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<document id="14709C408120633671B52E1B25AD0EB2" ID-CLB-Dataset="53365" ID-DOI="10.11646/zootaxa.5291.1.1" ID-GBIF-Dataset="0da652d5-1d8c-42f0-8b56-88151bbbdb9c" ID-ISSN="1175-5326" ID-Zenodo-Dep="7959311" ID-ZooBank="34500B9B-694C-49EE-A194-609AC6A617C7" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1684795151188" checkinUser="plazi" docAuthor="Okajima, Shûji &amp; Masumoto, Masami" docDate="2023" docId="E96787F4FFE38801FF06F8EAFD7CB82E" docLanguage="en" docName="zootaxa.5291.1.1.pdf" docOrigin="Zootaxa 5291 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5291.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Fernothrips femoralis Okajima &amp; Masumoto 2023, sp. n." docType="treatment" docVersion="6" lastPageNumber="66" masterDocId="155EFF8CFFDC8840FF91FFE1FFACBB3B" masterDocTitle="Six genera of the subtribe Macrothripina from Southeast Asia to Taiwan (Thysanoptera, Phlaeothripidae, Idolothripinae)" masterLastPageNumber="74" masterPageNumber="1" pageNumber="64" updateTime="1699507302642" updateUser="plazi">
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<mods:title id="5674E6A227AB407E5775CFABE1ABAD03">Six genera of the subtribe Macrothripina from Southeast Asia to Taiwan (Thysanoptera, Phlaeothripidae, Idolothripinae)</mods:title>
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<mods:namePart id="57F355B09EB80548C9C89D9AA8D3EC47">Okajima, Shûji</mods:namePart>
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<mods:namePart id="1C41A61BD3FB03F985E5975DCF61EFD7">Masumoto, Masami</mods:namePart>
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<treatment id="E96787F4FFE38801FF06F8EAFD7CB82E" ID-DOI="http://doi.org/10.5281/zenodo.7959491" ID-GBIF-Taxon="209630730" ID-Zenodo-Dep="7959491" LSID="urn:lsid:plazi:treatment:E96787F4FFE38801FF06F8EAFD7CB82E" httpUri="http://treatment.plazi.org/id/E96787F4FFE38801FF06F8EAFD7CB82E" lastPageId="65" lastPageNumber="66" pageId="63" pageNumber="64">
<subSubSection id="29D46569FFE3887FFF06F8EAFE47BC1E" box="[151,491,1803,1830]" pageId="63" pageNumber="64" type="nomenclature">
<paragraph id="617136E2FFE3887FFF06F8EAFE47BC1E" blockId="63.[151,491,1803,1866]" box="[151,491,1803,1830]" pageId="63" pageNumber="64">
<heading id="3A39818EFFE3887FFF06F8EAFE47BC1E" bold="true" box="[151,491,1803,1830]" fontSize="11" level="2" pageId="63" pageNumber="64" reason="4">
<emphasis id="53BAEAF0FFE3887FFF06F8EAFE47BC1E" bold="true" box="[151,491,1803,1830]" pageId="63" pageNumber="64">
<taxonomicName id="A6CE4D61FFE3887FFF06F8EAFE0EBC1D" authority="Okajima &amp; Masumoto, 2023" authorityName="Okajima &amp; Masumoto" authorityYear="2023" box="[151,418,1803,1830]" class="Insecta" family="Phlaeothripidae" genus="Fernothrips" kingdom="Animalia" order="Thysanoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="femoralis" status="sp. nov.">
<emphasis id="53BAEAF0FFE3887FFF06F8EAFE0EBC1D" bold="true" box="[151,418,1803,1830]" italics="true" pageId="63" pageNumber="64">Fernothrips femoralis</emphasis>
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<taxonomicNameLabel id="4889578BFFE3887FFE38F8EAFE47BC1E" box="[425,491,1803,1829]" pageId="63" pageNumber="64" rank="species">sp. n.</taxonomicNameLabel>
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<subSubSection id="29D46569FFE38801FF06F8CEFDA6B906" lastPageId="65" lastPageNumber="66" pageId="63" pageNumber="64" type="description">
