treatments-xml/data/EA/3F/13/EA3F133F0B93428C11669D94069C012D.xml
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<mods:title id="052FF9C5D320BCF5D8DC9BEC06D0BF29">Onthophagus cervicornis Kirby, 1825, new synonym under Onthophagus dama (Fabricius, 1798) (Coleoptera, Scarabaeidae, Scarabaeinae)</mods:title>
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<mods:namePart id="946F8CCE37E528C9EBA1C6FE3F1CBF28">Rossini, Michele</mods:namePart>
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<mods:namePart id="22275D0C1C0733854D35C35C382C59FC">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:namePart id="3FDBE03B355FCE72EDA02B0D52D33957">Mann, Darren J.</mods:namePart>
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<paragraph id="D7B0B6F8DD0082EFEB0C4DD32E7A7653" pageId="2" pageNumber="113">Taxon classification Animalia Coleoptera Scarabaeidae</paragraph>
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<subSubSection id="AA19A0A0866B58B9D8C62B30C4DE8484" pageId="2" pageNumber="113" type="nomenclature">
<paragraph id="2CFA18EDF3F49F89C843649C634E0CDA" pageId="2" pageNumber="113">
<taxonomicName id="DB32291DBE3B72EB96BE12249181F939" ID-CoL="74ML7" authority="Fabricius, 1798" authorityName="Fabricius" authorityYear="1798" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onthophagus dama" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="dama">Onthophagus dama (Fabricius, 1798)</taxonomicName>
Fig. 1-4
</paragraph>
</subSubSection>
<subSubSection id="2B7B25133F0F6BB16F13AD13D3ABFFD0" pageId="2" pageNumber="113" type="reference_group">
<paragraph id="929173C231416EF3B57C59543A638F61" pageId="2" pageNumber="113">
<taxonomicName id="D6077D84717011A7634726DE3EB04E14" class="Insecta" family="Scarabaeidae" genus="Copris" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copris dama" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="dama">Copris dama</taxonomicName>
<bibRefCitation id="DBA40F2357ABF50DD44C07886721A8AE" author="Fabricius, JC" journalOrPublisher="Proet and Storch, Hafniae" pageId="4" pageNumber="115" title="Entomologiae systematicae supplementum." year="1798">Fabricius 1798</bibRefCitation>
: 32
</paragraph>
<paragraph id="D5BFA90FD8521B7D30AE6E2993804D1B" pageId="2" pageNumber="113">
<taxonomicName id="4DC624658CCEE33102211DE841EF6A20" class="Insecta" family="Scarabaeidae" genus="Scarabaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scarabaeus aeneus" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="aeneus">Scarabaeus aeneus</taxonomicName>
<bibRefCitation id="217AEBCFA6477E19DE5DB3E788707AA3" author="Olivier, AG" journalOrPublisher="Coleopteres Baudouin (Paris)" pageId="5" pageNumber="116" pagination="1 - 190" title="Entomologie, ou histoire naturelle des insectes, avec leurs caracteres generiques et specifiques, leur description, leur synonymie, et leur figure enluminee." url="10.5962/bhl.title.49479" volume="1" year="1789">Olivier 1789</bibRefCitation>
: 131
</paragraph>
<paragraph id="87E265312F4553CEDBC8283AC7E4A97E" pageId="2" pageNumber="113">
<taxonomicName id="72E96931414BFA93C559C33A5D811AEA" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onthophagus dama" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="dama">Onthophagus dama</taxonomicName>
(
<bibRefCitation id="E2BFE58B1368B46742A64423D7B7A885" author="Fabricius, JC" journalOrPublisher="Proet and Storch, Hafniae" pageId="4" pageNumber="115" title="Entomologiae systematicae supplementum." year="1798">Fabricius 1798</bibRefCitation>
)
<bibRefCitation id="FFDD074606B19D336929360E738DDCE1" author="Arrow, GJ" journalOrPublisher="Taylor and Francis, London" pageId="4" pageNumber="115" title="The Fauna of British India including Ceylon and Burma, Coleoptera, Lamellicornia, Part III. (Coprinae)." year="1931">Arrow 1931</bibRefCitation>
: 280
</paragraph>
<paragraph id="10CC89AD15BCF7086BC0B03AE76F3BA9" pageId="2" pageNumber="113">
<taxonomicName id="A09507274C09416C5D56FEAEC0712366" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onthophagus zubaci" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="zubaci">Onthophagus zubaci</taxonomicName>
<bibRefCitation id="6AEA68E12A5C08C53BC8F028D709D5C7" author="Balthasar, V" journalOrPublisher="Stettiner entomologische Zeitung, Stettin" pageId="4" pageNumber="115" pagination="151 - 153" title="Eine neue Onthophagus-Art aus Ost-Indien." volume="93" year="1932">Balthasar 1932</bibRefCitation>
: 151
</paragraph>
<paragraph id="E305D2491D119AD0B7FDC664A0377386" pageId="2" pageNumber="113">
