2" pageNumber="360">Y</collectingCountry>
30046
</accessionNumber>
, attached on stones, collected on 
<date id="176D103DA76F4C7431C5F80045D46C00" box="[1005,1224,327,352]" pageId="2" pageNumber="360" value="2014-12-16">16 December 2014</date>
from FX-Dive 17 (
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, 
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), 
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, foraminiferal ooze bottom. 
<typeStatus id="BC68885FA76F4C743154F82A42F16CE5" box="[892,1005,365,389]" pageId="2" pageNumber="360" type="paratype">Paratypes</typeStatus>
: 
<accessionNumber id="7C80AB1EA76F4C7431D7F82B45406CE5" box="[1023,1116,364,389]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/Y30051" pageId="2" pageNumber="360" type="EnaNcbi">
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30051
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, two specimens, embraced sponge spicules, collected on 
<date id="176D103DA76F4C743027F8C843E56CC8" box="[527,761,399,424]" pageId="2" pageNumber="360" value="2014-12-16">16 December 2014</date>
from FX-Dive 17 (
<geoCoordinate id="06E7503AA76F4C7431D5F8C845976CC8" box="[1021,1163,399,424]" direction="east" orientation="longitude" pageId="2" pageNumber="360" precision="9" value="137.83017">137°49.81′E</geoCoordinate>
, 
<geoCoordinate id="06E7503AA76F4C74368AF8C844046CC8" box="[1186,1304,399,424]" direction="north" orientation="latitude" pageId="2" pageNumber="360" precision="9" value="8.996667">8°59.80′N</geoCoordinate>
), 
<quantity id="A42B9B18A76F4C74371EF8D7448B6CC7" box="[1334,1431,400,424]" metricMagnitude="3" metricUnit="m" metricValue="1.928" pageId="2" pageNumber="360" unit="m" value="1928.0">1928 m</quantity>
, foraminiferal ooze bottom.
</paragraph>
<paragraph id="636C36FDA76F4C7432EFF89041EF6E70" blockId="2.[151,1437,327,1037]" pageId="2" pageNumber="360">
<emphasis id="51A7EAEFA76F4C7432EFF89040646C90" bold="true" box="[199,376,471,496]" pageId="2" pageNumber="360">Body and Size.</emphasis>
Pedal disc attached to stone or grasping sponge spicules. Column covered with brown cuticle, darker proximally than distally (
<figureCitation id="FBE82A78A76F4C743038F8BB43446F74" box="[528,600,508,533]" captionStart="FIGURE 2" captionStartId="2.[151,250,1928,1950]" captionTargetBox="[151,1436,1089,1906]" captionTargetId="figure@2.[151,1436,1089,1907]" captionTargetPageId="2" captionText="FIGURE 2. External morphology of Paraphelliactis tangi n. sp. A, B. The holotype in situ and in preservation. C. The larger paratype in preservation. Scale bars = 40 mm." httpUri="https://zenodo.org/record/255246/files/figure.png" pageId="2" pageNumber="360">Fig. 2</figureCitation>
). In preservation, column sub-cylindrical, height 
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(
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in 
<typeStatus id="BC68885FA76F4C7432BFFB5841E16F58" box="[151,253,543,568]" pageId="2" pageNumber="360" type="holotype">holotype</typeStatus>
), diameter of pedal disc 
<quantity id="A42B9B18A76F4C743025FB67439A6F57" box="[525,646,544,568]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" metricValueMax="8.9" metricValueMin="2.7" pageId="2" pageNumber="360" unit="mm" value="58.0" valueMax="89.0" valueMin="27.0">27–89 mm</quantity>
(
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in 
<typeStatus id="BC68885FA76F4C74312FFB5842726F58" box="[775,878,543,568]" pageId="2" pageNumber="360" type="holotype">holotype</typeStatus>
), larger than that of proximal column (
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; 
<quantity id="A42B9B18A76F4C7432BFFB0341F56F3C" box="[151,233,580,605]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="360" unit="mm" value="45.0">45 mm</quantity>
in 
<typeStatus id="BC68885FA76F4C74333BFB0340656F3D" box="[275,377,580,605]" pageId="2" pageNumber="360" type="holotype">holotype</typeStatus>
), greatest diameter of distal column 
<quantity id="A42B9B18A76F4C74310BFB0342836F3C" box="[803,927,580,604]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" metricValueMax="7.4" metricValueMin="1.7" pageId="2" pageNumber="360" unit="mm" value="45.5" valueMax="74.0" valueMin="17.0">17–74 mm</quantity>
(
<quantity id="A42B9B18A76F4C743198FB0345186F3C" box="[944,1028,580,604]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="2" pageNumber="360" unit="mm" value="74.0">74 mm</quantity>
in 
<typeStatus id="BC68885FA76F4C743606FB03458E6F3D" box="[1070,1170,580,605]" pageId="2" pageNumber="360" type="holotype">holotype</typeStatus>
) (
<figureCitation id="FBE82A78A76F4C743684FB0345EE6F3C" box="[1196,1266,580,605]" captionStart="FIGURE 2" captionStartId="2.[151,250,1928,1950]" captionTargetBox="[151,1436,1089,1906]" captionTargetId="figure@2.[151,1436,1089,1907]" captionTargetPageId="2" captionText="FIGURE 2. External morphology of Paraphelliactis tangi n. sp. A, B. The holotype in situ and in preservation. C. The larger paratype in preservation. Scale bars = 40 mm." httpUri="https://zenodo.org/record/255246/files/figure.png" pageId="2" pageNumber="360">Fig. 2</figureCitation>
B, C). Column divisible into scapus and scapulus: former provided with layer of cuticle and larger tubercles; latter short, without cuticle, longitudinally furrowed, with smaller tubercles (
<figureCitation id="FBE82A78A76F4C743107FBCB42666FC4" box="[815,890,652,677]" captionStart="FIGURE 2" captionStartId="2.[151,250,1928,1950]" captionTargetBox="[151,1436,1089,1906]" captionTargetId="figure@2.[151,1436,1089,1907]" captionTargetPageId="2" captionText="FIGURE 2. External morphology of Paraphelliactis tangi n. sp. A, B. The holotype in situ and in preservation. C. The larger paratype in preservation. Scale bars = 40 mm." httpUri="https://zenodo.org/record/255246/files/figure.png" pageId="2" pageNumber="360">Fig. 2</figureCitation>
). Tubercles mainly in distal column, typically conical in 
<typeStatus id="BC68885FA76F4C743323FBE840716FA8" box="[267,365,687,712]" pageId="2" pageNumber="360" type="holotype">holotype</typeStatus>
and rounded in smaller specimens, of different sizes, diameters of base to 
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and height to 
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, irregularly arranged. Mesogloea of column thick, to 
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between tubercles and about same thickness along column.
</paragraph>
<paragraph id="636C36FDA76F4C7432EFFA5B4217696D" blockId="2.[151,1437,327,1037]" pageId="2" pageNumber="360">
<emphasis id="51A7EAEFA76F4C7432EFFA5B40E76E55" bold="true" box="[199,507,796,821]" pageId="2" pageNumber="360">Oral Disc and Tentacles.</emphasis>
Oral disc pink, elliptical, long axis diameter 
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(
<quantity id="A42B9B18A76F4C74368DFA5B45E76E54" box="[1189,1275,796,821]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="2" pageNumber="360" unit="mm" value="54.0">54 mm</quantity>
in 
<typeStatus id="BC68885FA76F4C743702FA5B448D6E55" box="[1322,1425,796,821]" pageId="2" pageNumber="360" type="holotype">holotype</typeStatus>
). Actinopharynx well developed, length to 
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, occupying about 1/3 of column length in 
<typeStatus id="BC68885FA76F4C7436E8FA78443A6E38" box="[1216,1318,831,856]" pageId="2" pageNumber="360" type="holotype">holotype</typeStatus>
. Tentacles marginal, retractile, smooth, tapered, and without mesogloeal thickenings on aboral side (
<figureCitation id="FBE82A78A76F4C7436A7FA2345C36E1C" box="[1167,1247,868,893]" captionStart="FIGURE 2" captionStartId="2.[151,250,1928,1950]" captionTargetBox="[151,1436,1089,1906]" captionTargetId="figure@2.[151,1436,1089,1907]" captionTargetPageId="2" captionText="FIGURE 2. External morphology of Paraphelliactis tangi n. sp. A, B. The holotype in situ and in preservation. C. The larger paratype in preservation. Scale bars = 40 mm." httpUri="https://zenodo.org/record/255246/files/figure.png" pageId="2" pageNumber="360">Figs. 2</figureCitation>
A). Length to 
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in life, to 
<quantity id="A42B9B18A76F4C743315FACF40916EFF" box="[317,397,904,928]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="360" unit="mm" value="10.0">10 mm</quantity>
long and 
<quantity id="A42B9B18A76F4C743029FACF435F6EFF" box="[513,579,904,928]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="360" unit="mm" value="3.0">3 mm</quantity>
wide in preservation. Tentacles hexamerously arranged in six cycles, inner ones wider and longer than outer ones, 
<quantity id="A42B9B18A76F4C743073FAEB43B86EA5" box="[603,676,940,965]" metricMagnitude="0" metricUnit="m" metricValue="4.5974" pageId="2" pageNumber="360" unit="in" value="181.0">181 in</quantity>
<typeStatus id="BC68885FA76F4C743085FAEB42136EA5" box="[685,783,940,965]" pageId="2" pageNumber="360" type="holotype">holotype</typeStatus>
(12 cut off with margin for histological sections), 
<quantity id="A42B9B18A76F4C74377BFAEB44806EA5" box="[1363,1436,940,965]" metricMagnitude="0" metricUnit="m" metricValue="4.8514" pageId="2" pageNumber="360" unit="in" value="191.0">191 in</quantity>
larger 
<typeStatus id="BC68885FA76F4C7432C8FA97405C6E88" box="[224,320,976,1000]" pageId="2" pageNumber="360" type="paratype">paratype</typeStatus>
(15 cut off with margin for histological sections), and 
<quantity id="A42B9B18A76F4C74319CFA9742F26E88" box="[948,1006,975,1000]" metricMagnitude="0" metricUnit="m" metricValue="2.3368" pageId="2" pageNumber="360" unit="in" value="92.0">92 in</quantity>
smaller 
<typeStatus id="BC68885FA76F4C74367AFA9745AA6E88" box="[1106,1206,976,1000]" pageId="2" pageNumber="360" type="paratype">paratype</typeStatus>
; full complement of large individuals likely 192 (6 + 6 + 12 + 24 + 48 + 96).