<paragraph id="617136E2FFE3887FFF06F8CEFED7BC72" blockId="63.[151,491,1803,1866]" box="[151,379,1839,1866]" pageId="63" pageNumber="64">
(
<figureCitation id="F9F52A67FFE3887FFF0EF8CEFE91BC71" box="[159,317,1839,1866]" captionStart="FIGURES 190–195" captionStartId="64.[151,264,1535,1559]" captionTargetBox="[168,1417,181,1511]" captionTargetId="figure-23@64.[168,1419,181,1511]" captionTargetPageId="64" captionText="FIGURES 190–195. Fernothrips femoralis gen. et sp. n., female. (190) Head &amp; prothorax; (191) antennal segments III–VIII; (192) fore leg; (193) tube; (194) metanotum &amp; pelta; (195) prosternum." figureDoi="http://doi.org/10.5281/zenodo.7959407" httpUri="https://zenodo.org/record/7959407/files/figure.png" pageId="63" pageNumber="64">Figs 190–195</figureCitation>
,
<figureCitation id="F9F52A67FFE3887FFED9F8CEFED8BC72" box="[328,372,1839,1865]" captionStart="FIGURES 235–246" captionStartId="71.[151,264,1413,1437]" captionTargetBox="[168,1420,181,1389]" captionTargetId="figure-17@71.[168,1420,181,1389]" captionTargetPageId="71" captionText="FIGURES 235–246. Macrothripina genera and species. Ethirothrips species 235–243: (235) E. stenomelas, male; (236) E. (?) stenomelas, female from Mindanao; (237) E. tenuis sp. n., female; (238) E. terminalis sp. n., female; (239) E. thai sp. n., female; (240) E. tibialis, female; (241) E. tibialis, micropterous male; (242) E. tibialis, macropterous male; (243) E. virgulae, female. (244) Fernothrips femoralis gen. et sp. n., female. (245) Ischyrothrips crassus, female. (246) Polytrichothrips malayanus sp. n., female." figureDoi="http://doi.org/10.5281/zenodo.7959423" httpUri="https://zenodo.org/record/7959423/files/figure.png" pageId="63" pageNumber="64">244</figureCitation>
)
</paragraph>
<paragraph id="617136E2FFE38801FF06F891FAD5BA26" blockId="63.[151,1436,1904,2039]" lastBlockId="65.[151,1437,151,789]" lastPageId="65" lastPageNumber="66" pageId="63" pageNumber="64">
<emphasis id="53BAEAF0FFE3887FFF06F891FE3CBCB0" box="[151,400,1904,1931]" italics="true" pageId="63" pageNumber="64">Female (macroptera).</emphasis>
<materialsCitation id="D1A63CBFFFE38801FE08F891FAD8BA26" ID-GBIF-Occurrence="4116953382" collectionCode="III, VIII, IV, CPS, IX" lastPageId="65" lastPageNumber="66" pageId="63" pageNumber="64" specimenCode="III 2.8" specimenCount="1" typeStatus="holotype">
Distended body length:
<quantity id="A6369B07FFE3887FFD3CF891FD53BCB0" box="[685,767,1904,1931]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="63" pageNumber="64" unit="mm" value="5.4">5.4mm</quantity>
. Colour uniformly dark brown. Antennal segment
<collectionCode id="07DFAE27FFE3887FFAD4F890FACDBCB0" box="[1349,1377,1905,1931]" pageId="63" pageNumber="64">III</collectionCode>
with extreme base yellowish. All tarsi brown. Fore wings weakly shaded with brown, with a longitudinal brown streak on basal 3/5. Prominent setae almost clear, yellowish. Head (
<figureCitation id="F9F52A67FFE3887FFCA7F859FC3EBCE8" box="[822,914,1976,2003]" captionStart="FIGURES 190–195" captionStartId="64.[151,264,1535,1559]" captionTargetBox="[168,1417,181,1511]" captionTargetId="figure-23@64.[168,1419,181,1511]" captionTargetPageId="64" captionText="FIGURES 190–195. Fernothrips femoralis gen. et sp. n., female. (190) Head &amp; prothorax; (191) antennal segments III–VIII; (192) fore leg; (193) tube; (194) metanotum &amp; pelta; (195) prosternum." figureDoi="http://doi.org/10.5281/zenodo.7959407" httpUri="https://zenodo.org/record/7959407/files/figure.png" pageId="63" pageNumber="64">Fig. 190</figureCitation>
