257 lines
28 KiB
XML
257 lines
28 KiB
XML
<document ID-CLB-Dataset="9619" ID-DOI="http://dx.doi.org/10.3897/zookeys.988.56188" ID-GBIF-Dataset="a38f6648-1c49-486b-a7ee-a877315d4e09" ID-PMC="PMC7666111" ID-Pensoft-Pub="1313-2970-988-63" ID-Pensoft-UUID="D5BC323AA5F05894AE8FBB10A3F0960D" ID-PubMed="33223891" ID-ZooBank="6E64D69BDD734A7EAE2B3CD21247A5E3" ModsDocID="1313-2970-988-63" checkinTime="1604753996846" checkinUser="pensoft" docAuthor="Li, Ya, Lin, Yucheng & Li, Shuqiang" docDate="2020" docId="EAF964568E8A54799AD34A848E56144A" docLanguage="en" docName="ZooKeys 988: 63-128" docOrigin="ZooKeys 988" docSource="http://dx.doi.org/10.3897/zookeys.988.56188" docTitle="Crassignatha quadriventris Li & Lin & Li 2020, comb. nov." docType="treatment" docVersion="6" id="D5BC323AA5F05894AE8FBB10A3F0960D" lastPageNumber="63" masterDocId="D5BC323AA5F05894AE8FBB10A3F0960D" masterDocTitle="A review of Crassignatha (Araneae, Symphytognathidae)" masterLastPageNumber="128" masterPageNumber="63" pageNumber="63" updateTime="1732684104256" updateUser="ExternalLinkService">
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<mods:titleInfo id="B69083390C92FB2563C224B1CD407D5D">
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<mods:title id="D43CDDE86C2DF75135ED5B354DF25B5F">A review of Crassignatha (Araneae, Symphytognathidae)</mods:title>
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<mods:name id="1DCE6F671A0F84120708E95C30823E33" type="personal">
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<mods:roleTerm id="581F99EC27415F873B7BC38FDBD3FF06">Author</mods:roleTerm>
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<mods:namePart id="50FB03E9274964C2FA6A6831F05FAE01">Li, Ya</mods:namePart>
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<mods:affiliation id="53E835ADACCB3E43E1AEBC02EFAC44E2">Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China & The Sichuan Key Laboratory for Conservation Biology of Endangered Wildlife, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
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<mods:nameIdentifier id="2CA9DEE180700CCA88ACA9588C6C7A2E" type="orcid">https://orcid.org/0000-0002-9558-4154</mods:nameIdentifier>
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<mods:name id="CE80AA67914F8961B84D4309237E2DB3" type="personal">
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<mods:roleTerm id="DCC8487FF4323E130822E7C75A33CC16">Author</mods:roleTerm>
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<mods:namePart id="7A6986C24A4220DBE1C063C45CB3424C">Lin, Yucheng</mods:namePart>
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<mods:affiliation id="0052CFD4690D7E0E91E48EB27E8E5E14">Key Laboratory of Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, Sichuan 610064, China & The Sichuan Key Laboratory for Conservation Biology of Endangered Wildlife, Sichuan University, Chengdu, Sichuan 610064, China</mods:affiliation>
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<mods:nameIdentifier id="6D1AA7FDA716330C13B8173C3C39ABF0" type="email">linyucheng@scu.edu.cn</mods:nameIdentifier>
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<mods:roleTerm id="ABE23D367DC2F80F358DE930ECB7BA91">Author</mods:roleTerm>
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<mods:namePart id="2FBC8A3E447439107F2BD37631E0C792">Li, Shuqiang</mods:namePart>
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<mods:affiliation id="88FE8C142EC9B48E22A6A44F86699318">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:nameIdentifier id="638E283A04B2C8606395281E5EAEBF8A" type="orcid">https://orcid.org/0000-0002-3290-5416</mods:nameIdentifier>
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<mods:nameIdentifier id="77E378976FF6A28FE88A68882E0574F1" type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
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<mods:typeOfResource id="7A8CC420E47BF42DAB91C2BE595CDC0F">text</mods:typeOfResource>
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<mods:title id="E927173FBEBC3DE51F7103761ABE70F6">ZooKeys</mods:title>
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<mods:part id="5D8E3EEFFDA82A37A29F5FC0D9D9A8B1">
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<mods:date id="71327719EDF046613EBF4F1C35303948">2020</mods:date>
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<mods:number id="BD5B7AE441F92D158A82AFB83853E802">988</mods:number>