<taxonomicName id="6A34A55F69600D31702905BB6024F8E2" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onthophagus (Onthophagus) dama" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="dama" subGenus="Onthophagus">Onthophagus (Onthophagus) dama</taxonomicName>
(
<bibRefCitation id="8B227CDA369743119154A10E833B87AC" author="Fabricius, JC" journalOrPublisher="Proet and Storch, Hafniae" pageId="4" pageNumber="115" title="Entomologiae systematicae supplementum." year="1798">Fabricius 1798</bibRefCitation>
)
<bibRefCitation id="CECC39209848DED915083F09A2AA8BE0" author="Balthasar, V" journalOrPublisher="Tschechoslowakische Akademie der Wissenschaften, Prague" pageId="4" pageNumber="115" title="Monographie der Scarabaeidae und Aphodidae der palaearktischen und Orientalischen Region, Coleoptera: Lamellicornia, Band 2, Coprinae (Onitini, Oniticellini, Onthophagini)." year="1963">Balthasar 1963</bibRefCitation>
: 325
</paragraph>
<paragraph id="002E7D16BE694CF6103895BBF95F87E2" pageId="2" pageNumber="113">
<taxonomicName id="F3A337BCB06854BDC85A19814E79953F" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onthophagus cervicornis" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="cervicornis">Onthophagus cervicornis</taxonomicName>
<bibRefCitation id="26FB17A8383ADC5883B47C439A2AE70B" author="Kirby, W" journalOrPublisher="The Transactions of the Linnean Society of London" pageId="5" pageNumber="116" pagination="563 - 572" title="A description of such genera and species of insects, alluded to in the &quot; Introduction to Entomology &quot; of Messrs. Kirby and Spence, as appear not to have been before sufficiently noticed or described." url="10.1111/j.1095-8339.1823.tb00103.x" volume="14" year="1825">Kirby 1825</bibRefCitation>
: 565, syn. n.
</paragraph>
</subSubSection>
<subSubSection id="CF420547E251CCCD49E3BA0BEA0FDAD2" pageId="2" pageNumber="113" type="remarks">
<paragraph id="293D00FF37924319098CAFE449FA651A" pageId="2" pageNumber="113">Remarks.</paragraph>
<paragraph id="21980222963A7B5ED8487A1FB634E67B" pageId="2" pageNumber="113">
As no major nomenclatural concern can affect the current taxonomic status of
<taxonomicName id="A9CFB3AC4EB3232241C616D95C2BB64C" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onthophagus cervicornis" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="cervicornis">Onthophagus cervicornis</taxonomicName>
, we decided to maintain the syntypic status (Art. 73.2.1) for the examined specimens (
<bibRefCitation id="4EC02BE0E969FFA9DB1609E2AEA2EFE3" author="Howden, HF" journalOrPublisher="The International Trust for Zoological Nomenclature, London" pageId="4" pageNumber="115" title="International Code of Zoological Nomenclature. Fourth Edition." year="1999">ICZN 1999</bibRefCitation>
: 74.7.3). Instead, in order to maintain the nomenclature stability for
<taxonomicName id="E5FEFBB5BA85155B8F36FEF512424F66" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onthophagus dama" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="dama">Onthophagus dama</taxonomicName>
, as well as the correct identification of further specimens, a male lectotype is here designated by choosing a name-bearing type specimen.
</paragraph>
</subSubSection>
<subSubSection id="66CC331628A3BD4259C9A73D27B12B37" pageId="2" pageNumber="113" type="geographical distribution">
<paragraph id="43D436F84B7F55A165074F3CF813D6EB" pageId="2" pageNumber="113">Geographical distribution.</paragraph>
<paragraph id="DA90863EC78E7A94BFDDE9F78900FCDE" pageId="2" pageNumber="113">
Nepal, Bhutan, Sri Lanka and India: Chhattisgarh, Haryana, Himachal Pradesh, Karnataka, Madhya Pradesh, Maharashtra, Odisha, Tamil Nadu, Uttar Pradesh, Uttarakhand (
<bibRefCitation id="CD62CA7E0B92F3C921458E819CF1BB56" author="Chandra, K" journalOrPublisher="Journal of Threatened Taxa" pageId="4" pageNumber="115" pagination="4660 - 4671" title="Scarab beetles (Coleoptera Scarabaeoidea) of Barnawapara Wildlife Sanctuary, Chhattisgarh, India." url="10.11609/jott.o3251.4660-71" volume="5" year="2013">Chandra and Gupta 2013</bibRefCitation>
: 4665), and Sikkim (
<bibRefCitation id="5D6BD1895520C4566DD0A4D2A51F9239" author="Balthasar, V" journalOrPublisher="Tschechoslowakische Akademie der Wissenschaften, Prague" pageId="4" pageNumber="115" title="Monographie der Scarabaeidae und Aphodidae der palaearktischen und Orientalischen Region, Coleoptera: Lamellicornia, Band 2, Coprinae (Onitini, Oniticellini, Onthophagini)." year="1963">Balthasar 1963</bibRefCitation>
: 326).