) elongate, 1.96 times as long as broad, broadest across eyes; dorsal surface sculptured with indistinct transverse row of striae, but weakly sculptured with reticulation posteriorly. Cheeks gradually, but distinctly, constricted behind eyes, then swollen at basal 1/3 before sub-basal constriction; each with some small and stout setae, postocular cheek setae 40–50μm. Postocular setae situated behind inner margins of eyes, 140μm apart, much longer than eyes, about 0.4 times as long as head, nearly pointed; ocellar setae minute, postocellar setae situated behind eyes, wider apart from each other, 125μm apart. Eyes bulged, 0.24 times as long as head, 145μm apart from each other; posterior ocelli close to eyes, 32–37μm in diameter, 80μm apart from each other, 42–45μm apart from anterior one. Antennae (
<figureCitation id="F9F52A67FF9C8800FC3EF8A0FBBEBC60" box="[943,1042,1857,1883]" captionStart="FIGURES 190–195" captionStartId="64.[151,264,1535,1559]" captionTargetBox="[168,1417,181,1511]" captionTargetId="figure-23@64.[168,1419,181,1511]" captionTargetPageId="64" captionText="FIGURES 190–195. Fernothrips femoralis gen. et sp. n., female. (190) Head &amp; prothorax; (191) antennal segments III–VIII; (192) fore leg; (193) tube; (194) metanotum &amp; pelta; (195) prosternum." figureDoi="http://doi.org/10.5281/zenodo.7959407" httpUri="https://zenodo.org/record/7959407/files/figure.png" pageId="64" pageNumber="65">Fig. 191</figureCitation>
) about 1.6 times as long as head; segment
<collectionCode id="07DFAE27FF9C8800FF6DF887FE80BCBB" box="[252,300,1894,1920]" pageId="64" pageNumber="65">VIII</collectionCode>
slender, not strongly constricted basally; segment
<specimenCode id="31689E99FF9C8800FCFAF887FC1EBC44" box="[875,946,1893,1920]" collectionCode="III" pageId="64" pageNumber="65">III 2.8</specimenCode>
times as long as broad, a little shorter than segment
<collectionCode id="07DFAE27FF9C8800FF6DF86BFEB7BC9F" box="[252,283,1930,1956]" pageId="64" pageNumber="65">IV</collectionCode>
; sense-cones short and curved. Prothorax (
<figureCitation id="F9F52A67FF9C8800FD6EF868FCCCBC98" box="[767,864,1929,1955]" captionStart="FIGURES 190–195" captionStartId="64.[151,264,1535,1559]" captionTargetBox="[168,1417,181,1511]" captionTargetId="figure-23@64.[168,1419,181,1511]" captionTargetPageId="64" captionText="FIGURES 190–195. Fernothrips femoralis gen. et sp. n., female. (190) Head &amp; prothorax; (191) antennal segments III–VIII; (192) fore leg; (193) tube; (194) metanotum &amp; pelta; (195) prosternum." figureDoi="http://doi.org/10.5281/zenodo.7959407" httpUri="https://zenodo.org/record/7959407/files/figure.png" pageId="64" pageNumber="65">Fig. 190</figureCitation>
) stout; pronotum much longer than half the length of head, 0.68 times as long as head, 1.17 times as broad as long, sculptured with fine reticulation anteriorly and laterally; prominent setae comparatively short, am and ml setae shorter than 40μm. Ferna (
<figureCitation id="F9F52A67FF9C8800FB15F830FB4FBCD7" box="[1156,1251,2001,2028]" captionStart="FIGURES 190–195" captionStartId="64.[151,264,1535,1559]" captionTargetBox="[168,1417,181,1511]" captionTargetId="figure-23@64.[168,1419,181,1511]" captionTargetPageId="64" captionText="FIGURES 190–195. Fernothrips femoralis gen. et sp. n., female. (190) Head &amp; prothorax; (191) antennal segments III–VIII; (192) fore leg; (193) tube; (194) metanotum &amp; pelta; (195) prosternum." figureDoi="http://doi.org/10.5281/zenodo.7959407" httpUri="https://zenodo.org/record/7959407/files/figure.png" pageId="64" pageNumber="65">Fig. 195</figureCitation>