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<mods:start id="57CE299D96C5AADE49EEBB3D3F746404">63</mods:start>
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<mods:end id="E8C8F14E2914F6AD34AD6AC22DDF44BF">128</mods:end>
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<mods:url id="44D1F5629526B66FA853067762332CA0">http://dx.doi.org/10.3897/zookeys.988.56188</mods:url>
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<mods:classification id="79FDA9CF1EBB07315A7CFC27FFC1F79E">journal article</mods:classification>
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<mods:identifier id="34C1216B6A3F04A70F8CE881D149D868" type="DOI">http://dx.doi.org/10.3897/zookeys.988.56188</mods:identifier>
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<treatment id="EAF964568E8A54799AD34A848E56144A" ID-GBIF-Taxon="168821873" LSID="urn:lsid:plazi:treatment:EAF964568E8A54799AD34A848E56144A" httpUri="http://treatment.plazi.org/id/EAF964568E8A54799AD34A848E56144A" lastPageNumber="63" pageId="0" pageNumber="63" scope_family="Symphytognathidae" scope_order="Araneae">
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<subSubSection id="F906125CE354F0CA0F28EF85A1421AFB" pageId="0" pageNumber="63" type="nomenclature">
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<paragraph id="94F29F501EE17DC73E4F72C6DFA19C38" pageId="0" pageNumber="63">
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<taxonomicName id="C39643AD930094BD7853C884DE5217BB" ID-CoL="98ZFD" LSID="EAF96456-8E8A-5479-9AD3-4A848E56144A" authority="(Lin & Li, 2009)" authorityName="Li & Lin & Li" authorityYear="2020" baseAuthorityName="Lin & Li" baseAuthorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha quadriventris" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="quadriventris" status="comb. nov.">Crassignatha quadriventris (Lin & Li, 2009)</taxonomicName>
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<taxonomicNameLabel id="7E27A2B2651AA0EE10BEB6FA822D6B42" pageId="0" pageNumber="63">comb. nov.</taxonomicNameLabel>
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<figureCitation id="722A1CCD2A367984A960391990334861" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Crassignatha quadriventris (Lin & Li, 2009) comb. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Scale bars: 0.50 mm (A-F)." figureDoi="10.3897/zookeys.988.56188.figure20" httpUri="https://binary.pensoft.net/fig/472605" pageId="0" pageNumber="63">Figs 20</figureCitation>
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<figureCitation id="9F71F5731E972EB620E2603ED1D1F070" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Crassignatha quadriventris (Lin & Li, 2009) comb. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure21" httpUri="https://binary.pensoft.net/fig/472606" pageId="0" pageNumber="63">, 21</figureCitation>
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<figureCitation id="59EB0D36E45DC06DFAFA299CCC4849A9" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Distribution records of Crassignatha spp. in Asia. 1 C. baihua sp. nov. 2 C. bangbie sp. nov. 3 C. bicorniventris 4 C. changyan sp. nov. 5 C. danaugirangensis 6 C. dongnai sp. nov. 7 C. ertou 8 C. gucheng sp. nov. 9 C. gudu 10 C. haeneli 11 C. mengla sp. nov. 12 C. nantou sp. nov. 13 C. nasalis sp. nov. 14 C. pianma 15 C. quadriventris 16 C. quanqu 17 C. rostriformis sp. nov. 18 C. shiluensis 19 C. shunani sp. nov. 20 C. si sp. nov. 21 C. thamphra sp. nov. 22 C. xichou sp. nov. 23 C. yamu 24 C. yinzhi." figureDoi="10.3897/zookeys.988.56188.figure38" httpUri="https://binary.pensoft.net/fig/472623" pageId="0" pageNumber="63">, 38</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="3C1661F1DC43C8F4B36B213E6AEA810A" pageId="0" pageNumber="63" type="reference_group">
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<paragraph id="D6A6145B251AF5094CB632250B8B4F2B" pageId="0" pageNumber="63">
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<taxonomicName id="47C9548FAB1845F5A2D7F26FF23B2E8D" authorityName="Lin & Li" authorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu quadriventris" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="quadriventris">Patu quadriventris</taxonomicName>
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Lin & Li, 2009: 55, figs 7A, B, 8A, B, 9A-E, 10A, B (♂♀).