</paragraph>
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<paragraph id="636C36FDA76F4C7432BFFECF43746ADD" blockId="2.[151,1436,1928,1981]" pageId="2" pageNumber="360">
<emphasis id="51A7EAEFA76F4C7432BFFECF400F6AFD" bold="true" box="[151,275,1928,1950]" pageId="2" pageNumber="360">FIGURE 2.</emphasis>
External morphology of 
<taxonomicName id="A4D34D7EA76F4C74303DFECF43F06AFE" box="[533,748,1928,1950]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="2" pageNumber="360" phylum="Cnidaria" rank="species" species="tangi" status="sp. nov.">
<emphasis id="51A7EAEFA76F4C74303DFECF43F06AFE" box="[533,748,1928,1950]" italics="true" pageId="2" pageNumber="360">Paraphelliactis tangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4A945794A76F4C7430DBFECE42316AFE" box="[755,813,1929,1950]" pageId="2" pageNumber="360" rank="species">
<emphasis id="51A7EAEFA76F4C7430DBFECE42316AFE" bold="true" box="[755,813,1929,1950]" pageId="2" pageNumber="360">n. sp.</emphasis>
</taxonomicNameLabel>
A, B. The holotype 
<emphasis id="51A7EAEFA76F4C7431D7FECE45226AFE" box="[1023,1086,1929,1950]" italics="true" pageId="2" pageNumber="360">in situ</emphasis>
and in preservation. C. The larger paratype in preservation. Scale bars = 40 mm.
</paragraph>
</caption>
<caption id="37AC6675A76E4C7532BFFE7443546AA4" httpUri="https://zenodo.org/record/255247/files/figure.png" pageId="3" pageNumber="361" targetBox="[151,1436,193,1820]" targetPageId="3">
<paragraph id="636C36FDA76E4C7532BFFE7443546AA4" blockId="3.[151,1437,1843,1988]" pageId="3" pageNumber="361">
<emphasis id="51A7EAEFA76E4C7532BFFE74400A6A28" bold="true" box="[151,278,1843,1865]" pageId="3" pageNumber="361">FIGURE 3.</emphasis>
Internal anatomy of 
<taxonomicName id="A4D34D7EA76E4C7533D0FE7443CE6A29" box="[504,722,1843,1865]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="3" pageNumber="361" phylum="Cnidaria" rank="species" species="tangi" status="sp. nov.">
<emphasis id="51A7EAEFA76E4C7533D0FE7443CE6A29" box="[504,722,1843,1865]" italics="true" pageId="3" pageNumber="361">Paraphelliactis tangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4A945794A76E4C7530F4FE7342056A29" box="[732,793,1844,1865]" pageId="3" pageNumber="361" rank="species">
<emphasis id="51A7EAEFA76E4C7530F4FE7342056A29" bold="true" box="[732,793,1844,1865]" pageId="3" pageNumber="361">n. sp.</emphasis>
</taxonomicNameLabel>
A. Longitudinal section through the margin of the holotype, showing marginal sphincter muscle (S). B. Transverse section through the oral disc of the holotype, note much thicker mesogloea over an endocoel (arrow). C. Cross sections of mesenteries through the actinopharynx of the larger paratype, showing weak parietobasilar muscle (P), diffuse retractors (R) and oocytes (O). Numbers indicate the cycles of mesenteries. Scale bars = 0.5 mm (A, B) and 1 mm (C).
</paragraph>
</caption>
<paragraph id="636C36FDA7694C7232EFF9D0441B6CB0" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="362">
<emphasis id="51A7EAEFA7694C7232EFF9D040B76DD0" bold="true" box="[199,427,151,176]" pageId="4" pageNumber="362">Internal Anatomy.</emphasis>
Two symmetrical siphonoglyphs well developed, each attached to pair of directive mesenteries. Mesenteries not divisible into macrocnemes and microcnemes. Equal number of mesenteries at margin and limbus. In large specimens (
<typeStatus id="BC68885FA7694C723044F99843CC6D98" box="[620,720,223,248]" pageId="4" pageNumber="362" type="holotype">holotype</typeStatus>
and larger 
<typeStatus id="BC68885FA7694C723171F9A742A26D98" box="[857,958,224,248]" pageId="4" pageNumber="362" type="paratype">paratype</typeStatus>
), 96 pairs hexamerously arranged in five cycles. In 
<typeStatus id="BC68885FA7694C723326F843406F6C7D" box="[270,371,260,285]" pageId="4" pageNumber="362" type="holotype">holotype</typeStatus>
, six pairs of mesenteries of first cycle and one mesentery of second cycle perfect and sterile (mesentery pairing with perfect one of second cycle fertile). In larger 
<typeStatus id="BC68885FA7694C7231B7F86F451E6C20" box="[927,1026,296,320]" pageId="4" pageNumber="362" type="paratype">paratype</typeStatus>
, mesenteries of first cycle except for one perfect and sterile (imperfect mesentery also sterile). Mesenteries of second-fourth cycle fertile (except for perfect mesentery in 
<typeStatus id="BC68885FA7694C7233AEF82840F06CE8" box="[390,492,367,392]" pageId="4" pageNumber="362" type="holotype">holotype</typeStatus>
), with oocytes larger than 250 Μm in diameter. Mesenteries of fifth cycle sterile. In smaller 
<typeStatus id="BC68885FA7694C7232DAF8D240496CCD" box="[242,341,405,429]" pageId="4" pageNumber="362" type="paratype">paratype</typeStatus>
, 48 pairs of mesenteries regularly arranged in four cycles: first cycle perfect and sterile, second and third cycles fertile, fourth cycle sterile. No cinclides. Larger mesenteries bear acontia proximally.
</paragraph>
<paragraph id="636C36FDA7694C7232EFF89B40BE6F00" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="362">
Sphincter mesogloeal, alveolar, not very strong (
<figureCitation id="FBE82A78A7694C723121F89B424D6C94" box="[777,849,476,501]" captionStart="FIGURE 3" captionStartId="3.[151,250,1843,1865]" captionTargetBox="[151,1436,193,1820]" captionTargetId="figure@3.[151,1436,193,1822]" captionTargetPageId="3" captionText="FIGURE 3. Internal anatomy of Paraphelliactis tangi n. sp. A. Longitudinal section through the margin of the holotype, showing marginal sphincter muscle (S). B. Transverse section through the oral disc of the holotype, note much thicker mesogloea over an endocoel (arrow). C. Cross sections of mesenteries through the actinopharynx of the larger paratype, showing weak parietobasilar muscle (P), diffuse retractors (R) and oocytes (O). Numbers indicate the cycles of mesenteries. Scale bars = 0.5 mm (A, B) and 1 mm (C)." httpUri="https://zenodo.org/record/255247/files/figure.png" pageId="4" pageNumber="362">Fig. 3</figureCitation>
A). Longitudinal muscles of tentacles ectodermal. Radial muscles of oral disc meso-ectodermal, and mesogloea much thicker in parts corresponding to stronger endocoels than in exocoels (
<figureCitation id="FBE82A78A7694C7233C2FB63432F6F5C" box="[490,563,548,573]" captionStart="FIGURE 3" captionStartId="3.[151,250,1843,1865]" captionTargetBox="[151,1436,193,1820]" captionTargetId="figure@3.[151,1436,193,1822]" captionTargetPageId="3" captionText="FIGURE 3. Internal anatomy of Paraphelliactis tangi n. sp. A. Longitudinal section through the margin of the holotype, showing marginal sphincter muscle (S). B. Transverse section through the oral disc of the holotype, note much thicker mesogloea over an endocoel (arrow). C. Cross sections of mesenteries through the actinopharynx of the larger paratype, showing weak parietobasilar muscle (P), diffuse retractors (R) and oocytes (O). Numbers indicate the cycles of mesenteries. Scale bars = 0.5 mm (A, B) and 1 mm (C)." httpUri="https://zenodo.org/record/255247/files/figure.png" pageId="4" pageNumber="362">Fig. 3</figureCitation>
B). Parietobasilar muscles weak (
<figureCitation id="FBE82A78A7694C7231E9FB6345156F5C" box="[961,1033,548,573]" captionStart="FIGURE 3" captionStartId="3.[151,250,1843,1865]" captionTargetBox="[151,1436,193,1820]" captionTargetId="figure@3.[151,1436,193,1822]" captionTargetPageId="3" captionText="FIGURE 3. Internal anatomy of Paraphelliactis tangi n. sp. A. Longitudinal section through the margin of the holotype, showing marginal sphincter muscle (S). B. Transverse section through the oral disc of the holotype, note much thicker mesogloea over an endocoel (arrow). C. Cross sections of mesenteries through the actinopharynx of the larger paratype, showing weak parietobasilar muscle (P), diffuse retractors (R) and oocytes (O). Numbers indicate the cycles of mesenteries. Scale bars = 0.5 mm (A, B) and 1 mm (C)." httpUri="https://zenodo.org/record/255247/files/figure.png" pageId="4" pageNumber="362">Fig. 3</figureCitation>
C). Longitudinal retractor muscles diffuse, weak (
<figureCitation id="FBE82A78A7694C723315FB0040636F00" box="[317,383,583,608]" captionStart="FIGURE 3" captionStartId="3.[151,250,1843,1865]" captionTargetBox="[151,1436,193,1820]" captionTargetId="figure@3.[151,1436,193,1822]" captionTargetPageId="3" captionText="FIGURE 3. Internal anatomy of Paraphelliactis tangi n. sp. A. Longitudinal section through the margin of the holotype, showing marginal sphincter muscle (S). B. Transverse section through the oral disc of the holotype, note much thicker mesogloea over an endocoel (arrow). C. Cross sections of mesenteries through the actinopharynx of the larger paratype, showing weak parietobasilar muscle (P), diffuse retractors (R) and oocytes (O). Numbers indicate the cycles of mesenteries. Scale bars = 0.5 mm (A, B) and 1 mm (C)." httpUri="https://zenodo.org/record/255247/files/figure.png" pageId="4" pageNumber="362">Fig. 3</figureCitation>
C).