</paragraph>
</subSubSection>
<subSubSection id="190A4CD821140992F4663EFA29D56E58" lastPageId="3" lastPageNumber="114" pageId="2" pageNumber="113" type="type material examined">
<paragraph id="741473BEED07960E97335BA12E395F55" pageId="2" pageNumber="113">Type material examined.</paragraph>
<paragraph id="1EC5E57A6EA775A155D60EDF5487C50E" lastPageId="3" lastPageNumber="114" pageId="2" pageNumber="113">
Syntypes (1♂ + 1♀): 1♂, dry pinned. Original label:
<normalizedToken id="04F96C5CE4B0F0334584391B655EEED6" originalValue="“cervicornis”">&quot;cervicornis&quot;</normalizedToken>
[cream label, W. Kirby black handwritten]; &quot;♂ syntype
<taxonomicName id="305E6813A615C783EDB1CE66C7C001AC" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onthophagus cervicornis" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="cervicornis">Onthophagus cervicornis</taxonomicName>
Kirby 1825: 565 =
<taxonomicName id="7A75611270269B7E857F5E012FCBFD3B" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onthophagus dama" order="Coleoptera" pageId="2" pageNumber="113" phylum="Arthropoda" rank="species" species="dama">Onthophagus dama</taxonomicName>
(F.) Rossini &amp; Mann, 2014&quot; [printed] (MM). 1♀, dry pinned. Original labels:
<normalizedToken id="708FA02FF563CADBD4CC9C1E10548D03" originalValue="“♀”">&quot;&quot;</normalizedToken>
[cream label, black hand
<pageBreakToken id="9A0625067DA8BF3999151969351107DC" pageId="3" pageNumber="114" start="start">written</pageBreakToken>
]/ cervicornis coll. Kirby [cream label with black border, black handwritten] / &quot;Ex. Coll. Hincks &amp; Dibb&quot; [white label, black written printed] / &quot;Manchester Museum, SYNTYPE&quot; [blue label, black written printed] &quot;♀ syntype
<taxonomicName id="9B287AE4D48DA88B9A5698675887E786" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onthophagus cervicornis" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="cervicornis">Onthophagus cervicornis</taxonomicName>
Kirby 1825: 565 =
<taxonomicName id="447FF0374014E9EEFC5DEA5325281E61" class="Insecta" family="Scarabaeidae" genus="Onthophagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Onthophagus dama" order="Coleoptera" pageId="3" pageNumber="114" phylum="Arthropoda" rank="species" species="dama">Onthophagus dama</taxonomicName>
(F.) Rossini &amp; Mann, 2014&quot; [printed] (MM).
</paragraph>
</subSubSection>
<subSubSection id="5F64DB2454E3DDFE9DDFE5E3FFF0B6B0" pageId="3" pageNumber="114" type="ecology">
<paragraph id="91B82744EE107711C7BC4A491B3F2150" pageId="3" pageNumber="114">Ecology.</paragraph>
<paragraph id="9E734A4F48265FDD34C0AEB152B5F3CE" pageId="3" pageNumber="114">
coprophagus, mostly attracted by cattle and human excrements, tunneler, active in any seasons and diurnal (
<bibRefCitation id="21E5FDA01F1ADA3CD7A77D05F480930F" author="Venugopal, KS" journalOrPublisher="Journal of Threatened Taxa" pageId="5" pageNumber="116" pagination="2685 - 2692" title="Diversity and community structure of dung beetles (Coleoptera, Scarabaeinae) associated with semi-urban fragmented agricultural land in the Malabar coast in southern India." url="10.11609/jott.o3074.2685-92" volume="4" year="2012">Venugopal et al. 2012</bibRefCitation>
). Very widespread and abundant in tropical dry forest and agricultural habitats.
</paragraph>
</subSubSection>
</treatment>
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