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</paragraph>
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</subSubSection>
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<subSubSection id="9EAFFC8790E2DE64EE466761092B92B3" pageId="0" pageNumber="63" type="type material">
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<paragraph id="02BFF3887483B03DD1B2FAA52E5B59FB" pageId="0" pageNumber="63">Type material.</paragraph>
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<paragraph id="62941D6F9BC2A05D754131088C2AB51E" pageId="0" pageNumber="63">
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<emphasis id="45DCECFF083DEE8115B5828593F294C2" bold="true" italics="true" pageId="0" pageNumber="63">Holotype</emphasis>
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♂ and
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<emphasis id="CEBB803CB752EC26793D47AD31447110" bold="true" italics="true" pageId="0" pageNumber="63">paratypes</emphasis>
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2♂ 2♀ (IZCAS),
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<emphasis id="04882C4CDFAE55CEACD296ACB4355D00" bold="true" pageId="0" pageNumber="63">China</emphasis>
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: Hainan Province, Wuzhishan City, Mt. Wuzhishan Nature Reserve (
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<geoCoordinate id="28E87859A08644795DE37DA5F7250607" degrees="18.90000" direction="north" orientation="latitude" precision="1" value="18.9">18.90000°N</geoCoordinate>
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,
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<geoCoordinate id="5AFBD2E7F76430E945AAA77BD8053787" degrees="109.65000" direction="east" orientation="longitude" precision="1" value="109.65">109.65000°E</geoCoordinate>
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), 9.VIII.2007, S. Li and C. Wang leg.;
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<emphasis id="BBADEB03A73010181FA3988284775364" bold="true" italics="true" pageId="0" pageNumber="63">paratype</emphasis>
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1♀ (IZCAS),
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<emphasis id="D6FA6D6298B7FF5ADA4D24E42400937D" bold="true" pageId="0" pageNumber="63">China</emphasis>
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: Hainan Province, Qiongzhong County, Limushan Nature Reserve (
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<geoCoordinate id="1193B74E8D1C83B726EFB77F8C10BA03" degrees="19.18333" direction="north" orientation="latitude" precision="1" value="19.18333">19.18333°N</geoCoordinate>
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,
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<geoCoordinate id="6CE8B3ADAE97C05D5DCC515B288A5017" degrees="109.73333" direction="east" orientation="longitude" precision="1" value="109.73333">109.73333°E</geoCoordinate>
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; 655 m), 12.VIII.2007, S. Li and C. Wang leg. Examined.
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</paragraph>
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<caption id="4F4CD7259CF21130743F245674F9B81E" doi="10.3897/zookeys.988.56188.figure20" httpUri="https://binary.pensoft.net/fig/472605" pageId="0" pageNumber="63" start="Figure 20" startId="F20">
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<paragraph id="C0237EC3DAA05DADD22FE380CC2CE94C" pageId="0" pageNumber="63">
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<emphasis id="348BC7C643B9006E02578540DC95F792" bold="true" pageId="0" pageNumber="63">Figure 20.</emphasis>
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<taxonomicName id="23696A03D3A92E15D7041E99F0A03466" authorityName="Li & Lin & Li" authorityYear="2020" baseAuthorityName="Lin & Li" baseAuthorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha quadriventris" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="quadriventris">
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<emphasis id="D85C771C1C45768063ED66D50B7D0228" italics="true" pageId="0" pageNumber="63">Crassignatha quadriventris</emphasis>
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</taxonomicName>
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(Lin & Li, 2009) comb. nov.