</paragraph>
<paragraph id="636C36FDA7694C7232EFFB2B446E6FE5" blockId="4.[151,1437,151,789]" box="[199,1394,620,645]" pageId="4" pageNumber="362">
<emphasis id="51A7EAEFA7694C7232EFFB2B40326FE5" bold="true" box="[199,302,620,645]" pageId="4" pageNumber="362">Cnidom.</emphasis>
Spirocysts, basitrichs, microbasic 
<emphasis id="51A7EAEFA7694C72309BFB2A43DD6FE4" box="[691,705,621,644]" italics="true" pageId="4" pageNumber="362">p</emphasis>
-mastigophores (
<figureCitation id="FBE82A78A7694C723155FB2B42DE6FE4" box="[893,962,620,645]" captionStart="FIGURE 4" captionStartId="5.[151,250,1023,1045]" captionTargetBox="[167,1421,339,1007]" captionTargetId="figure@5.[160,1426,331,1012]" captionTargetPageId="5" captionText="FIGURE 4. Cnidae from Paraphelliactis tangi n. sp. Tentacles: A. Robust spirocyst. B. Gracile spirocyst. C. Basitrich. D. Basitrich. Column: E. Basitrich. F. Basitrich. G. Microbasic p - mastigophore. Actinopharynx: H. Basitrich. I. Basitrich. J. Microbasic p - mastigophore. Filament: K. Basitrich. L. Basitrich. M. Microbasic p - mastigophore. Acontia: N. Basitrich. O. Basitrich. Scale bar = 20 µm." httpUri="https://zenodo.org/record/255248/files/figure.png" pageId="4" pageNumber="362">Fig. 4</figureCitation>
). See 
<tableCitation id="2E510346A7694C72362EFB2B45456FE4" box="[1030,1113,620,645]" captionStart="TABLE 1" captionStartId="4.[151,239,832,855]" captionText="TABLE 1. Distribution and size of the cnidae of Paraphelliactis tangi n. sp. Letters after each type of cnida refer to Fig. 4. N is the proportion of examined individuals with a particular type of cnida; n is the number of cnidae measured. *, no acontium was examined in the smaller paratype." pageId="4" pageNumber="362">Table 1</tableCitation>
for size and distribution.
</paragraph>
</subSubSection>
<subSubSection id="2BC96576A7694C7232EFFBC843CD6BD2" pageId="4" pageNumber="362" type="distribution">
<paragraph id="636C36FDA7694C7232EFFBC8429C6E75" blockId="4.[151,1437,151,789]" pageId="4" pageNumber="362">
<emphasis id="51A7EAEFA7694C7232EFFBC840E06FC8" bold="true" box="[199,508,655,680]" pageId="4" pageNumber="362">Distribution and Habitat.</emphasis>
<taxonomicName id="A4D34D7EA7694C72302EFBD743EA6FC7" box="[518,758,656,680]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="species" species="tangi" status="sp. nov.">
<emphasis id="51A7EAEFA7694C72302EFBD743EA6FC7" box="[518,758,656,680]" italics="true" pageId="4" pageNumber="362">Paraphelliactis tangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4A945794A7694C723128FBC8425D6FC8" box="[768,833,655,680]" pageId="4" pageNumber="362" rank="species">
<emphasis id="51A7EAEFA7694C723128FBC8425D6FC8" bold="true" box="[768,833,655,680]" pageId="4" pageNumber="362">n. sp.</emphasis>
</taxonomicNameLabel>
has been found only from a seamount near the Yap Trench in the tropical Western Pacific, where the water depth ranged from 
<quantity id="A42B9B18A7694C7231D8FBF345526FAC" box="[1008,1102,692,716]" metricMagnitude="3" metricUnit="m" metricValue="1.928" pageId="4" pageNumber="362" unit="m" value="1928.0">1,928 m</quantity>
to 
<quantity id="A42B9B18A7694C72365DFBF345CF6FAC" box="[1141,1235,692,716]" metricMagnitude="3" metricUnit="m" metricValue="1.98" pageId="4" pageNumber="362" unit="m" value="1980.0">1,980 m</quantity>
and the sediment was foraminiferal ooze. Some other individuals were also observed by ROV nearby, and the 
<typeStatus id="BC68885FA7694C723689FB90441F6F90" box="[1185,1283,727,752]" pageId="4" pageNumber="362" type="holotype">holotype</typeStatus>
attached on a stone and the 
<typeStatus id="BC68885FA7694C72331AFBBA40816E75" box="[306,413,765,789]" pageId="4" pageNumber="362" type="paratype">paratypes</typeStatus>
embraced sponge spicules with pedal disc.
</paragraph>
<caption id="37AC6675A7694C7232BFFA0743866EF7" pageId="4" pageNumber="362">
<paragraph id="636C36FDA7694C7232BFFA0743866EF7" blockId="4.[151,1436,831,919]" pageId="4" pageNumber="362">
<emphasis id="51A7EAEFA7694C7232BFFA0740156E37" bold="true" box="[151,265,832,855]" pageId="4" pageNumber="362">TABLE 1.</emphasis>
Distribution and size of the cnidae of 
<taxonomicName id="A4D34D7EA7694C72308AFA78429F6E37" box="[674,899,831,855]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="species" species="tangi" status="sp. nov.">
<emphasis id="51A7EAEFA7694C72308AFA78429F6E37" box="[674,899,831,855]" italics="true" pageId="4" pageNumber="362">Paraphelliactis tangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4A945794A7694C7231A2FA0742DA6E37" box="[906,966,832,855]" pageId="4" pageNumber="362" rank="species">
<emphasis id="51A7EAEFA7694C7231A2FA0742DA6E37" bold="true" box="[906,966,832,855]" pageId="4" pageNumber="362">n. sp.</emphasis>
</taxonomicNameLabel>
Letters after each type of cnida refer to Fig. 4. N is the proportion of examined individuals with a particular type of cnida; n is the number of cnidae measured. *, no acontium was examined in the smaller paratype.
</paragraph>
</caption>
<paragraph id="636C36FDA7694C7232B3FAF5458A6EA8" blockId="4.[155,1303,946,1588]" box="[155,1174,946,968]" pageId="4" pageNumber="362">
Tissue Cnida 
<typeStatus id="BC68885FA7694C723026FAF443266EA8" box="[526,570,947,968]" pageId="4" pageNumber="362">type</typeStatus>
N n Range, in µm
</paragraph>
<paragraph id="636C36FDA7694C7232B3FA9B440B6E92" blockId="4.[155,1303,946,1588]" box="[155,1303,988,1010]" pageId="4" pageNumber="362">
Tentacle Robust spirocysts (A) 
<date id="176D103DA7694C723173FA9B42CD6E92" box="[859,977,988,1010]" pageId="4" pageNumber="362" value="1965-03-03">3/3 65</date>
(21.5) 25.5–75.0 × 3.2–9.0
</paragraph>
<paragraph id="636C36FDA7694C7233E3FD4245C8690E" blockId="4.[155,1303,946,1588]" pageId="4" pageNumber="362">
Gracile spirocysts (B) 
<date id="176D103DA7694C723173FD4242CD697B" box="[859,977,1029,1051]" pageId="4" pageNumber="362" value="1945-03-03">3/3 45</date>
25.0–49.0 × 3.0–4.0 Basitrichs (C) 
<date id="176D103DA7694C723173FD6942CD6924" box="[859,977,1070,1092]" pageId="4" pageNumber="362" value="1952-03-03">3/3 52</date>
26.0–52.0 (56.0) × 2.5–4.0 Basitrichs (D) 2/3 7 16.0–21.0 × 1.5–3.0
</paragraph>
<paragraph id="636C36FDA7694C7232B3FDC645DB69A0" blockId="4.[155,1303,946,1588]" pageId="4" pageNumber="362">
Middle column Basitrichs (E) 
<date id="176D103DA7694C723173FDC642CD69F7" box="[859,977,1153,1175]" pageId="4" pageNumber="362" value="1940-03-03">3/3 40</date>
14.0–24.0 × 2.0–4.0 Basitrichs (F) 
<date id="176D103DA7694C723173FDED42CD69A0" box="[859,977,1194,1216]" pageId="4" pageNumber="362" value="1933-03-03">3/3 33</date>
9.5–14.0 × 1.5–2.0
</paragraph>
<paragraph id="636C36FDA7694C7233E3FD9345D1698A" blockId="4.[155,1303,946,1588]" box="[459,1229,1236,1258]" pageId="4" pageNumber="362">
Microbasic 
<emphasis id="51A7EAEFA7694C72306CFD92434D698A" box="[580,593,1237,1258]" italics="true" pageId="4" pageNumber="362">p</emphasis>
-mastigophores (G) 
<date id="176D103DA7694C723173FD9342CD698A" box="[859,977,1236,1258]" pageId="4" pageNumber="362" value="1918-03-01">1/3 18</date>
18.2–21.0 ×3.0–4.0
</paragraph>
<paragraph id="636C36FDA7694C7232B3FDBA45B4685C" blockId="4.[155,1303,946,1588]" pageId="4" pageNumber="362">
Actinopharynx Basitrichs (H) 
<date id="176D103DA7694C723173FDBA42CD6873" box="[859,977,1277,1299]" pageId="4" pageNumber="362" value="1961-03-03">3/3 61</date>
25.5–55.0 × 2.9–4.0 Basitrichs (I) 
<date id="176D103DA7694C723173FC6142CD685C" box="[859,977,1318,1340]" pageId="4" pageNumber="362" value="1925-03-03">3/3 25</date>
17.5–24.5 × 2.0
</paragraph>
<paragraph id="636C36FDA7694C7233E3FC17440B6806" blockId="4.[155,1303,946,1588]" box="[459,1303,1360,1382]" pageId="4" pageNumber="362">
Microbasic 
<emphasis id="51A7EAEFA7694C72306CFC16434D6806" box="[580,593,1361,1382]" italics="true" pageId="4" pageNumber="362">p</emphasis>
-mastigophores (J) 
<date id="176D103DA7694C723173FC1742CD6806" box="[859,977,1360,1382]" pageId="4" pageNumber="362" value="1946-03-03">3/3 46</date>
25.0–41.0 (50.0) × 4.0–4.9
</paragraph>
<paragraph id="636C36FDA7694C7232B3FC3E441668EF" blockId="4.[155,1303,946,1588]" box="[155,1290,1401,1423]" pageId="4" pageNumber="362">
Mesenterial filament Basitrichs (K) 
<date id="176D103DA7694C723173FC3E42CD68EF" box="[859,977,1401,1423]" pageId="4" pageNumber="362" value="1935-03-03">3/3 35</date>
35.0–60.0 × 3.0–4.0 (5.0)
</paragraph>
<paragraph id="636C36FDA7694C7233E3FCE545C86882" blockId="4.[155,1303,946,1588]" pageId="4" pageNumber="362">