) about 1.2 times as long as broad. Mesopresternum narrow, but invisible in
<typeStatus id="BE758840FF9D8801FC8DFF76FC2EBB8A" box="[796,898,151,177]" pageId="65" pageNumber="66" type="holotype">holotype</typeStatus>
. Metanotum (
<figureCitation id="F9F52A67FF9D8801FB8BFF76FBD4BB8A" box="[1050,1144,151,177]" captionStart="FIGURES 190–195" captionStartId="64.[151,264,1535,1559]" captionTargetBox="[168,1417,181,1511]" captionTargetId="figure-23@64.[168,1419,181,1511]" captionTargetPageId="64" captionText="FIGURES 190–195. Fernothrips femoralis gen. et sp. n., female. (190) Head &amp; prothorax; (191) antennal segments III–VIII; (192) fore leg; (193) tube; (194) metanotum &amp; pelta; (195) prosternum." figureDoi="http://doi.org/10.5281/zenodo.7959407" httpUri="https://zenodo.org/record/7959407/files/figure.png" pageId="65" pageNumber="66">Fig. 194</figureCitation>
) without
<collectionCode id="07DFAE27FF9D8801FB73FF76FABBBB8A" box="[1250,1303,151,177]" country="0" httpUri="http://grbio.org/cool/xgvw-ide6" name="Wyoming-Colorado Paleontological Society" pageId="65" pageNumber="66">CPS</collectionCode>
. Fore wings each with 54–57 duplicated cilia; sub-basal wing setae S1 and S2 minute, S3 elongate. Pelta (
<figureCitation id="F9F52A67FF9D8801FB3BFF5AFAA5BBEE" box="[1194,1289,187,213]" captionStart="FIGURES 190–195" captionStartId="64.[151,264,1535,1559]" captionTargetBox="[168,1417,181,1511]" captionTargetId="figure-23@64.[168,1419,181,1511]" captionTargetPageId="64" captionText="FIGURES 190–195. Fernothrips femoralis gen. et sp. n., female. (190) Head &amp; prothorax; (191) antennal segments III–VIII; (192) fore leg; (193) tube; (194) metanotum &amp; pelta; (195) prosternum." figureDoi="http://doi.org/10.5281/zenodo.7959407" httpUri="https://zenodo.org/record/7959407/files/figure.png" pageId="65" pageNumber="66">Fig. 194</figureCitation>
) 2.5 times as broad as long, with a pair of
<collectionCode id="07DFAE27FF9D8801FE46FF3EFDA0BBC2" box="[471,524,223,249]" country="0" httpUri="http://grbio.org/cool/xgvw-ide6" name="Wyoming-Colorado Paleontological Society" pageId="65" pageNumber="66">CPS</collectionCode>
. Posteromarginal setae on abdominal tergite
<collectionCode id="07DFAE27FF9D8801FB91FF3EFBB2BBC2" box="[1024,1054,223,249]" pageId="65" pageNumber="66">IX</collectionCode>
shorter than tube. Tube (
<figureCitation id="F9F52A67FF9D8801FAA4FF3EFA38BBC2" box="[1333,1428,223,249]" captionStart="FIGURES 190–195" captionStartId="64.[151,264,1535,1559]" captionTargetBox="[168,1417,181,1511]" captionTargetId="figure-23@64.[168,1419,181,1511]" captionTargetPageId="64" captionText="FIGURES 190–195. Fernothrips femoralis gen. et sp. n., female. (190) Head &amp; prothorax; (191) antennal segments III–VIII; (192) fore leg; (193) tube; (194) metanotum &amp; pelta; (195) prosternum." figureDoi="http://doi.org/10.5281/zenodo.7959407" httpUri="https://zenodo.org/record/7959407/files/figure.png" pageId="65" pageNumber="66">Fig. 193</figureCitation>
) weakly convex, distinctly constricted distally, about 1.1 times as long as head, about 4.0 times as long as broad
</materialsCitation>
.