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<emphasis id="1C09A6D6BA61AADCB22C3AAFD39EE314" bold="true" pageId="0" pageNumber="63">A</emphasis>
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male habitus, dorsal
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<emphasis id="FCA038F9257FB776C3D60882B45BC911" bold="true" pageId="0" pageNumber="63">B</emphasis>
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male habitus, ventral
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<emphasis id="7AE549198E03085C37AE8BB56AB0A243" bold="true" pageId="0" pageNumber="63">C</emphasis>
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male habitus, lateral
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<emphasis id="04578DC182E893F3F9FAE29109916863" bold="true" pageId="0" pageNumber="63">D</emphasis>
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female habitus, dorsal
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<emphasis id="DB13E819FDED87646BAEE8712CF3890E" bold="true" pageId="0" pageNumber="63">E</emphasis>
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female habitus, ventral
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<emphasis id="46D5C0A1B72C2B5279A872EA377A30F4" bold="true" pageId="0" pageNumber="63">F</emphasis>
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female habitus, lateral. Scale bars: 0.50 mm (
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<emphasis id="5DFAD76F5B479B407D7A961FF43F40BB" bold="true" pageId="0" pageNumber="63">A-F</emphasis>
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).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="3CF6A8521316C53E89C483B0D047D94F" pageId="0" pageNumber="63" type="materials_examined">
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<paragraph id="33C07B61FEF779C54953D3FE0677A3B3" pageId="0" pageNumber="63">Other material examined.</paragraph>
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<paragraph id="0A21FACAA33DADE385818DAAE7FFB682" pageId="0" pageNumber="63">
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1♂ 4♀ (IZCAS-Ar 41030-41034),
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<emphasis id="FCF2AEF4C6537417BD900B91E29241AD" bold="true" pageId="0" pageNumber="63">China</emphasis>
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: Hainan Province, Dongfang City, Donghe Town, Nanlang Village, at the foot of Exianling Mountain (
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<geoCoordinate id="B00690CA1EC2F643A56E6D977B10E10E" degrees="19.00633" direction="north" orientation="latitude" precision="1" value="19.00633">19.00633°N</geoCoordinate>
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,
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<geoCoordinate id="7B72FD2E5E0AD6F7D796E24E41D93BC0" degrees="109.08383" direction="east" orientation="longitude" precision="1" value="109.08383">109.08383°E</geoCoordinate>
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; 214 m), 16.XII.2014, Q. Zhao and L. Shao leg.; 1♀ (NHMSU-HA025) used for sequencing, GenBank: MT991990, same data as for preceding.
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</paragraph>
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</subSubSection>
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<subSubSection id="1EFA063570D64F580BD3DD6F89C47DF4" pageId="0" pageNumber="63" type="diagnosis">
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<paragraph id="19BF532D3469A681662AB2F35F111977" pageId="0" pageNumber="63">Diagnosis.</paragraph>
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<paragraph id="686C85D303BA5E30D353338BB9E818DC" pageId="0" pageNumber="63">
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This species differs from other congeneric species except
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<taxonomicName id="A229A8F698945C47B13BC208CE1D3554" lsidName="C. baihua" pageId="0" pageNumber="63" rank="species" species="baihua">
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<emphasis id="F230EB8718F7C9A7BEE7A7BC82768C02" italics="true" pageId="0" pageNumber="63">C. baihua</emphasis>
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</taxonomicName>
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sp. nov. by the short, stiff embolus (Fig.