Basitrichs (L) 
<date id="176D103DA7694C723173FCE542CD68D8" box="[859,977,1442,1464]" pageId="4" pageNumber="362" value="1943-03-03">3/3 43</date>
16.0–26.0 × 1.5–3.0 Microbasic 
<emphasis id="51A7EAEFA7694C72306CFC8A434D6882" box="[580,593,1485,1506]" italics="true" pageId="4" pageNumber="362">p</emphasis>
-mastigophores (M) 
<date id="176D103DA7694C723173FC8B42CD6882" box="[859,977,1484,1506]" pageId="4" pageNumber="362" value="1924-03-03">3/3 24</date>
24.0–37.0 × 3.0–5.5
</paragraph>
<paragraph id="636C36FDA7694C7232B3FCB243CD6BD2" blockId="4.[155,1303,946,1588]" lastBlockId="4.[151,1436,1653,2038]" pageId="4" pageNumber="362">
Acontia Basitrichs (N) 2/2* 65 49.0–62.0 × 3.0–4.1 Basitrichs (
<collectingCountry id="1BC4766DA7694C723016FF59434E6B54" box="[574,594,1566,1588]" name="Iceland" pageId="4" pageNumber="362">O</collectingCountry>
) 
<date id="176D103DA7694C723173FF5942CD6B54" box="[859,977,1566,1588]" pageId="4" pageNumber="362" value="1916-02-02">2/2 16</date>
17.0–27.0 × 2.0 
<emphasis id="51A7EAEFA7694C7232EFFF3240526BEE" bold="true" box="[199,334,1653,1678]" pageId="4" pageNumber="362">Etymology.</emphasis>
Named in honor of the late Chinese marine biologist, Prof. Zhican TANG, for his great contribution to the taxonomy of Cnidaria in 
<collectingCountry id="1BC4766DA7694C7230AEFFDE43D16BD2" box="[646,717,1689,1714]" name="China" pageId="4" pageNumber="362">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="2BC96576A7694C7332EFFFFA40C76AE7" lastPageId="5" lastPageNumber="363" pageId="4" pageNumber="362" type="discussion">
<paragraph id="636C36FDA7694C7332EFFFFA43726C7C" blockId="4.[151,1436,1653,2038]" lastBlockId="5.[151,1436,151,285]" lastPageId="5" lastPageNumber="363" pageId="4" pageNumber="362">
<emphasis id="51A7EAEFA7694C7232EFFFFA40276BB6" bold="true" box="[199,315,1725,1750]" pageId="4" pageNumber="362">Remarks.</emphasis>
The genus 
<taxonomicName id="A4D34D7EA7694C7233E7FFFA43606BB5" box="[463,636,1725,1749]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7694C7233E7FFFA43606BB5" box="[463,636,1725,1749]" italics="true" pageId="4" pageNumber="362">Paraphelliactis</emphasis>
</taxonomicName>
was established by 
<bibRefCitation id="07424B0CA7694C72315CFFFA45266BB6" author="Carlgren" box="[884,1082,1725,1750]" pageId="4" pageNumber="362" refString="Carlgren, O. (1928 a) Ceriantharier, Zoantharier och Actiniarier. Meddelelser om Gronland, Supplement 23, 253 - 308." type="book chapter" year="1928" yearSuffix="a">Carlgren (1928a)</bibRefCitation>
for 
<taxonomicName id="A4D34D7EA7694C72365DFFF945E26BB5" box="[1141,1278,1726,1749]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="species" species="spinosa">
<emphasis id="51A7EAEFA7694C72365DFFF945E26BB5" box="[1141,1278,1726,1749]" italics="true" pageId="4" pageNumber="362">Pa. spinosa</emphasis>
</taxonomicName>
based on its resemblance of appearance and structure with 
<taxonomicName id="A4D34D7EA7694C7230EEFFA642F86B9A" authority="Simon, 1892" authorityName="Simon" authorityYear="1892" box="[710,996,1761,1786]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7694C7230EEFFA642226B99" box="[710,830,1761,1785]" italics="true" pageId="4" pageNumber="362">Phelliactis</emphasis>
<bibRefCitation id="07424B0CA7694C723163FFA642F86B9A" author="Simon" box="[843,996,1761,1786]" pageId="4" pageNumber="362" refString="Simon, J. A. (1892) Ein Beitrag zur Anatomie und Systematik der Hexactinien. Druck von Val. Hofling, Kapellenstrasse 3, Munchen, 102 pp. [pp. 5 - 106]" type="book chapter" year="1892">Simon, 1892</bibRefCitation>
</taxonomicName>
. Subsequently a second species, 
<taxonomicName id="A4D34D7EA7694C72375EFFA540C56A7E" authority="Carlgren, 1934" authorityName="Carlgren" authorityYear="1934" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="species" species="michaelsarsi">
<emphasis id="51A7EAEFA7694C72375EFFA5403A6A7D" italics="true" pageId="4" pageNumber="362">Pa. michaelsarsi</emphasis>
<bibRefCitation id="07424B0CA7694C723305FE4240C56A7E" author="Carlgren" box="[301,473,1797,1822]" pageId="4" pageNumber="362" refString="Carlgren, O. (1934) Ceriantharia, Zoantharia and Actiniaria from the &quot; Michael Sars &quot; North Atlantic Deep-sea Expedition 1910. Report on the Scientific Results of the &quot; Michael Sars &quot; North Atlantic Deep-Sea Expedition 1910, 5 (6), 1 - 27." type="journal article" year="1934">Carlgren, 1934</bibRefCitation>
</taxonomicName>
, was described. 
<bibRefCitation id="07424B0CA7694C7230BAFE42425C6A7E" author="Carlgren" box="[658,832,1797,1822]" pageId="4" pageNumber="362" refString="Carlgren, O. (1942) Actiniaria II. Danish Ingolf-Expedition, 5 (12), 1 - 92." type="journal article" year="1942">Carlgren (1942</bibRefCitation>
, 
<bibRefCitation id="07424B0CA7694C723164FE41429A6A7E" author="Carlgren" box="[844,902,1798,1822]" pageId="4" pageNumber="362" refString="Carlgren, O. (1949) A survey of the Ptychodactaria, Corallimorpharia, and Actiniaria. Kungliga Svenska Vetenskapsakadamiens Handlingar, 1 (1), 1 - 121." type="journal article" year="1949">1949</bibRefCitation>
) distinguished 
<taxonomicName id="A4D34D7EA7694C72361AFE4245C36A7D" box="[1074,1247,1797,1821]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7694C72361AFE4245C36A7D" box="[1074,1247,1797,1821]" italics="true" pageId="4" pageNumber="362">Paraphelliactis</emphasis>
</taxonomicName>
from 
<taxonomicName id="A4D34D7EA7694C72370CFE4244806A7D" box="[1316,1436,1797,1821]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7694C72370CFE4244806A7D" box="[1316,1436,1797,1821]" italics="true" pageId="4" pageNumber="362">Phelliactis</emphasis>
</taxonomicName>
mainly by the different arrangement of the radial muscles of the oral disc and the (probably) greater number of mesenteries at the margin than at the limbus. 
<bibRefCitation id="07424B0CA7694C7230A6FE0A42B76A06" author="Riemann-Zurneck" box="[654,939,1869,1894]" pageId="4" pageNumber="362" refString="Riemann-Zurneck, K. (1973) Actiniaria des Sudwestatlantik I. Hormathiidae. Helgol wiss Meeres, 25, 273 - 325. http: // dx. doi. org / 10.1007 / BF 01611200" type="journal article" year="1973">Riemann-Zürneck (1973)</bibRefCitation>
suggested the radial muscles of the oral disc used to separate 
<taxonomicName id="A4D34D7EA7694C723342FE36430B6AE9" box="[362,535,1905,1929]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7694C723342FE36430B6AE9" box="[362,535,1905,1929]" italics="true" pageId="4" pageNumber="362">Paraphelliactis</emphasis>
</taxonomicName>
and 
<taxonomicName id="A4D34D7EA7694C723075FE3643C96AE9" box="[605,725,1905,1929]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7694C723075FE3643C96AE9" box="[605,725,1905,1929]" italics="true" pageId="4" pageNumber="362">Phelliactis</emphasis>
</taxonomicName>
were variable and thus synonymized 
<taxonomicName id="A4D34D7EA7694C723686FE3644476AE9" box="[1198,1371,1905,1929]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7694C723686FE3644476AE9" box="[1198,1371,1905,1929]" italics="true" pageId="4" pageNumber="362">Paraphelliactis</emphasis>
</taxonomicName>
with 
<taxonomicName id="A4D34D7EA7694C7232BFFED240136ACD" box="[151,271,1941,1965]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7694C7232BFFED240136ACD" box="[151,271,1941,1965]" italics="true" pageId="4" pageNumber="362">Phelliactis</emphasis>
</taxonomicName>
. Later, 
<bibRefCitation id="07424B0CA7694C72334CFED140E86ACE" author="Dunn" box="[356,500,1942,1966]" pageId="4" pageNumber="362" refString="Dunn, D. F. (1982) Paraphelliactis pabista, a new species of hormathiid sea anemone from abyssal northeastern Pacific waters (Coelenterata: Actiniaria). Syesis, 15, 51 - 56." type="journal article" year="1982">Dunn (1982)</bibRefCitation>
described the third species of 
<taxonomicName id="A4D34D7EA7694C72316FFED242E86ACD" box="[839,1012,1941,1965]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7694C72316FFED242E86ACD" box="[839,1012,1941,1965]" italics="true" pageId="4" pageNumber="362">Paraphelliactis</emphasis>
</taxonomicName>
, 
<taxonomicName id="A4D34D7EA7694C72362AFED1459E6ACD" box="[1026,1154,1941,1965]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="species" species="pabista">
<emphasis id="51A7EAEFA7694C72362AFED1459E6ACD" box="[1026,1154,1941,1965]" italics="true" pageId="4" pageNumber="362">Pa. pabista</emphasis>
</taxonomicName>
. 