</paragraph>
<caption id="35B1666AFF9C8800FF06FA1EFCC0BD01" ID-DOI="http://doi.org/10.5281/zenodo.7959407" ID-Zenodo-Dep="7959407" httpUri="https://zenodo.org/record/7959407/files/figure.png" pageId="64" pageNumber="65" startId="64.[151,264,1535,1559]" targetBox="[168,1417,181,1511]" targetPageId="64" targetType="figure">
<paragraph id="617136E2FF9C8800FF06FA1EFCC0BD01" blockId="64.[151,1436,1534,1595]" pageId="64" pageNumber="65">
<emphasis id="53BAEAF0FF9C8800FF06FA1EFEC4BD2C" bold="true" box="[151,360,1535,1559]" pageId="64" pageNumber="65">FIGURES 190–195</emphasis>
.
<taxonomicName id="A6CE4D61FF9C8800FEE5FA1EFDF8BD2C" authority="Okajima &amp; Masumoto, 2023" authorityName="Okajima &amp; Masumoto" authorityYear="2023" box="[372,596,1535,1559]" class="Insecta" family="Phlaeothripidae" genus="Fernothrips" kingdom="Animalia" order="Thysanoptera" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="femoralis" status="gen. et sp. n.">
<emphasis id="53BAEAF0FF9C8800FEE5FA1EFDF8BD2C" box="[372,596,1535,1559]" italics="true" pageId="64" pageNumber="65">Fernothrips femoralis</emphasis>
</taxonomicName>
<emphasis id="53BAEAF0FF9C8800FDCBFA1EFD4DBD2C" bold="true" box="[602,737,1535,1559]" pageId="64" pageNumber="65">
<taxonomicNameLabel id="4889578BFF9C8800FDCBFA1EFD4DBD2C" box="[602,737,1535,1559]" pageId="64" pageNumber="65" rank="species">gen. et sp. n.</taxonomicNameLabel>
</emphasis>
, female. (190) Head &amp; prothorax; (191) antennal segments III–VIII; (192) fore leg; (193) tube; (194) metanotum &amp; pelta; (195) prosternum.
</paragraph>
</caption>
<paragraph id="617136E2FF9D8801FF56FEC6FDA6B906" blockId="65.[151,1437,151,789]" pageId="65" pageNumber="66">
<emphasis id="53BAEAF0FF9D8801FF56FEC6FEDABA7A" bold="true" box="[199,374,295,321]" pageId="65" pageNumber="66">Measurements</emphasis>
(
<typeStatus id="BE758840FF9D8801FE14FEC6FE45BA7A" box="[389,489,295,321]" pageId="65" pageNumber="66" type="holotype">holotype</typeStatus>
female in μm). Body length about 5400 (distended). Head length 620, from anterior margin of eyes 585, width across eyes 317, maximum width across cheeks 282, minimum width across cheeks 260, across sub-basal constriction 255; eyes length 150, width 85; postocular setae 250. Antenna total length 1000, segments I–VIII length (width) as follows: 125 (70), 110 (60), 170 (60), 188 (61), 175 (52), 145 (43), 90 (34), 65 (18). Pronotum length 420, width 492. Ferna length about 120, width 100. Setae on prothorax: am less than 20, aa about 50, ml less than 30, pa 100, epim?100. Fore wing length?. Sub-basal wing setae S1 and S2 shorter than 30, S3 180–185. Pelta length 180, width 450. Setae on tergite IX: S1?650, S2?600. Tube length 680, basal width 170, apical width 70; terminal setae?.
</paragraph>
</subSubSection>
<subSubSection id="29D46569FF9D8801FF56FDA6FAD8B95A" box="[199,1396,582,609]" pageId="65" pageNumber="66" type="materials_examined">
<paragraph id="617136E2FF9D8801FF56FDA6FAD8B95A" blockId="65.[151,1437,151,789]" box="[199,1396,582,609]" pageId="65" pageNumber="66">
<emphasis id="53BAEAF0FF9D8801FF56FDA6FEE0B95B" bold="true" box="[199,332,582,609]" pageId="65" pageNumber="66">Type series</emphasis>
.