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<figureCitation id="71C3C5521B6EE30D24B4CFB86EFD927D" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Crassignatha quadriventris (Lin & Li, 2009) comb. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure21" httpUri="https://binary.pensoft.net/fig/472606" pageId="0" pageNumber="63">21A, B</figureCitation>
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). It is most similar to
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<taxonomicName id="EA2B2C6E18C538263F29A9ED88165B0E" lsidName="C. baihua" pageId="0" pageNumber="63" rank="species" species="baihua">
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<emphasis id="F60E65B8C056DBAAEF64DE29D87ABA2F" italics="true" pageId="0" pageNumber="63">C. baihua</emphasis>
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</taxonomicName>
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sp. nov. in the form of the palp and the vulva configuration but can be easily distinguished by the sloped embolic apex, rather than flat as in the latter, and by having six twists of the copulatory ducts, rather than four in the latter (Figs
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<figureCitation id="229E580EA7C908AE7372D03C689118FA" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Crassignatha quadriventris (Lin & Li, 2009) comb. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure21" httpUri="https://binary.pensoft.net/fig/472606" pageId="0" pageNumber="63">21A, B, F, G</figureCitation>
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,
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<figureCitation id="1E09A1E56A36A0E1E75B3F9101CB1A34" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Crassignatha baihua sp. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure2" httpUri="https://binary.pensoft.net/fig/472587" pageId="0" pageNumber="63">2A, B, F, G</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="9499B75B08FBCE0A5087EB7DF98056BB" pageId="0" pageNumber="63" type="description">
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<paragraph id="C855B4F92ADCAD76D821F261075B7F82" pageId="0" pageNumber="63">Description.</paragraph>
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<paragraph id="8FD83DDF718C70B366487CAD12E18C83" pageId="0" pageNumber="63">
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<emphasis id="138DCFE721992ED79289870D07243FCD" bold="true" pageId="0" pageNumber="63">Male</emphasis>
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(IZCAS-Ar 41030). Total length 0.80. Carapace 0.36 long, 0.36 wide, 0.40 high. Clypeus 0.16 high. Sternum 0.28 long, 0.24 wide. Abdomen 0.52 long, 0.48 wide, 0.52 high. Length of legs: I 1.34 (0.42, 0.14, 0.34, 0.20, 0.24); II 1.10 (0.30, 0.14, 0.26, 0.16, 0.24); III 0.90 (0.24, 0.12, 0.18, 0.14, 0.22); IV 1.02 (0.30, 0.12, 0.22, 0.18, 0.20).
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</paragraph>
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<paragraph id="9473781093D97CA17658BB78708B5982" pageId="0" pageNumber="63">
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<emphasis id="D0B2ED6ECD4D9713A18A3289E632206A" bold="true" pageId="0" pageNumber="63">Somatic characters</emphasis>
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(Fig.
|
||
<figureCitation id="0BA4D1348D95FE70B607F2D827B8D936" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Crassignatha quadriventris (Lin & Li, 2009) comb. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Scale bars: 0.50 mm (A-F)." figureDoi="10.3897/zookeys.988.56188.figure20" httpUri="https://binary.pensoft.net/fig/472605" pageId="0" pageNumber="63">20A-C</figureCitation>
|
||
).
|
||
<emphasis id="182752C03F7279B0DD22484D677544AE" bold="true" italics="true" pageId="0" pageNumber="63">Coloration</emphasis>
|
||
: prosoma brown. Legs pale brown. Abdomen dorsally dark, with pale stripes and speckles, ventrally pale brown.
|
||
<emphasis id="37128FC32C1A16EE1CD435F036AB4AC1" bold="true" italics="true" pageId="0" pageNumber="63">Prosoma</emphasis>
|
||
: carapace sub-rounded, granular, sculptured, with two cephalic setae. Cephalic area elevated. Clypeus concave. ALE protruded. PER strongly recurved. Chelicerae covered with few setae anteriorly. Labium tongue shaped, wider than long. Sternum heart shaped, surface rough, slightly plump, truncated posteriorly.
|
||
<emphasis id="F2084C0F416E8B7CC0B3D3C4BDFA6F02" bold="true" italics="true" pageId="0" pageNumber="63">Legs</emphasis>
|
||
: tibia II with two clasping spurs.