<bibRefCitation id="07424B0CA7694C7236A7FED141FD6AB2" author="Sanamyan" pageId="4" pageNumber="362" refString="Sanamyan, N. P. &amp; Sanamyan, K. E. (2007) Deep-water Actiniaria from East Pacific hydrothermal vents and cold seeps. Invertebrate Zoology, 4 (1), 83 - 102." type="journal article" year="2007">Sanamyan &amp; Sanamyan (2007)</bibRefCitation>
regarded 
<taxonomicName id="A4D34D7EA7694C72337CFEFE431D6AB1" box="[340,513,1977,2001]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7694C72337CFEFE431D6AB1" box="[340,513,1977,2001]" italics="true" pageId="4" pageNumber="362">Paraphelliactis</emphasis>
</taxonomicName>
as a valid genus after discussing the relation of the number of mesenteries at the margin to that at the limbus in 
<taxonomicName id="A4D34D7EA7694C7233DCFE9A43BD6A95" box="[500,673,2013,2037]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7694C7233DCFE9A43BD6A95" box="[500,673,2013,2037]" italics="true" pageId="4" pageNumber="362">Paraphelliactis</emphasis>
</taxonomicName>
and 
<taxonomicName id="A4D34D7EA7694C7230F1FE9A424D6A95" box="[729,849,2013,2037]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7694C7230F1FE9A424D6A95" box="[729,849,2013,2037]" italics="true" pageId="4" pageNumber="362">Phelliactis</emphasis>
</taxonomicName>
, where 
<taxonomicName id="A4D34D7EA7694C723184FE9A45D36A96" authority="Simon, 1892" authorityName="Simon" authorityYear="1892" box="[940,1231,2013,2038]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="4" pageNumber="362" phylum="Cnidaria" rank="species" species="hertwigi">
<emphasis id="51A7EAEFA7694C723184FE9A452A6A95" box="[940,1078,2013,2037]" italics="true" pageId="4" pageNumber="362">
<collectingCountry id="1BC4766DA7694C723184FE9A42D26A95" box="[940,974,2013,2037]" name="Philippines" pageId="4" pageNumber="362">Ph</collectingCountry>
. hertwigi
</emphasis>
<bibRefCitation id="07424B0CA7694C723617FE9A45D36A96" author="Simon" box="[1087,1231,2013,2038]" pageId="4" pageNumber="362" refString="Simon, J. A. (1892) Ein Beitrag zur Anatomie und Systematik der Hexactinien. Druck von Val. Hofling, Kapellenstrasse 3, Munchen, 102 pp. [pp. 5 - 106]" type="book chapter" year="1892">Simon, 1892</bibRefCitation>
</taxonomicName>
(the 
<typeStatus id="BC68885FA7694C723722FE9944266A96" box="[1290,1338,2014,2038]" pageId="4" pageNumber="362">type</typeStatus>
species) and many other species of 
<taxonomicName id="A4D34D7EA7684C7333E5F9DF43596DD0" box="[461,581,152,176]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7684C7333E5F9DF43596DD0" box="[461,581,152,176]" italics="true" pageId="5" pageNumber="363">Phelliactis</emphasis>
</taxonomicName>
have fewer mesenteries at the margin than at the limbus, while species of 
<taxonomicName id="A4D34D7EA7684C7332BFF9FB40586DB4" box="[151,324,188,212]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7684C7332BFF9FB40586DB4" box="[151,324,188,212]" italics="true" pageId="5" pageNumber="363">Paraphelliactis</emphasis>
</taxonomicName>
have more mesenteries at the margin than at the limbus. 
<bibRefCitation id="07424B0CA7684C7331DEF9FB44076DB4" box="[1014,1307,188,213]" pageId="5" pageNumber="363" refString="Molodtsova, T. N, Sanamyan, N. P. &amp; Keller, N. B. (2008) Anthozoa from the northern Mid-Atlantic Ridge and Charlie-Gibbs Fracture Zone. Marine Biology Research, 4, 112 - 130. http: // dx. doi. org / 10.1080 / 17451000701821744" type="journal article">
Molodtsova 
<emphasis id="51A7EAEFA7684C7336A0F9FA45D96DB4" box="[1160,1221,188,212]" italics="true" pageId="5" pageNumber="363">et al.</emphasis>
(2008)
</bibRefCitation>
reported a specimen, which has almost the equal number of mesenteries at the margin and at the limbus, and identified it as 
<taxonomicName id="A4D34D7EA7684C7332BFF84343756C7C" authority="Carlgren, 1934" authorityName="Carlgren" authorityYear="1934" box="[151,617,260,285]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="species" species="michaelsarsi">
<emphasis id="51A7EAEFA7684C7332BFF84340B86C7C" box="[151,420,260,284]" italics="true" pageId="5" pageNumber="363">Phelliactis michaelsarsi</emphasis>
(
<bibRefCitation id="07424B0CA7684C73339BF843437D6C7C" author="Carlgren" box="[435,609,260,285]" pageId="5" pageNumber="363" refString="Carlgren, O. (1934) Ceriantharia, Zoantharia and Actiniaria from the &quot; Michael Sars &quot; North Atlantic Deep-sea Expedition 1910. Report on the Scientific Results of the &quot; Michael Sars &quot; North Atlantic Deep-Sea Expedition 1910, 5 (6), 1 - 27." type="journal article" year="1934">Carlgren, 1934</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<caption id="37AC6675A7684C7332BFFAB840A36911" httpUri="https://zenodo.org/record/255248/files/figure.png" pageId="5" pageNumber="363" targetBox="[167,1421,339,1007]" targetPageId="5">
<paragraph id="636C36FDA7684C7332BFFAB840A36911" blockId="5.[151,1436,1023,1137]" pageId="5" pageNumber="363">
<emphasis id="51A7EAEFA7684C7332BFFAB840096974" bold="true" box="[151,277,1023,1045]" pageId="5" pageNumber="363">FIGURE 4.</emphasis>
Cnidae from 
<taxonomicName id="A4D34D7EA7684C733381FAB8439D6976" box="[425,641,1023,1046]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="species" species="tangi" status="sp. nov.">
<emphasis id="51A7EAEFA7684C733381FAB8439D6976" box="[425,641,1023,1046]" italics="true" pageId="5" pageNumber="363">Paraphelliactis tangi</emphasis>
</taxonomicName>
<emphasis id="51A7EAEFA7684C7330A2FD4743DA6975" bold="true" box="[650,710,1024,1045]" pageId="5" pageNumber="363">
<taxonomicNameLabel id="4A945794A7684C7330A2FD4743DA6975" box="[650,710,1024,1045]" pageId="5" pageNumber="363" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Tentacles: A. Robust spirocyst. B. Gracile spirocyst. C. Basitrich. D. Basitrich. Column: E. Basitrich. F. Basitrich. G. Microbasic 
<emphasis id="51A7EAEFA7684C733133FD5842346954" box="[795,808,1055,1076]" italics="true" pageId="5" pageNumber="363">p</emphasis>
-mastigophore. Actinopharynx: H. Basitrich. I. Basitrich. J. Microbasic 
<emphasis id="51A7EAEFA7684C73333AFD7940036933" box="[274,287,1086,1107]" italics="true" pageId="5" pageNumber="363">p</emphasis>
-mastigophore. Filament: K. Basitrich. L. Basitrich. M. Microbasic 
<emphasis id="51A7EAEFA7684C7331CEFD7942EE6933" box="[998,1010,1086,1107]" italics="true" pageId="5" pageNumber="363">p</emphasis>
-mastigophore. Acontia: N. Basitrich. O. Basitrich. Scale bar = 20 µm.
</paragraph>
</caption>
<paragraph id="636C36FDA7684C7332EFFDD940F56B23" blockId="5.[151,1437,1182,2035]" pageId="5" pageNumber="363">
The new species 
<taxonomicName id="A4D34D7EA7684C7333A4FDD9436769D6" box="[396,635,1182,1206]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="species" species="tangi" status="sp. nov.">
<emphasis id="51A7EAEFA7684C7333A4FDD9436769D6" box="[396,635,1182,1206]" italics="true" pageId="5" pageNumber="363">Paraphelliactis tangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4A945794A7684C7330ACFDD943D869D7" box="[644,708,1182,1207]" pageId="5" pageNumber="363" rank="species">
<emphasis id="51A7EAEFA7684C7330ACFDD943D869D7" bold="true" box="[644,708,1182,1207]" pageId="5" pageNumber="363">n. sp.</emphasis>
</taxonomicNameLabel>
has two unusual features: a complete fifth cycle of mesenteries and an equal number of mesenteries throughout the column. The former feature is absent in all three known species of 
<taxonomicName id="A4D34D7EA7684C733290FDA14079699E" box="[184,357,1254,1278]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7684C733290FDA14079699E" box="[184,357,1254,1278]" italics="true" pageId="5" pageNumber="363">Paraphelliactis</emphasis>
</taxonomicName>
, but exists in three of 20 known species of 
<taxonomicName id="A4D34D7EA7684C733144FDA142F8699E" box="[876,996,1254,1278]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7684C733144FDA142F8699E" box="[876,996,1254,1278]" italics="true" pageId="5" pageNumber="363">Phelliactis</emphasis>
</taxonomicName>
(
<taxonomicName id="A4D34D7EA7684C7331DFFDA14474699F" authority="Widersten, 1976" authorityName="Widersten" authorityYear="1976" box="[1015,1384,1254,1279]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="species" species="americana">
<emphasis id="51A7EAEFA7684C7331DFFDA14583699E" box="[1015,1183,1254,1278]" italics="true" pageId="5" pageNumber="363">
<collectingCountry id="1BC4766DA7684C7331DFFDA14505699E" box="[1015,1049,1254,1278]" name="Philippines" pageId="5" pageNumber="363">Ph</collectingCountry>
. americana
</emphasis>
Widersten, 1976
</taxonomicName>
, 
<taxonomicName id="A4D34D7EA7684C73375EFDA1432F6843" authority="Riemann-Zurneck, 1973" authorityName="Riemann-Zurneck" authorityYear="1973" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="species" species="callicyclus">
<emphasis id="51A7EAEFA7684C73375EFDA140136842" italics="true" pageId="5" pageNumber="363">
<collectingCountry id="1BC4766DA7684C73375EFDA14484699E" box="[1398,1432,1254,1278]" name="Philippines" pageId="5" pageNumber="363">Ph</collectingCountry>
. callicyclus
</emphasis>
<bibRefCitation id="07424B0CA7684C733333FC4D432F6843" author="Riemann-Zurneck" box="[283,563,1290,1315]" pageId="5" pageNumber="363" refString="Riemann-Zurneck, K. (1973) Actiniaria des Sudwestatlantik I. Hormathiidae. Helgol wiss Meeres, 25, 273 - 325. http: // dx. doi. org / 10.1007 / BF 01611200" type="journal article" year="1973">Riemann-Zürneck, 1973</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="A4D34D7EA7684C73305CFC4D455B6843" authority="Riemann-Zurneck, 1973" authorityName="Riemann-Zurneck" authorityYear="1973" box="[628,1095,1290,1315]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="species" species="lophohelia">
<emphasis id="51A7EAEFA7684C73305CFC4D42036842" box="[628,799,1290,1314]" italics="true" pageId="5" pageNumber="363">
<collectingCountry id="1BC4766DA7684C73305CFC4D438A6842" box="[628,662,1290,1314]" name="Philippines" pageId="5" pageNumber="363">Ph</collectingCountry>
. lophohelia
</emphasis>
<bibRefCitation id="07424B0CA7684C733103FC4D455B6843" author="Riemann-Zurneck" box="[811,1095,1290,1315]" pageId="5" pageNumber="363" refString="Riemann-Zurneck, K. (1973) Actiniaria des Sudwestatlantik I. Hormathiidae. Helgol wiss Meeres, 25, 273 - 325. http: // dx. doi. org / 10.1007 / BF 01611200" type="journal article" year="1973">Riemann-Zürneck, 1973</bibRefCitation>
</taxonomicName>
). The latter feature probably exists also in the three known species of 
<taxonomicName id="A4D34D7EA7684C73304EFC6943C26826" box="[614,734,1326,1350]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7684C73304EFC6943C26826" box="[614,734,1326,1350]" italics="true" pageId="5" pageNumber="363">Phelliactis</emphasis>
</taxonomicName>
and in the specimen identified by 
<bibRefCitation id="07424B0CA7684C73364EFC6944616827" box="[1126,1405,1326,1351]" pageId="5" pageNumber="363" refString="Molodtsova, T. N, Sanamyan, N. P. &amp; Keller, N. B. (2008) Anthozoa from the northern Mid-Atlantic Ridge and Charlie-Gibbs Fracture Zone. Marine Biology Research, 4, 112 - 130. http: // dx. doi. org / 10.1080 / 17451000701821744" type="journal article">Molodtsova et al. (2008)</bibRefCitation>
as 
<taxonomicName id="A4D34D7EA7684C7332BFFC154045680A" box="[151,345,1362,1386]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="species" species="michaelsarsi">
<emphasis id="51A7EAEFA7684C7332BFFC154045680A" box="[151,345,1362,1386]" italics="true" pageId="5" pageNumber="363">
<collectingCountry id="1BC4766DA7684C7332BFFC1541A5680A" box="[151,185,1362,1386]" name="Philippines" pageId="5" pageNumber="363">Ph</collectingCountry>
. michaelsarsi
</emphasis>
</taxonomicName>
. In addition, 
<taxonomicName id="A4D34D7EA7684C73302AFC144373680A" box="[514,623,1363,1386]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="species" species="tangi" status="sp. nov.">
<emphasis id="51A7EAEFA7684C73302AFC144373680A" box="[514,623,1363,1386]" italics="true" pageId="5" pageNumber="363">Pa. tangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4A945794A7684C733054FC1543DE680B" box="[636,706,1362,1387]" pageId="5" pageNumber="363" rank="species">
<emphasis id="51A7EAEFA7684C733054FC1543DE680B" bold="true" box="[636,706,1362,1387]" pageId="5" pageNumber="363">n. sp.</emphasis>
</taxonomicNameLabel>
has another feature not observed in other 
<taxonomicName id="A4D34D7EA7684C7336E3FC154464680A" box="[1227,1400,1362,1386]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7684C7336E3FC154464680A" box="[1227,1400,1362,1386]" italics="true" pageId="5" pageNumber="363">Paraphelliactis</emphasis>
</taxonomicName>
or 
<taxonomicName id="A4D34D7EA7684C7332BFFC31401368EE" box="[151,271,1398,1422]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7684C7332BFFC31401368EE" box="[151,271,1398,1422]" italics="true" pageId="5" pageNumber="363">Phelliactis</emphasis>
</taxonomicName>
, viz., tentacles without mesogloeal thickenings on the aboral side. Nonetheless, the new species is covered with a thick cuticle, a typical feature of the genus 
<taxonomicName id="A4D34D7EA7684C73311DFCDD42FE68D2" box="[821,994,1434,1458]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7684C73311DFCDD42FE68D2" box="[821,994,1434,1458]" italics="true" pageId="5" pageNumber="363">Paraphelliactis</emphasis>
</taxonomicName>
, while in the species of 
<taxonomicName id="A4D34D7EA7684C7336D1FCDD446D68D2" box="[1273,1393,1434,1458]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7684C7336D1FCDD446D68D2" box="[1273,1393,1434,1458]" italics="true" pageId="5" pageNumber="363">Phelliactis</emphasis>
</taxonomicName>
the cuticle is usually thin and is easily stripped off. Thus, we assigned the new species to the genus 
<taxonomicName id="A4D34D7EA7684C7336C7FCF9448068B6" box="[1263,1436,1470,1494]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7684C7336C7FCF9448068B6" box="[1263,1436,1470,1494]" italics="true" pageId="5" pageNumber="363">Paraphelliactis</emphasis>
</taxonomicName>
rather than 
<taxonomicName id="A4D34D7EA7684C73333EFCA54092689A" box="[278,398,1506,1530]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7684C73333EFCA54092689A" box="[278,398,1506,1530]" italics="true" pageId="5" pageNumber="363">Phelliactis</emphasis>
</taxonomicName>
. We extend the generic diagnosis of 
<taxonomicName id="A4D34D7EA7684C733100FCA542C9689A" box="[808,981,1506,1530]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7684C733100FCA542C9689A" box="[808,981,1506,1530]" italics="true" pageId="5" pageNumber="363">Paraphelliactis</emphasis>
</taxonomicName>
as follows to include the new species by stating that the number of tentacles can be equal to that of mesenteries at the limbus and the fifth cycle of mesenteries may be complete.
</paragraph>
<paragraph id="636C36FDA7684C7332EFFF0940C76AE7" blockId="5.[151,1437,1182,2035]" pageId="5" pageNumber="363">
<emphasis id="51A7EAEFA7684C7332EFFF09424C6B07" bold="true" box="[199,848,1613,1639]" pageId="5" pageNumber="363">
Improved diagnosis of 
<taxonomicName id="A4D34D7EA7684C7333CAFF0A424C6B07" authority="Carlgren, 1928" authorityName="Carlgren" authorityYear="1928" box="[482,848,1613,1639]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA7684C7333CAFF0A438C6B06" bold="true" box="[482,656,1613,1638]" italics="true" pageId="5" pageNumber="363">Paraphelliactis</emphasis>
Carlgren, 1928
</taxonomicName>
</emphasis>
(adapted from 
<bibRefCitation id="07424B0CA7684C73362EFF0945B36B07" author="Carlgren" box="[1030,1199,1614,1639]" pageId="5" pageNumber="363" refString="Carlgren, O. (1949) A survey of the Ptychodactaria, Corallimorpharia, and Actiniaria. Kungliga Svenska Vetenskapsakadamiens Handlingar, 1 (1), 1 - 121." type="journal article" year="1949">Carlgren 1949</bibRefCitation>
). 
<taxonomicName id="A4D34D7EA7684C7336EEFF09447C6B07" box="[1222,1376,1614,1639]" class="Anthozoa" family="Hormathiidae" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="family">Hormathiidae</taxonomicName>
with well developed pedal disc. Column divisible into scapus and scapulus, the former strongly tuberculated and provided with a thick cuticle. Sphincter mesogloeal, alveolar. Tentacles arranged in more than five cycles, more than or almost equal to mesenteries at the limbus, with or without mesogloeal thickenings on the aboral side. Longitudinal muscles of tentacles ectodermal, radial muscles of oral disc ectodermal or more or less mesogloeal. Two well developed siphonoglyphs. Mesenteries hexamerously arranged in five cycles, usually six pairs perfect and sterile, and the last cycle incomplete or complete. Retractors of mesenteries diffuse and weak. Parietobasilar muscles weak. Acontia well developed. No cinclides. Cnidom: Robust and gracile spirocysts, basitrichs and microbasic 
<emphasis id="51A7EAEFA7684C733331FE28403B6AE6" box="[281,295,1903,1926]" italics="true" pageId="5" pageNumber="363">p</emphasis>
-mastigophores.