<materialsCitation id="D1A63CBFFF9D8801FECBFDA6FAD8B95A" ID-GBIF-Occurrence="4116953339" box="[346,1396,582,609]" collectingDate="1992-03-17" collectorName="S. Itoh." location="Keningau" municipality="Borneo" pageId="65" pageNumber="66" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="BE758840FF9D8801FECBFDA6FE6BB95A" box="[346,455,583,609]" pageId="65" pageNumber="66" type="holotype">Holotype</typeStatus>
: macropterous female,
<emphasis id="53BAEAF0FF9D8801FD5BFDA6FC8DB95A" bold="true" box="[714,801,583,609]" pageId="65" pageNumber="66">
<collectingMunicipality id="8115AC98FF9D8801FD5BFDA6FC8DB95A" box="[714,801,583,609]" pageId="65" pageNumber="66">Borneo</collectingMunicipality>
</emphasis>
,
<collectingRegion id="A30AF800FF9D8801FCBEFDA6FCDAB95A" box="[815,886,583,609]" country="Malaysia" name="Sabah" pageId="65" pageNumber="66">Sabah</collectingRegion>
, 10.5mls from
<location id="64116039FF9D8801FBB1FDA6FB3CB95A" LSID="urn:lsid:plazi:treatment:E96787F4FFE38801FF06F8EAFD7CB82E:64116039FF9D8801FBB1FDA6FB3CB95A" box="[1056,1168,583,609]" municipality="Borneo" name="Keningau" pageId="65" pageNumber="66" stateProvince="Sabah">Keningau</location>
,
<date id="15701022FF9D8801FB0BFDA6FABAB95A" box="[1178,1302,583,609]" pageId="65" pageNumber="66" value="1992-03-17">
<collectingDate id="0534E9CAFF9D8801FB0BFDA6FABAB95A" box="[1178,1302,583,609]" pageId="65" pageNumber="66" value="1992-03-17">17.iii.1992</collectingDate>
</date>
,
<collectorName id="CC3B5334FF9D8801FAB3FDA6FAD8B95A" box="[1314,1396,583,609]" pageId="65" pageNumber="66">S. Itoh.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="29D46569FF9D8801FF56FD8AFD7CB82E" pageId="65" pageNumber="66" type="discussion">
<paragraph id="617136E2FF9D8801FF56FD8AFD7CB82E" blockId="65.[151,1437,151,789]" pageId="65" pageNumber="66">
<emphasis id="53BAEAF0FF9D8801FF56FD8AFE98B9BE" bold="true" box="[199,308,619,645]" pageId="65" pageNumber="66">Remarks</emphasis>
. The wings and some prominent setae of the unique
<typeStatus id="BE758840FF9D8801FC03FD8AFC58B9BE" box="[914,1012,619,645]" pageId="65" pageNumber="66" type="holotype">holotype</typeStatus>
female of
<taxonomicName id="A6CE4D61FF9D8801FBE0FD8AFB54B9BE" authorityName="Okajima &amp; Masumoto" authorityYear="2023" box="[1137,1272,619,645]" class="Insecta" family="Phlaeothripidae" genus="Fernothrips" kingdom="Animalia" order="Thysanoptera" pageId="65" pageNumber="66" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="53BAEAF0FF9D8801FBE0FD8AFB54B9BE" box="[1137,1272,619,645]" italics="true" pageId="65" pageNumber="66">F. femoralis</emphasis>
</taxonomicName>
are somewhat damaged by maceration. Furthermore, the right antennal segment III has an inner small sense-cone in addition to two usual ones, and the left antennal segment IV has an outer small sense-cone in addition to four usual ones. There is a possibility that these character states are normal, and the left antennal segment III and right antennal segment IV are missing artificially these small sense-cones.
</paragraph>
</subSubSection>
</treatment>
</document>
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