|
||
<emphasis id="1A1D99B98B928582DE0E5373DB41A71F" bold="true" italics="true" pageId="0" pageNumber="63">Abdomen</emphasis>
|
||
: anteriorly round, nearly square posteriorly, with sparse, long setae, lateral scutum distinct. Spinnerets slightly sclerotized, surrounded by a circular plate.
|
||
</paragraph>
|
||
<paragraph id="B138C6D9EA0BA0191C9E55A2CA1CE5CB" pageId="0" pageNumber="63">
|
||
<emphasis id="B961B79A59CE29EF4EDFF8849BE9634C" bold="true" italics="true" pageId="0" pageNumber="63">Palp</emphasis>
|
||
(Fig.
|
||
<figureCitation id="20DAE12C9E1152FC4173915269A90B25" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Crassignatha quadriventris (Lin & Li, 2009) comb. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure21" httpUri="https://binary.pensoft.net/fig/472606" pageId="0" pageNumber="63">21A-C</figureCitation>
|
||
): tibia with four distal, dorsal, short setae. Cymbium bears a few distal setae. Apical cymbial tooth hook shaped. Tegulum broad, smooth, globular. Plate-like median apophysis with a sclerotized, longitudinal central ridge line and a prolaterally odontoid process. Embolic membrane arises near anterior part of median apophysis. Embolus short, stiff, basally constricted, mesally wide, distally coracoid.
|
||
</paragraph>
|
||
<paragraph id="15B48C9825433D8F23D6AA01B4C1AD6C" pageId="0" pageNumber="63">
|
||
<emphasis id="22630658B2418F9D25FC3166C4395508" bold="true" pageId="0" pageNumber="63">Female</emphasis>
|
||
(IZCAS-Ar 41031). Total length 1.04. Carapace 0.44 long, 0.40 wide, 0.40 high. Clypeus 0.14 high. Sternum 0.28 long, 0.28 wide. Abdomen 0.72 long, 0.68 wide, 0.80 high. Length of legs: I 1.72 (0.60, 0.18, 0.40, 0.26, 0.28); II 1.42 (0.44, 0.16, 0.34, 0.20, 0.28); III 0.98 (0.26, 0.10, 0.22, 0.16, 0.24); IV 1.18 (0.34, 0.12, 0.28, 0.18, 0.26).
|
||
</paragraph>
|
||
<paragraph id="7CD66E1D1A8F3BEC4B2EC1657C783B35" pageId="0" pageNumber="63">
|
||
<emphasis id="3CC321A3D27EEB2BDE6AA7F366839B04" bold="true" pageId="0" pageNumber="63">Somatic characters</emphasis>
|
||
(Fig.
|
||
<figureCitation id="0A83EB548A3325782F993217C2920D20" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Crassignatha quadriventris (Lin & Li, 2009) comb. nov. A male habitus, dorsal B male habitus, ventral C male habitus, lateral D female habitus, dorsal E female habitus, ventral F female habitus, lateral. Scale bars: 0.50 mm (A-F)." figureDoi="10.3897/zookeys.988.56188.figure20" httpUri="https://binary.pensoft.net/fig/472605" pageId="0" pageNumber="63">20D-F</figureCitation>
|
||
).
|
||
<emphasis id="9BEE6F0D800256E2B7CBDA2A4507C31B" bold="true" italics="true" pageId="0" pageNumber="63">Coloration</emphasis>
|
||
: prosoma and legs as in male. Abdomen dorsally darker than in male, ventrally lighter.
|
||
<emphasis id="3010E5B94BD9D25FF7182C55051E5A61" bold="true" italics="true" pageId="0" pageNumber="63">Prosoma</emphasis>
|
||
: modification and arrangement of eyes as in male, cephalic area lower than in male. Clypeus slightly concave. Mouthparts and sternum as in male.