</paragraph>
</subSubSection>
<subSubSection id="2BC96576A7684C7032EFFED541E86805" lastPageId="6" lastPageNumber="364" pageId="5" pageNumber="363" type="reference_group">
<paragraph id="636C36FDA7684C7032EFFED544A66FEF" blockId="5.[151,1437,1182,2035]" lastBlockId="6.[125,1888,630,1562]" lastPageId="6" lastPageNumber="364" pageId="5" pageNumber="363">
<taxonomicName id="A4D34D7EA7684C7332EFFED540A96ACA" ID-CoL="4D7ZF" box="[199,437,1938,1962]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="species" species="tangi" status="sp. nov.">
<emphasis id="51A7EAEFA7684C7332EFFED540A96ACA" box="[199,437,1938,1962]" italics="true" pageId="5" pageNumber="363">Paraphelliactis tangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4A945794A7684C733395FED540E16ACB" box="[445,509,1938,1963]" pageId="5" pageNumber="363" rank="species">
<emphasis id="51A7EAEFA7684C733395FED540E16ACB" bold="true" box="[445,509,1938,1963]" pageId="5" pageNumber="363">n. sp.</emphasis>
</taxonomicNameLabel>
differs from 
<taxonomicName id="A4D34D7EA7684C7330BCFED442D76ACB" ID-CoL="6TR9F" authority="Carlgren, 1928" authorityName="Carlgren" authorityYear="1928" box="[660,971,1938,1963]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="species" species="spinosa">
<emphasis id="51A7EAEFA7684C7330BCFED4420B6ACA" box="[660,791,1939,1962]" italics="true" pageId="5" pageNumber="363">Pa. spinosa</emphasis>
Carlgren, 1928
</taxonomicName>
, 
<typeStatus id="BC68885FA7684C7331FFFED4451B6ACB" box="[983,1031,1939,1963]" pageId="5" pageNumber="363">type</typeStatus>
of the genus, in the presence of the large basitrichs of its mesenterial filaments (vs. absent) and larger basitrichs of acontia (length 49–62 Μm vs. 33.8– 42 Μm) (
<bibRefCitation id="07424B0CA7684C7332DFFE9D40816A93" author="Carlgren" box="[247,413,2010,2035]" pageId="5" pageNumber="363" refString="Carlgren, O. (1942) Actiniaria II. Danish Ingolf-Expedition, 5 (12), 1 - 92." type="journal article" year="1942">Carlgren 1942</bibRefCitation>
). It differs from 
<taxonomicName id="A4D34D7EA7684C73307DFE9C42DC6A93" ID-CoL="4FZ8K" authority="Carlgren, 1934" authorityName="Carlgren" authorityYear="1934" box="[597,960,2010,2035]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="5" pageNumber="363" phylum="Cnidaria" rank="species" species="michaelsarsi">
<emphasis id="51A7EAEFA7684C73307DFE9C420E6A92" box="[597,786,2010,2034]" italics="true" pageId="5" pageNumber="363">Pa. michaelsarsi</emphasis>
<bibRefCitation id="07424B0CA7684C733130FE9D42DC6A93" author="Carlgren" box="[792,960,2010,2035]" pageId="5" pageNumber="363" refString="Carlgren, O. (1934) Ceriantharia, Zoantharia and Actiniaria from the &quot; Michael Sars &quot; North Atlantic Deep-sea Expedition 1910. Report on the Scientific Results of the &quot; Michael Sars &quot; North Atlantic Deep-Sea Expedition 1910, 5 (6), 1 - 27." type="journal article" year="1934">
Carlgren, 
<quantity id="A42B9B18A7684C7331A0FE9C42DC6A93" box="[904,960,2011,2035]" metricMagnitude="1" metricUnit="m" metricValue="4.91236" pageId="5" pageNumber="363" unit="in" value="1934.0">1934</quantity>
</bibRefCitation>
</taxonomicName>
in its smaller body size (maximum 
<emphasis id="51A7EAEFA76B4C703255FB3141F06FEF" bold="true" box="[125,236,630,655]" pageId="6" pageNumber="364">TABLE 2.</emphasis>
Comparison of 
<taxonomicName id="A4D34D7EA76B4C7033B8FB31437B6FF0" ID-CoL="4D7ZF" box="[400,615,630,656]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="6" pageNumber="364" phylum="Cnidaria" rank="species" species="tangi" status="sp. nov.">
<emphasis id="51A7EAEFA76B4C7033B8FB31437B6FF0" box="[400,615,630,656]" italics="true" pageId="6" pageNumber="364">Paraphelliactis tangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4A945794A76B4C703046FB3143BB6FEF" box="[622,679,630,655]" pageId="6" pageNumber="364" rank="species">
<emphasis id="51A7EAEFA76B4C703046FB3143BB6FEF" bold="true" box="[622,679,630,655]" pageId="6" pageNumber="364">n. sp.</emphasis>
</taxonomicNameLabel>
with known species of 
<taxonomicName id="A4D34D7EA76B4C7031BFFB3145CA6FEF" ID-CoL="7PPXY" authority="Carlgren, 1928" authorityName="Carlgren" authorityYear="1928" box="[919,1238,630,656]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="6" pageNumber="364" phylum="Cnidaria" rank="genus">
<emphasis id="51A7EAEFA76B4C7031BFFB3145286FF0" box="[919,1076,630,656]" italics="true" pageId="6" pageNumber="364">Paraphelliactis</emphasis>
Carlgren, 1928
</taxonomicName>
. −, Data not available.
</paragraph>
<paragraph id="636C36FDA76B4C703255FBE347C86FDD" blockId="6.[125,1888,630,1562]" box="[125,1748,676,702]" pageId="6" pageNumber="364">
Characteristics 
<taxonomicName id="A4D34D7EA76B4C703379FBE340B26FDE" ID-CoL="4D7ZF" box="[337,430,676,702]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="6" pageNumber="364" phylum="Cnidaria" rank="species" species="tangi" status="sp. nov.">
<emphasis id="51A7EAEFA76B4C703379FBE340B26FDE" box="[337,430,676,702]" italics="true" pageId="6" pageNumber="364">Pa. tangi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="4A945794A76B4C70339CFBE340F16FDD" box="[436,493,676,701]" pageId="6" pageNumber="364" rank="species">
<emphasis id="51A7EAEFA76B4C70339CFBE340F16FDD" bold="true" box="[436,493,676,701]" pageId="6" pageNumber="364">n. sp.</emphasis>
</taxonomicNameLabel>
<taxonomicName id="A4D34D7EA76B4C7030AEFBE342876FDD" ID-CoL="6TR9F" authority="Carlgren, 1928" authorityName="Carlgren" authorityYear="1928" box="[646,923,676,702]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="6" pageNumber="364" phylum="Cnidaria" rank="species" species="spinosa">
<emphasis id="51A7EAEFA76B4C7030AEFBE343E16FDE" box="[646,765,676,702]" italics="true" pageId="6" pageNumber="364">Pa. spinosa</emphasis>
Carlgren, 1928
</taxonomicName>
<taxonomicName id="A4D34D7EA76B4C703639FBE344466FDD" ID-CoL="4FZ8K" authority="Carlgren, 1934" authorityName="Carlgren" authorityYear="1934" box="[1041,1370,676,702]" class="Anthozoa" family="Hormathiidae" genus="Phelliactis" kingdom="Animalia" order="Actiniaria" pageId="6" pageNumber="364" phylum="Cnidaria" rank="species" species="michaelsarsi">
<emphasis id="51A7EAEFA76B4C703639FBE345A76FDE" box="[1041,1211,676,702]" italics="true" pageId="6" pageNumber="364">Pa. michaelsarsi</emphasis>
<bibRefCitation id="07424B0CA76B4C7036EAFBE344466FDD" author="Carlgren" box="[1218,1370,676,701]" pageId="6" pageNumber="364" refString="Carlgren, O. (1934) Ceriantharia, Zoantharia and Actiniaria from the &quot; Michael Sars &quot; North Atlantic Deep-sea Expedition 1910. Report on the Scientific Results of the &quot; Michael Sars &quot; North Atlantic Deep-Sea Expedition 1910, 5 (6), 1 - 27." type="journal article" year="1934">Carlgren, 1934</bibRefCitation>
</taxonomicName>
<taxonomicName id="A4D34D7EA76B4C7037CAFBE347C86FDD" ID-CoL="4D7ZD" authority="Dunn, 1982" authorityName="Dunn" authorityYear="1982" box="[1506,1748,676,702]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="6" pageNumber="364" phylum="Cnidaria" rank="species" species="pabista">
<emphasis id="51A7EAEFA76B4C7037CAFBE3474A6FDE" box="[1506,1622,676,702]" italics="true" pageId="6" pageNumber="364">Pa. pabista</emphasis>
<bibRefCitation id="07424B0CA76B4C703474FBE347C86FDD" author="Dunn" box="[1628,1748,676,701]" pageId="6" pageNumber="364" refString="Dunn, D. F. (1982) Paraphelliactis pabista, a new species of hormathiid sea anemone from abyssal northeastern Pacific waters (Coelenterata: Actiniaria). Syesis, 15, 51 - 56." type="journal article" year="1982">Dunn, 1982</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="636C36FDA76B4C703255FB9447ED6F8C" blockId="6.[125,1888,630,1562]" box="[125,1777,723,748]" pageId="6" pageNumber="364">Size (mm) Up to 70 high and 74 wide Up to 50 high and wide Up to 140 high, over 100 wide Up to 80 high and 73 wide</paragraph>
<paragraph id="636C36FDA76B4C703255FA47467C6E79" blockId="6.[125,1888,630,1562]" box="[125,1888,768,794]" pageId="6" pageNumber="364">Body shape Sub-cylindrical, pedal disc − Cup shaped, pedal disc narrower than Cup shaped, pedal disc narrower than</paragraph>
<paragraph id="636C36FDA76B4C703379FA6A47326E26" blockId="6.[125,1888,630,1562]" box="[337,1582,813,838]" pageId="6" pageNumber="364">wider than limbus column column</paragraph>
<paragraph id="636C36FDA76B4C703255FA1D472E6E13" blockId="6.[125,1888,630,1562]" box="[125,1586,858,883]" pageId="6" pageNumber="364">Scapulus Distinct Distinct Indistinct Distinct</paragraph>
<paragraph id="636C36FDA76B4C703255FACF47046EC1" blockId="6.[125,1888,630,1562]" box="[125,1560,904,929]" pageId="6" pageNumber="364">Column Solid Solid Spongy Solid</paragraph>
<paragraph id="636C36FDA76B4C703255FAF241F56EAE" blockId="6.[125,1888,630,1562]" box="[125,233,949,974]" pageId="6" pageNumber="364">mesogloea</paragraph>
<paragraph id="636C36FDA76B4C703255FAA546286E9B" blockId="6.[125,1888,630,1562]" box="[125,1844,994,1019]" pageId="6" pageNumber="364">Tubercles Conical; irregularly Acuminated; irregularly arranged Columnar, some with pointed tips; Conical; regularly arranged in 24</paragraph>
<paragraph id="636C36FDA76B4C703379FD48470F6948" blockId="6.[125,1888,630,1562]" box="[337,1555,1039,1064]" pageId="6" pageNumber="364">arranged irregularly arranged rows</paragraph>
<paragraph id="636C36FDA76B4C703255FD7B479D6935" blockId="6.[125,1888,630,1562]" box="[125,1665,1084,1109]" pageId="6" pageNumber="364">Sphincter Moderately strong Strong Weak Weak and short</paragraph>
<paragraph id="636C36FDA76B4C703255FD2D465569E3" blockId="6.[125,1888,630,1562]" box="[125,1865,1130,1155]" pageId="6" pageNumber="364">Tentacles Near 192; unthickened Up to 168; thickened aborally Up to 152; thickened aborally Maximun&gt;150; thickened aborally</paragraph>
<paragraph id="636C36FDA76B4C703379FDD040B869D0" blockId="6.[125,1888,630,1562]" box="[337,420,1175,1200]" pageId="6" pageNumber="364">aborally</paragraph>
<paragraph id="636C36FDA76B4C703255FD83460E69BD" blockId="6.[125,1888,630,1562]" box="[125,1810,1220,1245]" pageId="6" pageNumber="364">Mesenteries Same number distally and More distally than proximally More distally than proximally, one More distally than proximally</paragraph>
<paragraph id="636C36FDA76B4C703379FDB6445C686A" blockId="6.[125,1888,630,1562]" box="[337,1344,1265,1290]" pageId="6" pageNumber="364">proximally specimen about equal number</paragraph>
<paragraph id="636C36FDA76B4C703255FC5847486858" blockId="6.[125,1888,630,1562]" box="[125,1620,1311,1336]" pageId="6" pageNumber="364">Fifth cycle Complete Incomplete Incomplete Incomplete</paragraph>
<paragraph id="636C36FDA76B4C703255FC0B41E86805" blockId="6.[125,1888,630,1562]" box="[125,244,1356,1381]" pageId="6" pageNumber="364">mesenteries</paragraph>
</subSubSection>
<subSubSection id="2BC96576A76B4C713255FC3E43686CE8" lastPageId="7" lastPageNumber="365" pageId="6" pageNumber="364" type="distribution">
<paragraph id="636C36FDA76B4C703255FC3E474C68F2" blockId="6.[125,1888,630,1562]" box="[125,1616,1401,1426]" pageId="6" pageNumber="364">Distribution W Pacific N Atlantic N Atlantic NE Pacific</paragraph>
<paragraph id="636C36FDA76B4C703255FCE144B16B7A" blockId="6.[125,1888,630,1562]" pageId="6" pageNumber="364">
References Present study 
<bibRefCitation id="07424B0CA76B4C7030AEFCE1423A68DF" author="Carlgren" box="[646,806,1446,1471]" pageId="6" pageNumber="364" refString="Carlgren, O. (1928 a) Ceriantharier, Zoantharier och Actiniarier. Meddelelser om Gronland, Supplement 23, 253 - 308." type="book chapter" year="1928" yearSuffix="a">Carlgren 1928a</bibRefCitation>
, 
<bibRefCitation id="07424B0CA76B4C703118FCE1427968DF" author="Carlgren" box="[816,869,1446,1471]" pageId="6" pageNumber="364" refString="Carlgren, O. (1942) Actiniaria II. Danish Ingolf-Expedition, 5 (12), 1 - 92." type="journal article" year="1942">1942</bibRefCitation>
, 
<bibRefCitation id="07424B0CA76B4C703147FCE142B968DF" author="Carlgren" box="[879,933,1446,1471]" pageId="6" pageNumber="364" refString="Carlgren, O. (1949) A survey of the Ptychodactaria, Corallimorpharia, and Actiniaria. Kungliga Svenska Vetenskapsakadamiens Handlingar, 1 (1), 1 - 121." type="journal article" year="1949">1949</bibRefCitation>
; Dunn 
<bibRefCitation id="07424B0CA76B4C703639FCE145BA68DF" author="Carlgren" box="[1041,1190,1446,1471]" pageId="6" pageNumber="364" refString="Carlgren, O. (1949) A survey of the Ptychodactaria, Corallimorpharia, and Actiniaria. Kungliga Svenska Vetenskapsakadamiens Handlingar, 1 (1), 1 - 121." type="journal article" year="1949">Carlgren 1949</bibRefCitation>
; 
<bibRefCitation id="07424B0CA76B4C703698FCE1443968DF" author="Dunn" box="[1200,1317,1446,1471]" pageId="6" pageNumber="364" refString="Dunn, D. F. (1982) Paraphelliactis pabista, a new species of hormathiid sea anemone from abyssal northeastern Pacific waters (Coelenterata: Actiniaria). Syesis, 15, 51 - 56." type="journal article" year="1982">Dunn 1982</bibRefCitation>
; 
<bibRefCitation id="07424B0CA76B4C7037CAFCE1474A68DF" author="Dunn" box="[1506,1622,1446,1471]" pageId="6" pageNumber="364" refString="Dunn, D. F. (1982) Paraphelliactis pabista, a new species of hormathiid sea anemone from abyssal northeastern Pacific waters (Coelenterata: Actiniaria). Syesis, 15, 51 - 56." type="journal article" year="1982">Dunn 1982</bibRefCitation>
; Sanamyan &amp; Sanamyan 1982; 
<bibRefCitation id="07424B0CA76B4C7030EEFC9442EF688C" author="Sanamyan" box="[710,1011,1491,1516]" pageId="6" pageNumber="364" refString="Sanamyan, N. P. &amp; Sanamyan, K. E. (2007) Deep-water Actiniaria from East Pacific hydrothermal vents and cold seeps. Invertebrate Zoology, 4 (1), 83 - 102." type="journal article" year="2007">Sanamyan &amp; Sanamyan 2007</bibRefCitation>
<bibRefCitation id="07424B0CA76B4C703639FC944419688C" author="Riemann-Zurneck" box="[1041,1285,1491,1516]" pageId="6" pageNumber="364" refString="Riemann-Zurneck, K. (1986) On some abyssal sea anemones of the North Atlantic (Actiniaria: Hormathiidae). Mitteilungen aus dem Hamburgischen Zoologischen Museum und Institut, 83, 7 - 29." type="journal article" year="1986">Riemann-Zürneck 1986</bibRefCitation>
; 
<bibRefCitation id="07424B0CA76B4C703738FC944704688C" author="Sanamyan" box="[1296,1560,1491,1516]" pageId="6" pageNumber="364" refString="Sanamyan, N. P. &amp; Sanamyan, K. E. (2007) Deep-water Actiniaria from East Pacific hydrothermal vents and cold seeps. Invertebrate Zoology, 4 (1), 83 - 102." type="journal article" year="2007">Sanamyan &amp; 2007</bibRefCitation>
; 
<bibRefCitation id="07424B0CA76B4C70340BFC944649688C" author="Eash-Loucks" box="[1571,1877,1491,1516]" pageId="6" pageNumber="364" refString="Eash-Loucks, W. E. &amp; Fautin, D. G. (2012) Taxonomy and distribution of sea anemones (Cnidaria: Actiniaria and Corallimorpharia) from deep water of the northeastern Pacific. Zootaxa, 3375, 1 - 80." type="journal article" year="2012">Eash-Loucks&amp; &amp; Fautin 2012</bibRefCitation>
<bibRefCitation id="07424B0CA76B4C703639FF4645AB6B7A" author="Sanamyan" box="[1041,1207,1537,1562]" pageId="6" pageNumber="364" refString="Sanamyan, N. P. &amp; Sanamyan, K. E. (2007) Deep-water Actiniaria from East Pacific hydrothermal vents and cold seeps. Invertebrate Zoology, 4 (1), 83 - 102." type="journal article" year="2007">Sanamyan 2007</bibRefCitation>
; 
<bibRefCitation id="07424B0CA76B4C7036E9FF4644B16B7A" author="Molodtsova" box="[1217,1453,1536,1562]" pageId="6" pageNumber="364" refString="Molodtsova, T. N, Sanamyan, N. P. &amp; Keller, N. B. (2008) Anthozoa from the northern Mid-Atlantic Ridge and Charlie-Gibbs Fracture Zone. Marine Biology Research, 4, 112 - 130. http: // dx. doi. org / 10.1080 / 17451000701821744" type="journal article" year="2008">
Molodtsova 
<emphasis id="51A7EAEFA76B4C703769FF4744686B7A" box="[1345,1396,1536,1562]" italics="true" pageId="6" pageNumber="364">et al.</emphasis>
2008
</bibRefCitation>
</paragraph>
<paragraph id="636C36FDA76A4C7132BFF9D043686CE8" blockId="7.[151,1436,151,392]" pageId="7" pageNumber="365">
height 
<quantity id="A42B9B18A76A4C7132CCF9DF402F6DCF" box="[228,307,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" pageId="7" pageNumber="365" unit="mm" value="70.0">70 mm</quantity>
and width 
<quantity id="A42B9B18A76A4C713399F9DF431C6DCF" box="[433,512,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="7.4" pageId="7" pageNumber="365" unit="mm" value="74.0">74 mm</quantity>
vs. height up to 
<quantity id="A42B9B18A76A4C713094F9DF42056DCF" box="[700,793,152,176]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="7" pageNumber="365" unit="mm" value="140.0">140 mm</quantity>
and width over 
<quantity id="A42B9B18A76A4C7131F8F9DF452D6DD0" box="[976,1073,152,176]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="365" unit="mm" value="100.0">100 mm</quantity>
), the sub-cylindrical body shape (vs. cup shaped with pedal disc narrower than column), the solid mesogloea of column (vs. spongy), distinct scapulus (vs. indistinct), and stronger sphincter. 
<taxonomicName id="A4D34D7EA76A4C713094F9A742B76D97" box="[700,939,224,248]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="7" pageNumber="365" phylum="Cnidaria" rank="species" species="tangi" status="sp. nov.">
<emphasis id="51A7EAEFA76A4C713094F9A742B76D97" box="[700,939,224,248]" italics="true" pageId="7" pageNumber="365">Paraphelliactis tangi</emphasis>
</taxonomicName>
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<emphasis id="51A7EAEFA76A4C71319CF99842E86D98" bold="true" box="[948,1012,223,248]" pageId="7" pageNumber="365">n. sp.</emphasis>
</taxonomicNameLabel>
differs from 
<taxonomicName id="A4D34D7EA76A4C7136A4F9A644806D98" authority="Dunn, 1982" authorityName="Dunn" authorityYear="1982" box="[1164,1436,224,248]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="7" pageNumber="365" phylum="Cnidaria" rank="species" species="pabista">
<emphasis id="51A7EAEFA76A4C7136A4F9A644126D98" box="[1164,1294,224,248]" italics="true" pageId="7" pageNumber="365">Pa. pabista</emphasis>
<bibRefCitation id="07424B0CA76A4C71373EF9A744806D98" author="Dunn" box="[1302,1436,224,248]" pageId="7" pageNumber="365" refString="Dunn, D. F. (1982) Paraphelliactis pabista, a new species of hormathiid sea anemone from abyssal northeastern Pacific waters (Coelenterata: Actiniaria). Syesis, 15, 51 - 56." type="journal article" year="1982">Dunn, 1982</bibRefCitation>
</taxonomicName>
by the sub-cylindrical body shape (vs. cup shaped with pedal disc narrower than column), the irregularly arranged tubercles (vs. regularly arranged), the stronger sphincter, and the larger basitrichs of mesenterial filaments (length 35–60 Μm vs. 23–34 Μm) (Table 2, and references therein). So far, 
<taxonomicName id="A4D34D7EA76A4C713155F80A42FF6C04" box="[893,995,333,356]" class="Anthozoa" family="Hormathiidae" genus="Paraphelliactis" kingdom="Animalia" order="Actiniaria" pageId="7" pageNumber="365" phylum="Cnidaria" rank="species" species="tangi" status="sp. nov.">
<emphasis id="51A7EAEFA76A4C713155F80A42FF6C04" box="[893,995,333,356]" italics="true" pageId="7" pageNumber="365">Pa. tangi</emphasis>
</taxonomicName>
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<emphasis id="51A7EAEFA76A4C7131C2F80B45356C05" bold="true" box="[1002,1065,332,357]" pageId="7" pageNumber="365">n. sp.</emphasis>
</taxonomicNameLabel>
is the fourth species of the genus and the first known in the Western Pacific.
</paragraph>
</subSubSection>
</treatment>
</document>