|
||
<emphasis id="D8679ACA0B94E9006308A473096AEF3B" bold="true" italics="true" pageId="0" pageNumber="63">Abdomen</emphasis>
|
||
: anteriorly round and posteriorly square, surface bears sparse, long setae, lateral scutum and circular plate absent. Spinnerets slightly sclerotized.
|
||
</paragraph>
|
||
<paragraph id="3EF3E692DCEE852238052F58C888BBC0" pageId="0" pageNumber="63">
|
||
<emphasis id="9E1E7DEFDA6D9C1D2C80020B0DF34017" bold="true" italics="true" pageId="0" pageNumber="63">Epigyne</emphasis>
|
||
(Fig.
|
||
<figureCitation id="1562097D8DCC0998297330578B293EFF" captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Crassignatha quadriventris (Lin & Li, 2009) comb. nov. A male palp, prolateral B male palp, ventral C male palp, retrolateral D epigyne, ventral E epigyne, lateral F vulva, ventral G vulva, dorsal. Scale bars: 0.10 mm (A-G)." figureDoi="10.3897/zookeys.988.56188.figure21" httpUri="https://binary.pensoft.net/fig/472606" pageId="0" pageNumber="63">21D-F</figureCitation>
|
||
): epigynal area weakly sclerotized, bears a few setae. Scape developed, protruded, longer than wide. Copulatory openings located at terminus. Internal structures faintly visible via translucent tegument. Paired spermathecae globose, widely separated by at least 1.5
|
||
<normalizedToken id="64587E54DCBE0C7996DDD6C2F4FADD5C" originalValue="×">x</normalizedToken>
|
||
their diameter. Fertilization ducts originating from the lower inside margin of spermathecae, bent downwards and laterally. Copulatory ducts long, connected to dorsal surface of spermathecae, from below spermathecae to the center of vulva, making six bends, then reaching copulatory openings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="08782A33BAC76740A14CCB67ACE83610" pageId="0" pageNumber="63" type="taxonomic justification">
|
||
<paragraph id="6428092B70A632477E1D32AF9A463FB9" pageId="0" pageNumber="63">Taxonomic justification.</paragraph>
|
||
<paragraph id="1E16C4B181B650D99F467DD5220B260F" pageId="0" pageNumber="63">
|
||
The shape of the male palps, the configuration of the epigyne, the modified carapace, and the male abdominal scutum and clasping spurs on tibia II leave no doubt that this species is a member of
|
||
<taxonomicName id="3463E5594F1A96EFF81311717684E7EB" authorityName="Wunderlich" authorityYear="1995" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="69C3650186B12081FFB90A39D1434947" italics="true" pageId="0" pageNumber="63">Crassignatha</emphasis>
|
||
</taxonomicName>
|
||
and not
|
||
<taxonomicName id="B0CA38E740502900752CAAB020D2678E" authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="658BC707498CE4064E32F048BBDA270B" italics="true" pageId="0" pageNumber="63">Patu</emphasis>
|
||
</taxonomicName>
|
||
. Therefore, we propose a new combination,
|
||
<taxonomicName id="D21C1A5FC893F5325F5706D7FC74CB26" lsidName="C. quadriventris" pageId="0" pageNumber="63" rank="species" species="quadriventris">
|
||
<emphasis id="0014D2F105B19AFBEBB853A93C99F2AF" italics="true" pageId="0" pageNumber="63">C. quadriventris</emphasis>
|
||
</taxonomicName>
|
||
(Lin & Li, 2009) comb. nov., transferring it from
|
||
<taxonomicName id="A0BFAAC0500B8156B8BFA6E5C691B07B" authorityName="Marples" authorityYear="1951" class="Arachnida" family="Symphytognathidae" genus="Patu" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Patu" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4C3FE4F5984094A5E88A14BEDEC4AD48" italics="true" pageId="0" pageNumber="63">Patu</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="25BE120F78BF9F33A7A006357F5BB05B" pageId="0" pageNumber="63" type="distribution">
|
||
<paragraph id="E9F4E10E8436DDBE89BB07BA7885B380" pageId="0" pageNumber="63">Distribution.</paragraph>
|
||
<paragraph id="88FBCF25A6B36BABDA0B0D88600ECF1C" pageId="0" pageNumber="63">
|
||
China (Hainan) (Fig.
|
||
<figureCitation id="9EB7CE081BDDAF4944630BB0840437E8" captionStart="Figure 38" captionStartId="F38" captionText="Figure 38. Distribution records of Crassignatha spp. in Asia. 1 C. baihua sp. nov. 2 C. bangbie sp. nov. 3 C. bicorniventris 4 C. changyan sp. nov. 5 C. danaugirangensis 6 C. dongnai sp. nov. 7 C. ertou 8 C. gucheng sp. nov. 9 C. gudu 10 C. haeneli 11 C. mengla sp. nov. 12 C. nantou sp. nov. 13 C. nasalis sp. nov. 14 C. pianma 15 C. quadriventris 16 C. quanqu 17 C. rostriformis sp. nov. 18 C. shiluensis 19 C. shunani sp. nov. 20 C. si sp. nov. 21 C. thamphra sp. nov. 22 C. xichou sp. nov. 23 C. yamu 24 C. yinzhi." figureDoi="10.3897/zookeys.988.56188.figure38" httpUri="https://binary.pensoft.net/fig/472623" pageId="0" pageNumber="63">38</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="444B0C158DC1BE20550429CB570FE1C7" doi="10.3897/zookeys.988.56188.figure21" httpUri="https://binary.pensoft.net/fig/472606" pageId="0" pageNumber="63" start="Figure 21" startId="F21">
|
||
<paragraph id="73F4F87341EA18D868847CAFD1B7D5FC" pageId="0" pageNumber="63">
|
||
<emphasis id="694387CF3DB04147B46C60F6CB7794CE" bold="true" pageId="0" pageNumber="63">Figure 21.</emphasis>
|
||
<taxonomicName id="B0E32057DA9BD6BBDE094E77750FE757" authorityName="Li & Lin & Li" authorityYear="2020" baseAuthorityName="Lin & Li" baseAuthorityYear="2009" class="Arachnida" family="Symphytognathidae" genus="Crassignatha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Crassignatha quadriventris" order="Araneae" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="quadriventris">
|
||
<emphasis id="B022D1B7C092F38AADAE9EAE4AF106F4" italics="true" pageId="0" pageNumber="63">Crassignatha quadriventris</emphasis>
|
||
</taxonomicName>
|
||
(Lin & Li, 2009) comb. nov.
|
||
<emphasis id="B28BDB07648282D29B5A545DBC9D3CA7" bold="true" pageId="0" pageNumber="63">A</emphasis>
|
||
male palp, prolateral
|
||
<emphasis id="937814ED833A1F43D29DFB5C324A4D80" bold="true" pageId="0" pageNumber="63">B</emphasis>
|
||
male palp, ventral
|
||
<emphasis id="18336B3DCDA0208DD499C372BA7716F4" bold="true" pageId="0" pageNumber="63">C</emphasis>
|
||
male palp, retrolateral
|
||
<emphasis id="4B7BF23E2DC398DAC3E2446845E37E96" bold="true" pageId="0" pageNumber="63">D</emphasis>
|
||
epigyne, ventral
|
||
<emphasis id="C7162BB19E9ACDA5221F41EA88C62FEC" bold="true" pageId="0" pageNumber="63">E</emphasis>
|
||
epigyne, lateral
|
||
<emphasis id="684B98DC1DC1F52B7C11104E2FB38C04" bold="true" pageId="0" pageNumber="63">F</emphasis>
|
||
vulva, ventral
|
||
<emphasis id="A1341334D79709DA9B64A78AC6C1ACF0" bold="true" pageId="0" pageNumber="63">G</emphasis>
|
||
vulva, dorsal. Scale bars: 0.10 mm (
|
||
<emphasis id="C21311C4EF65050FF0F99987E5434298" bold="true" pageId="0" pageNumber="63">A-G</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |