711 lines
88 KiB
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711 lines
88 KiB
XML
<document id="991D42742C431C41AED40A919D015C6C" ID-CLB-Dataset="37427" ID-DOI="10.5281/zenodo.293461" ID-GBIF-Dataset="ff02939e-3601-42a7-8eda-24e911e4b392" ID-ISSN="1175-5326" ID-Zenodo-Dep="293461" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481813413499" checkinUser="plazi" docAuthor="J. Pablo Jayat, Pablo E. Ortiz, Jorge Salazar-Bravo, Ulyses F. J. Pardiñas & Guillermo D’Elía" docDate="2010" docId="E91F87EDFFA9FF8AD9DA8CE2FE8CFDB2" docLanguage="en" docName="zt02409p061.pdf.imf" docOrigin="Zootaxa 2409" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Akodon boliviensis" docType="treatment" docVersion="8" lastPageNumber="45" masterDocId="1526FF95FF83FFA6D94D8E7FFFD3FF95" masterDocTitle="The Akodon boliviensis species group (Rodentia: Cricetidae: Sigmodontinae) in Argentina: species limits and distribution, with the description of a new entity" masterLastPageNumber="61" masterPageNumber="1" pageNumber="43" updateTime="1698434547521" updateUser="ExternalLinkService">
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<mods:title id="365033D8D74840F9BF34E12B08529FC4">The Akodon boliviensis species group (Rodentia: Cricetidae: Sigmodontinae) in Argentina: species limits and distribution, with the description of a new entity</mods:title>
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<mods:namePart id="62BC42914288E0EFEBB56DDEAD0D3E98">J. Pablo Jayat</mods:namePart>
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<mods:namePart id="0D4F672FEA9368BF0A9A93FC3D4CE693">Pablo E. Ortiz</mods:namePart>
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<mods:namePart id="07AEBEEDF3C2C6DDB9C865D964E4B79C">Jorge Salazar-Bravo</mods:namePart>
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<mods:namePart id="1D1C2FF9136F25356EFC237254BBEDB1">Ulyses F. J. Pardiñas</mods:namePart>
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<mods:namePart id="DBC4E8365D90977052E1EB3778A39A42">Guillermo D’Elía</mods:namePart>
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<mods:title id="0C9BDBC2ECBF6A0E75ADB2031BA512E1">Zootaxa</mods:title>
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<mods:date id="E3C0BE819D3B117D2D6133422951D664">2010</mods:date>
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<mods:number id="199D3DD7EBD31B44A8C4305E870E7B9D">2409</mods:number>
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<treatment id="E91F87EDFFA9FF8AD9DA8CE2FE8CFDB2" ID-DOI="http://doi.org/10.5281/zenodo.6196163" ID-GBIF-Taxon="125137295" ID-Zenodo-Dep="6196163" LSID="urn:lsid:plazi:treatment:E91F87EDFFA9FF8AD9DA8CE2FE8CFDB2" httpUri="http://treatment.plazi.org/id/E91F87EDFFA9FF8AD9DA8CE2FE8CFDB2" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43">
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<subSubSection id="29AC6570FFA9FF8CD9DA8CE2FF35FC2F" pageId="42" pageNumber="43" type="nomenclature">
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<paragraph id="610936FBFFA9FF8CD9DA8CE2FDF8FD22" blockId="42.[151,555,669,695]" box="[151,555,669,695]" pageId="42" pageNumber="43">
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<heading id="3A418197FFA9FF8CD9DA8CE2FDF8FD22" bold="true" box="[151,555,669,695]" fontSize="11" level="1" pageId="42" pageNumber="43" reason="1">
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<emphasis id="53C2EAE9FFA9FF8CD9DA8CE2FDF8FD22" bold="true" box="[151,555,669,695]" pageId="42" pageNumber="43">
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<taxonomicName id="A6B64D78FFA9FF8CD9DA8CE2FEA8FD22" box="[151,379,669,695]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="boliviensis">
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<emphasis id="53C2EAE9FFA9FF8CD9DA8CE2FEA8FD22" bold="true" box="[151,379,669,695]" italics="true" pageId="42" pageNumber="43">Akodon boliviensis</emphasis>
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species group
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="610936FBFFA9FF8CD9DA8C9AFF35FC2F" blockId="42.[151,1437,741,2036]" pageId="42" pageNumber="43">
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The great morphological similarity among species of the
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<taxonomicName id="A6B64D78FFA9FF8CDA2D8C9AFBD0FD6B" box="[864,1027,741,766]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="boliviensis">
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<emphasis id="53C2EAE9FFA9FF8CDA2D8C9AFBD0FD6B" box="[864,1027,741,766]" italics="true" pageId="42" pageNumber="43">A. boliviensis</emphasis>
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</taxonomicName>
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group, as well as high levels of intraspecific variability, has precluded a clear and stable taxonomy for the group and in particular for those forms present in northwestern
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<collectingCountry id="19A1766BFFA9FF8CDB448D4FFD52FCDF" box="[521,641,816,842]" name="Argentina" pageId="42" pageNumber="43">Argentina</collectingCountry>
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. This situation, together with the lack of studies of representative series of specimens, including the
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<typeStatus id="BE0D8859FFA9FF8CDB668D2AFD4EFCFA" box="[555,669,853,879]" pageId="42" pageNumber="43" type="holotype">holotypes</typeStatus>
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and additional specimens collected at the
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<typeStatus id="BE0D8859FFA9FF8CDDC18D2AFB6CFCFA" box="[1164,1215,853,879]" pageId="42" pageNumber="43">type</typeStatus>
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localities, resulted in the suggestion, along the years, of different taxanomic scenarios, sometimes quite distinct among themselves.
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</paragraph>
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</subSubSection>
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<subSubSection id="29AC6570FFA9FF8AD98B8DBAFE8CFDB2" lastPageId="44" lastPageNumber="45" pageId="42" pageNumber="43" type="description">
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<paragraph id="610936FBFFA9FF8CD98B8DBAFABDFBDA" blockId="42.[151,1437,741,2036]" pageId="42" pageNumber="43">
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As in previous instances (e.g.,
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<bibRefCitation id="05274B0AFFA9FF8CDB728DBAFCF0FC4A" author="Patton, J. L. & Da Silva, M. D. F. & Malcom, J. R." box="[575,803,965,991]" journalOrPublisher="Acta Theriologica" pageId="42" pageNumber="43" pagination="185 - 193" part="43" refString="Patton, J. L., Da Silva, M. D. F. & Malcom, J. R. (2000) Mammals of the rio Jurua and the evolutionary and ecological diversification of Amazonia. Bulletin of the American Museum of Natural History, 244, 1 - 306. Piantanida, M. J. & Barrantes, G. E. (1998) Growth studies in Akodon dolores (Rodentia: Muridae) in captivity. Acta Theriologica, 43, 185 - 193." title="Mammals of the rio Jurua and the evolutionary and ecological diversification of Amazonia. Bulletin of the American Museum of Natural History, 244, 1 - 306. Piantanida, M. J. & Barrantes, G. E. (1998) Growth studies in Akodon dolores (Rodentia: Muridae) in captivity" type="journal article" year="2000">
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Patton
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<emphasis id="53C2EAE9FFA9FF8CDBD98DBAFD18FC4B" box="[660,715,965,990]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
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., 2000
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</bibRefCitation>
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||
), the integration of morphologic and molecular data interpreted in a geographic context has proven usefull to assess species boundaries of a taxonomically confusing group. In this regard, the evidence presented herein indicates the existence of four species of the
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<taxonomicName id="A6B64D78FFA9FF8CDC628A6FFF01FBDB" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="boliviensis">
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<emphasis id="53C2EAE9FFA9FF8CDC628A6FFA92FBBC" box="[1327,1345,1040,1065]" italics="true" pageId="42" pageNumber="43">A</emphasis>
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.
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<emphasis id="53C2EAE9FFA9FF8CDC028A6FFF01FBDB" italics="true" pageId="42" pageNumber="43">boliviensis</emphasis>
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</taxonomicName>
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group in northwestern
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<collectingCountry id="19A1766BFFA9FF8CD8AE8A4AFD84FBDA" box="[483,599,1077,1103]" name="Argentina" pageId="42" pageNumber="43">Argentina</collectingCountry>
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with a new species belonging to this group from central
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<collectingCountry id="19A1766BFFA9FF8CDDBC8A4AFAB9FBDA" box="[1265,1386,1077,1103]" name="Argentina" pageId="42" pageNumber="43">Argentina</collectingCountry>
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.
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</paragraph>
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<paragraph id="610936FBFFA9FF8CD98B8A25FB01F931" blockId="42.[151,1437,741,2036]" pageId="42" pageNumber="43">
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The taxonomic status of
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<taxonomicName id="A6B64D78FFA9FF8CD8A58A24FD58FBE1" box="[488,651,1114,1140]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="spegazzinii">
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<emphasis id="53C2EAE9FFA9FF8CD8A58A24FE29FBE1" box="[488,506,1115,1140]" italics="true" pageId="42" pageNumber="43">A</emphasis>
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||
.
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||
<emphasis id="53C2EAE9FFA9FF8CDB448A24FD58FBE1" box="[521,651,1115,1140]" italics="true" pageId="42" pageNumber="43">spegazzinii</emphasis>
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</taxonomicName>
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(including
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<taxonomicName id="A6B64D78FFA9FF8CDA5E8A24FCB6FBE1" box="[787,869,1115,1140]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="alterus">
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<emphasis id="53C2EAE9FFA9FF8CDA5E8A24FCB6FBE1" box="[787,869,1115,1140]" italics="true" pageId="42" pageNumber="43">alterus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="A6B64D78FFA9FF8CDA3E8A24FBCFFBE1" box="[883,1052,1115,1140]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="leucolimnaeus">
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<emphasis id="53C2EAE9FFA9FF8CDA3E8A24FBCFFBE1" box="[883,1052,1115,1140]" italics="true" pageId="42" pageNumber="43">leucolimnaeus</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="A6B64D78FFA9FF8CDD118A24FB28FBE1" box="[1116,1275,1115,1140]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="tucumanensis">
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<emphasis id="53C2EAE9FFA9FF8CDD118A24FB28FBE1" box="[1116,1275,1115,1140]" italics="true" pageId="42" pageNumber="43">tucumanensis</emphasis>
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</taxonomicName>
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) with respect to
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<taxonomicName id="A6B64D78FFA9FF8CD9F88AFFFE87FB0C" box="[181,340,1152,1178]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="boliviensis">
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<emphasis id="53C2EAE9FFA9FF8CD9F88AFFFF14FB0C" box="[181,199,1152,1177]" italics="true" pageId="42" pageNumber="43">A</emphasis>
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.
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<emphasis id="53C2EAE9FFA9FF8CD99A8AFFFE87FB0C" box="[215,340,1152,1177]" italics="true" pageId="42" pageNumber="43">boliviensis</emphasis>
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</taxonomicName>
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is the most debatable of the taxonomic decisions we make. Based on a small series of specimens,
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<bibRefCitation id="05274B0AFFA9FF8CD9AE8ADAFE17FB2A" author="Myers, P. & Patton, J. L. & Smith, M. F." box="[227,452,1189,1215]" journalOrPublisher="Miscellaneous Publications of the Museum of Zoology, University of Michigan" pageId="42" pageNumber="43" pagination="1 - 89" part="177" refString="Myers, P., Patton, J. L. & Smith, M. F. (1990) A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 177, 1 - 89." title="A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia" type="journal article" year="1990">
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Myers
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<emphasis id="53C2EAE9FFA9FF8CD8788ADAFEBBFB2B" box="[309,360,1189,1214]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
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. (1990)
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</bibRefCitation>
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had earlier proposed the valid status of this form. In line, much of the available evidence here considered suggests that
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<taxonomicName id="A6B64D78FFA9FF8CDB108AB4FCD6FB71" box="[605,773,1226,1252]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="spegazzinii">
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||
<emphasis id="53C2EAE9FFA9FF8CDB108AB4FDBCFB71" box="[605,623,1227,1252]" italics="true" pageId="42" pageNumber="43">A</emphasis>
|
||
.
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<emphasis id="53C2EAE9FFA9FF8CDBCE8AB4FCD6FB71" box="[643,773,1227,1252]" italics="true" pageId="42" pageNumber="43">spegazzinii</emphasis>
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</taxonomicName>
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represents a distinct species from
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<taxonomicName id="A6B64D78FFA9FF8CDDF68AB4FA8DFB71" box="[1211,1374,1226,1252]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="boliviensis">
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||
<emphasis id="53C2EAE9FFA9FF8CDDF68AB4FB1EFB71" box="[1211,1229,1227,1252]" italics="true" pageId="42" pageNumber="43">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA9FF8CDDAC8AB4FA8DFB71" box="[1249,1374,1227,1252]" italics="true" pageId="42" pageNumber="43">boliviensis</emphasis>
|
||
</taxonomicName>
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||
. For example, they form reciprocally monophyletic groups (
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<figureCitation id="F98D2A7EFFA9FF8CDA658A8FFCBEFA9F" box="[808,877,1264,1290]" captionStart="FIGURE 1" captionStartId="7.[151,255,1731,1755]" captionTargetBox="[278,1283,182,1711]" captionTargetId="figure@7.[249,1323,152,1743]" captionTargetPageId="7" captionText="FIGURE 1. Results of the phylogenetic analyses based on the first 801 bases of the cyt b gene of 71 sequences of Akodon. Topotype specimens included in the analyses are marked as follows: (a) A. alterus; (b) A. puer caenosus; (c) A. leucolimnaeus; (d) A. polopi new species (including the holotype); (e) A. spegazzinii; (f) A. sylvanus; and (g) A. tucumanensis. For those sequences retrieved from Genbank, accession numbers are provided next to species labels. For those sequences gathered by us, specimen collection numbers are provided next to species label; see Appendix I for locality data and Genbank accession numbers. A) Strict consensus tree of the 220 most parsimonious trees (length 1563, CI = 0.287, RI = 0.675). Numbers indicate parsimony jackknife values of the nodes at their right. Only jackknife values above 50 % are shown. B) Majority-rule consensus resulting from the Bayesian analysis. Numbers indicate posterior probability values of the nodes at their right." httpUri="https://zenodo.org/record/293462/files/figure.png" pageId="42" pageNumber="43">Fig. 1</figureCitation>
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) that are allopatrically distributed; in addition, they are clearly identifiable in morphometric multivariate space and morphological grounds (see comparison above). Moreover, these species have different karyotypes, including the presence of heteromorphism in the largest autosomal pair and the X chromosome in
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<taxonomicName id="A6B64D78FFA9FF8CDB9C8B1FFCA1FAEC" box="[721,882,1376,1402]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="spegazzinii">
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||
<emphasis id="53C2EAE9FFA9FF8CDB9C8B1FFD30FAEC" box="[721,739,1376,1401]" italics="true" pageId="42" pageNumber="43">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA9FF8CDBBD8B1FFCA1FAEC" box="[752,882,1376,1401]" italics="true" pageId="42" pageNumber="43">spegazzinii</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="05274B0AFFA9FF8CDACC8B1FFB4DFAEF" author="Gardner, A. L. & Patton, J. L." box="[897,1182,1376,1402]" journalOrPublisher="Occasional Papers of the Museum of Zoology, Louisiana State University" pageId="42" pageNumber="43" pagination="1 - 48" part="49" refString="Gardner, A. L. & Patton, J. L. (1976) Karyotypic variation in oryzomyne rodents (Cricetinae) with comments on chromosomal evolution in the Neotropical cricetine complex. Occasional Papers of the Museum of Zoology, Louisiana State University, 49, 1 - 48." title="Karyotypic variation in oryzomyne rodents (Cricetinae) with comments on chromosomal evolution in the Neotropical cricetine complex" type="journal article" year="1976">Gardner & Patton, 1976</bibRefCitation>
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||
;
|
||
<bibRefCitation id="05274B0AFFA9FF8CDDE48B1FFA44FAEF" author="Barquez, R. M. & Williams, D. F. & Mares, M. A. & Genoways, H. H." box="[1193,1431,1376,1402]" journalOrPublisher="Annals of Carnegie Museum" pageId="42" pageNumber="43" pagination="379 - 403" part="49" refString="Barquez, R. M., Williams, D. F., Mares, M. A. & Genoways, H. H. (1980) Karyology and morphometrics of three species of Akodon (Mammalia: Muridae) from northwestern Argentina. Annals of Carnegie Museum, 49, 379 - 403." title="Karyology and morphometrics of three species of Akodon (Mammalia: Muridae) from northwestern Argentina" type="journal article" year="1980">
|
||
Barquez
|
||
<emphasis id="53C2EAE9FFA9FF8CDC5C8B1FFA97FAEC" box="[1297,1348,1376,1401]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
., 1980
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="05274B0AFFA9FF8CD9DA8BFAFEA5FA0A" author="Myers, P. & Patton, J. L. & Smith, M. F." box="[151,374,1413,1439]" journalOrPublisher="Miscellaneous Publications of the Museum of Zoology, University of Michigan" pageId="42" pageNumber="43" pagination="1 - 89" part="177" refString="Myers, P., Patton, J. L. & Smith, M. F. (1990) A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 177, 1 - 89." title="A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia" type="journal article" year="1990">
|
||
Myers
|
||
<emphasis id="53C2EAE9FFA9FF8CD9A68BFAFECCFA0B" box="[235,287,1413,1438]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
., 1990
|
||
</bibRefCitation>
|
||
), and different ecological preferences. Our records indicate disjunct distribution for these forms, with
|
||
<taxonomicName id="A6B64D78FFA9FF8CD8678BD4FE19FA51" box="[298,458,1450,1476]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="boliviensis">
|
||
<emphasis id="53C2EAE9FFA9FF8CD8678BD4FEEFFA51" box="[298,316,1451,1476]" italics="true" pageId="42" pageNumber="43">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA9FF8CD8008BD4FE19FA51" box="[333,458,1451,1476]" italics="true" pageId="42" pageNumber="43">boliviensis</emphasis>
|
||
</taxonomicName>
|
||
restricted to high altitudinal grasslands in the extreme north of the region and
|
||
<taxonomicName id="A6B64D78FFA9FF8CDCCE8BD4FEC9FA7C" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="spegazzinii">
|
||
<emphasis id="53C2EAE9FFA9FF8CDCCE8BD4FA46FA51" box="[1411,1429,1451,1476]" italics="true" pageId="42" pageNumber="43">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA9FF8CD9DA8BAFFEC9FA7C" box="[151,282,1488,1513]" italics="true" pageId="42" pageNumber="43">spegazzinii</emphasis>
|
||
</taxonomicName>
|
||
extended over many habitats and large areas in south-central portions of the region. In spite of these differences, the genetic divergence betweem them was the smallest among all species pairs of the
|
||
<taxonomicName id="A6B64D78FFA9FF8CDCCE8B8AFEC7F9A1" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="boliviensis">
|
||
<emphasis id="53C2EAE9FFA9FF8CDCCE8B8AFA46F99B" box="[1411,1429,1525,1550]" italics="true" pageId="42" pageNumber="43">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA9FF8CD9DA8864FEC7F9A1" box="[151,276,1563,1588]" italics="true" pageId="42" pageNumber="43">boliviensis</emphasis>
|
||
</taxonomicName>
|
||
group (
|
||
<tableCitation id="2C340340FFA9FF8CD8398865FE32F9A1" box="[372,481,1562,1588]" captionStart="TABLE 12" captionStartId="22.[151,239,151,175]" captionTargetBox="[165,1421,351,713]" captionText="TABLE 12. Average percentage of observerd sequence divergence within and between seven species of the Akodon boliviensis species group. Standard error estimates are shown between parentheses and were obtained by a bootstrap procedure (500 replicates) in MEGA 4. Positions containing missing data were excluded only in pairwise sequence comparisons. Intraspecific values for A. lutescens and A. subfuscus were not calculated because a single sequence was available for each species." httpUri="http://table.plazi.org/id/35C96673FF95FFB0D9DA8EE8FE63FEAC" pageId="42" pageNumber="43" tableUuid="35C96673FF95FFB0D9DA8EE8FE63FEAC">Table 12</tableCitation>
|
||
). In addition, some overlap in morphological and morphometric characters was evident. Moreover, we could not evaluate the constancy of these differences because we failed to capture both species in sympatry. So, additional studies are needed in the north of the northwestern Argentinean region and in possible areas of sympatry to test the constancy of the observed differences here reported.
|
||
</paragraph>
|
||
<paragraph id="610936FBFFA9FF8DD98B88CFFCBEFDE7" blockId="42.[151,1437,741,2036]" lastBlockId="43.[151,1437,152,2007]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
|
||
The taxonomic status of
|
||
<taxonomicName id="A6B64D78FFA9FF8CD8AB88CFFD76F95C" box="[486,677,1712,1738]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="tucumanensis">
|
||
<emphasis id="53C2EAE9FFA9FF8CD8AB88CFFE2BF95C" box="[486,504,1712,1737]" italics="true" pageId="42" pageNumber="43">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA9FF8CDB4B88CFFD76F95C" box="[518,677,1712,1737]" italics="true" pageId="42" pageNumber="43">tucumanensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A6B64D78FFA9FF8CDBF988CFFCF6F95C" box="[692,805,1712,1738]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="alterus">
|
||
<emphasis id="53C2EAE9FFA9FF8CDBF988CFFD15F95C" box="[692,710,1712,1737]" italics="true" pageId="42" pageNumber="43">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA9FF8CDB9988CFFCF6F95C" box="[724,805,1712,1737]" italics="true" pageId="42" pageNumber="43">alterus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="A6B64D78FFA9FF8CDA2888CFFBFDF95C" box="[869,1070,1712,1738]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="leucolimnaeus">
|
||
<emphasis id="53C2EAE9FFA9FF8CDA2888CFFCA4F95C" box="[869,887,1712,1737]" italics="true" pageId="42" pageNumber="43">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA9FF8CDAC888CFFBFDF95C" box="[901,1070,1712,1737]" italics="true" pageId="42" pageNumber="43">leucolimnaeus</emphasis>
|
||
</taxonomicName>
|
||
remained controversial despite the pioneering work by
|
||
<bibRefCitation id="05274B0AFFA9FF8CD8F888AAFD73F97A" author="Myers, P. & Patton, J. L. & Smith, M. F." box="[437,672,1749,1775]" journalOrPublisher="Miscellaneous Publications of the Museum of Zoology, University of Michigan" pageId="42" pageNumber="43" pagination="1 - 89" part="177" refString="Myers, P., Patton, J. L. & Smith, M. F. (1990) A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 177, 1 - 89." title="A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia" type="journal article" year="1990">
|
||
Myers
|
||
<emphasis id="53C2EAE9FFA9FF8CDB4488AAFDEDF97B" box="[521,574,1749,1774]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. (1990)
|
||
</bibRefCitation>
|
||
. While all these forms were treated as subspecies or allied to
|
||
<taxonomicName id="A6B64D78FFA9FF8CDCCE88AAFEC9F881" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="spegazzinii">
|
||
<emphasis id="53C2EAE9FFA9FF8CDCCE88AAFA46F97B" box="[1411,1429,1749,1774]" italics="true" pageId="42" pageNumber="43">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA9FF8CD9DA8884FEC9F881" box="[151,282,1787,1812]" italics="true" pageId="42" pageNumber="43">spegazzinii</emphasis>
|
||
</taxonomicName>
|
||
by some authors (e.g.
|
||
<bibRefCitation id="05274B0AFFA9FF8CDB6D8885FD23F881" author="Myers, P. & Patton, J. L. & Smith, M. F." box="[544,752,1786,1812]" journalOrPublisher="Miscellaneous Publications of the Museum of Zoology, University of Michigan" pageId="42" pageNumber="43" pagination="1 - 89" part="177" refString="Myers, P., Patton, J. L. & Smith, M. F. (1990) A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 177, 1 - 89." title="A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia" type="journal article" year="1990">
|
||
Myers
|
||
<emphasis id="53C2EAE9FFA9FF8CDB3C8884FD70F881" box="[625,675,1787,1812]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. 1990
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="05274B0AFFA9FF8CDBB68885FC6BF881" author="Diaz, M. M. & Braun, J. K. & Mares, M. A. & Barquez, R. M." box="[763,952,1786,1812]" journalOrPublisher="Occasional Papers of the Oklahoma Museum of Natural History" pageId="42" pageNumber="43" pagination="1 - 52" part="10" refString="Diaz, M. M., Braun, J. K., Mares, M. A. & Barquez, R. M. (2000) An update of the taxonomy, systematics, and distribution of the mammals of Salta province, Argentina. Occasional Papers of the Oklahoma Museum of Natural History, 10, 1 - 52." title="An update of the taxonomy, systematics, and distribution of the mammals of Salta province, Argentina" type="journal article" year="2000">
|
||
Díaz
|
||
<emphasis id="53C2EAE9FFA9FF8CDA748884FCBFF881" box="[825,876,1787,1812]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. 2000
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="05274B0AFFA9FF8CDA8E8885FB7EF881" author="Jayat, J. P. & Ortiz, P. E. & Teta, P. & Pardinas, U. F. J. & D'Elia, G." box="[963,1197,1786,1812]" journalOrPublisher="Mastozoologia Neotropical" pageId="42" pageNumber="43" pagination="51 - 67" part="13" refString="Jayat, J. P., Ortiz, P. E., Teta, P., Pardinas, U. F. J. & D'Elia, G. (2006) Nuevas localidades argentinas para algunos roedores sigmodontinos (Rodentia: Cricetidae). Mastozoologia Neotropical, 13, 51 - 67." title="Nuevas localidades argentinas para algunos roedores sigmodontinos (Rodentia: Cricetidae)" type="journal article" year="2006">
|
||
Pardiñas
|
||
<emphasis id="53C2EAE9FFA9FF8CDD638884FBB2F881" box="[1070,1121,1787,1812]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. 2006
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="05274B0AFFA9FF8CDDF58885FA5CF881" author="Jayat, J. P. & Ortiz, P. E. & Miotti, M. D." box="[1208,1423,1786,1812]" journalOrPublisher="Acta Zoologica Mexicana" pageId="42" pageNumber="43" pagination="137 - 177" part="24" refString="Jayat, J. P., Ortiz, P. E. & Miotti, M. D. (2008 a) Distribucion de sigmodontinos (Rodentia: Cricetidae) en pastizales de neblina del noroeste argentino. Acta Zoologica Mexicana, 24, 137 - 177." title="Distribucion de sigmodontinos (Rodentia: Cricetidae) en pastizales de neblina del noroeste argentino" type="journal article" year="2008" yearSuffix="a">
|
||
Jayat
|
||
<emphasis id="53C2EAE9FFA9FF8CDDB68884FAFDF881" box="[1275,1326,1787,1812]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
., 2008a
|
||
</bibRefCitation>
|
||
), others considered them as valid species (e.g.
|
||
<bibRefCitation id="05274B0AFFA9FF8CDBF4895FFC61F8AF" author="Barquez, R. M. & Mares, M. A. & Ojeda, R. A." bookContentInfo="282 pp." box="[697,946,1824,1850]" journalOrPublisher="Oklahoma Museum of Natural History, University of Oklahoma, Norman" pageId="42" pageNumber="43" refString="Barquez, R. M., Mares, M. A. & Ojeda, R. A. (1991) Mamiferos de Tucuman - Mammals of Tucuman. Oklahoma Museum of Natural History, University of Oklahoma, Norman, 282 pp." title="Mamiferos de Tucuman - Mammals of Tucuman" type="book" year="1991">
|
||
Barquez
|
||
<emphasis id="53C2EAE9FFA9FF8CDA68895FFC88F8AC" box="[805,859,1824,1849]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
., 1991
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="05274B0AFFA9FF8CDA8D895FFB73F8AF" author="Diaz, M. M. & Braun, J. K. & Mares, M. A. & Barquez, R. M." box="[960,1184,1824,1850]" journalOrPublisher="Occasional Papers of the Oklahoma Museum of Natural History" pageId="42" pageNumber="43" pagination="1 - 10" part="2" refString="Diaz, M. M., Braun, J. K., Mares, M. A. & Barquez, R. M. (1997) Key of Mammals of Salta Province, Argentina. Occasional Papers of the Oklahoma Museum of Natural History, 2, 1 - 10." title="Key of Mammals of Salta Province, Argentina" type="journal article" year="1997">
|
||
Mares
|
||
<emphasis id="53C2EAE9FFA9FF8CDD5E895FFB83F8AC" box="[1043,1104,1824,1849]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
|
||
, 1997
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="05274B0AFFA9FF8CDDE3895FFAE6F8AF" author="Diaz, M. M." bookContentInfo="640 pp." box="[1198,1333,1824,1850]" journalOrPublisher="Thesis, Universidad Nacional de Tucuman, Argentina" pageId="42" pageNumber="43" refString="Diaz, M. M. (1999) Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil. Thesis, Universidad Nacional de Tucuman, Argentina, 640 pp." title="Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil" type="book" year="1999">Díaz, 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="05274B0AFFA9FF8CDC0E895FFE97F8CA" author="Diaz, M. M. & Barquez, R. M." editor="Kelt, D. A." journalOrPublisher="University of California Publications in Zoology 134" pageId="42" pageNumber="43" pagination="417 - 578" refString="Diaz, M. M. & Barquez, R. M. (2007) The Wild Mammals of Jujuy Province, Argentina: Systematics and Distribution. In: Kelt, D. A., Lessa, E. P., Salazar-Bravo, J. & Patton, J. L. (Eds.), The Quintessential Naturalist: Honoring the Life and Legacy of Oliver P. Pearson. University of California Publications in Zoology 134, pp. 417 - 578." title="The Wild Mammals of Jujuy Province, Argentina: Systematics and Distribution" type="book chapter" volumeTitle="The Quintessential Naturalist: Honoring the Life and Legacy of Oliver P. Pearson" year="2007">Díaz & Barquez, 2007</bibRefCitation>
|
||
; but see
|
||
<bibRefCitation id="05274B0AFFA9FF8CD8E5893AFD59F8CA" author="Galliari, C. A. & Pardinas, U. F. J. & Goin, F. J." box="[424,650,1861,1887]" journalOrPublisher="Mastozoologia Neotropical" pageId="42" pageNumber="43" pagination="39 - 61" part="3" refString="Galliari, C. A., Pardinas, U. F. J. & Goin, F. J. (1996) Lista comentada de los mamiferos argentinos. Mastozoologia Neotropical, 3, 39 - 61." title="Lista comentada de los mamiferos argentinos" type="journal article" year="1996">
|
||
Galliari
|
||
<emphasis id="53C2EAE9FFA9FF8CDB45893AFDE9F8CB" box="[520,570,1861,1886]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
., 1996
|
||
</bibRefCitation>
|
||
about the status of
|
||
<taxonomicName id="A6B64D78FFA9FF8CDA26893AFBF9F8CB" box="[875,1066,1861,1887]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="tucumanensis">
|
||
<emphasis id="53C2EAE9FFA9FF8CDA26893AFCAEF8CB" box="[875,893,1861,1886]" italics="true" pageId="42" pageNumber="43">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA9FF8CDAC6893AFBF9F8CB" box="[907,1066,1861,1886]" italics="true" pageId="42" pageNumber="43">tucumanensis</emphasis>
|
||
</taxonomicName>
|
||
). In spite of the extended usage of the specific epithet
|
||
<taxonomicName id="A6B64D78FFA9FF8CD8E88914FE25F811" box="[421,502,1899,1924]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="alterus">
|
||
<emphasis id="53C2EAE9FFA9FF8CD8E88914FE25F811" box="[421,502,1899,1924]" italics="true" pageId="42" pageNumber="43">alterus</emphasis>
|
||
</taxonomicName>
|
||
in previous works (e.g.
|
||
<bibRefCitation id="05274B0AFFA9FF8CDA6D8915FBFBF811" author="Capllonch, P. & Autino, A. & Diaz, M. M. & Barquez, R. M. & Goytia, M." box="[800,1064,1898,1924]" journalOrPublisher="Mastozoologia Neotropical" pageId="42" pageNumber="43" pagination="49 - 71" part="4" refString="Capllonch, P., Autino, A., Diaz, M. M., Barquez, R. M. & Goytia, M. (1997) Los mamiferos del Parque Biologico Sierra de San Javier, Tucuman, Argentina: observaciones sobre su sistematica y distribucion. Mastozoologia Neotropical, 4, 49 - 71." title="Los mamiferos del Parque Biologico Sierra de San Javier, Tucuman, Argentina: observaciones sobre su sistematica y distribucion" type="journal article" year="1997">
|
||
Capllonch
|
||
<emphasis id="53C2EAE9FFA9FF8CDAEE8914FC0AF811" box="[931,985,1899,1924]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. 1997
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="05274B0AFFA9FF8CDD7B8915FB2EF811" author="Capllonch, P. & Autino, A. & Diaz, M. M. & Barquez, R. M. & Goytia, M." box="[1078,1277,1898,1924]" journalOrPublisher="Mastozoologia Neotropical" pageId="42" pageNumber="43" pagination="49 - 71" part="4" refString="Capllonch, P., Autino, A., Diaz, M. M., Barquez, R. M. & Goytia, M. (1997) Los mamiferos del Parque Biologico Sierra de San Javier, Tucuman, Argentina: observaciones sobre su sistematica y distribucion. Mastozoologia Neotropical, 4, 49 - 71." title="Los mamiferos del Parque Biologico Sierra de San Javier, Tucuman, Argentina: observaciones sobre su sistematica y distribucion" type="journal article" year="1997">
|
||
Díaz
|
||
<emphasis id="53C2EAE9FFA9FF8CDD358914FB7DF811" box="[1144,1198,1899,1924]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. 1997
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="05274B0AFFA9FF8CDC418915FF06F83F" author="Diaz, M. M. & Braun, J. K. & Mares, M. A. & Barquez, R. M." journalOrPublisher="Occasional Papers of the Oklahoma Museum of Natural History" pageId="42" pageNumber="43" pagination="1 - 10" part="2" refString="Diaz, M. M., Braun, J. K., Mares, M. A. & Barquez, R. M. (1997) Key of Mammals of Salta Province, Argentina. Occasional Papers of the Oklahoma Museum of Natural History, 2, 1 - 10." title="Key of Mammals of Salta Province, Argentina" type="journal article" year="1997">
|
||
Mares
|
||
<emphasis id="53C2EAE9FFA9FF8CDC128914FA46F811" box="[1375,1429,1899,1924]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. 1997
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="05274B0AFFA9FF8CD9AF89EFFE8CF83F" author="Diaz, M. M." bookContentInfo="640 pp." box="[226,351,1936,1962]" journalOrPublisher="Thesis, Universidad Nacional de Tucuman, Argentina" pageId="42" pageNumber="43" refString="Diaz, M. M. (1999) Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil. Thesis, Universidad Nacional de Tucuman, Argentina, 640 pp." title="Mamiferos de la Provincia de Jujuy: Sistematica, distribucion y ecologia. Unpublished D. Phil" type="book" year="1999">Díaz 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="05274B0AFFA9FF8CD82189EFFDA3F83F" author="Diaz, M. M. & Barquez, R. M." box="[364,624,1936,1962]" editor="Kelt, D. A." journalOrPublisher="University of California Publications in Zoology 134" pageId="42" pageNumber="43" pagination="417 - 578" refString="Diaz, M. M. & Barquez, R. M. (2007) The Wild Mammals of Jujuy Province, Argentina: Systematics and Distribution. In: Kelt, D. A., Lessa, E. P., Salazar-Bravo, J. & Patton, J. L. (Eds.), The Quintessential Naturalist: Honoring the Life and Legacy of Oliver P. Pearson. University of California Publications in Zoology 134, pp. 417 - 578." title="The Wild Mammals of Jujuy Province, Argentina: Systematics and Distribution" type="book chapter" volumeTitle="The Quintessential Naturalist: Honoring the Life and Legacy of Oliver P. Pearson" year="2007">Díaz & Barquez 2007</bibRefCitation>
|
||
) none of these references was based on a detailed study of extensive topotypical series nor the use of molecular characters.
|
||
<bibRefCitation id="05274B0AFFA9FF8CDA5C89CAFBC6F85A" author="Blaustein, S. A. & Liascovich, R. C. & Apfelbaum, L. I. & Daleffe, L. Barquez, R. M. & Reig, O. A." box="[785,1045,1973,1999]" journalOrPublisher="Zeitschrift fur Saugetierkunde" pageId="42" pageNumber="43" pagination="1 - 13" part="57" refString="Blaustein, S. A., Liascovich, R. C., Apfelbaum, L. I., Daleffe, L. Barquez, R. M. & Reig, O. A. (1992) Correlates of systematic differentiation between two closely related allopatric populations of the Akodon boliviensis group from NW Argentina (Rodentia, Cricetidae). Zeitschrift fur Saugetierkunde, 57, 1 - 13." title="Correlates of systematic differentiation between two closely related allopatric populations of the Akodon boliviensis group from NW Argentina (Rodentia, Cricetidae)" type="journal article" year="1992">
|
||
Blaustein
|
||
<emphasis id="53C2EAE9FFA9FF8CDACB89CAFC69F85B" box="[902,954,1973,1998]" italics="true" pageId="42" pageNumber="43">et al</emphasis>
|
||
. (1992)
|
||
</bibRefCitation>
|
||
were the only authors who tested the status of
|
||
<taxonomicName id="A6B64D78FFA9FF8CD86189A4FEADF861" box="[300,382,2011,2036]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="alterus">
|
||
<emphasis id="53C2EAE9FFA9FF8CD86189A4FEADF861" box="[300,382,2011,2036]" italics="true" pageId="42" pageNumber="43">alterus</emphasis>
|
||
</taxonomicName>
|
||
(with respect to
|
||
<taxonomicName id="A6B64D78FFA9FF8CDB0989A4FD37F861" box="[580,740,2011,2036]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="tucumanensis">
|
||
<emphasis id="53C2EAE9FFA9FF8CDB0989A4FD37F861" box="[580,740,2011,2036]" italics="true" pageId="42" pageNumber="43">tucumanensis</emphasis>
|
||
</taxonomicName>
|
||
). However, they erroneously assumed that specimens from El Infiernillo,
|
||
<collectingRegion id="A372F819FFA8FF8DD80C8EE7FE61FF27" box="[321,434,152,178]" country="Argentina" name="Tucuman" pageId="43" pageNumber="44">Tucumán</collectingRegion>
|
||
, belonged to
|
||
<taxonomicName id="A6B64D78FFA8FF8DDB1E8EE7FD77FF24" box="[595,676,152,177]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="alterus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDB1E8EE7FD77FF24" box="[595,676,152,177]" italics="true" pageId="43" pageNumber="44">alterus</emphasis>
|
||
</taxonomicName>
|
||
; then, they did not truly test the taxonomic status of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDC6C8EE7FA47FF24" box="[1313,1428,152,178]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="alterus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDC6C8EE7FAE0FF24" box="[1313,1331,152,177]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDC0E8EE7FA47FF24" box="[1347,1428,152,177]" italics="true" pageId="43" pageNumber="44">alterus</emphasis>
|
||
</taxonomicName>
|
||
. They also failed to separate these specimens from
|
||
<taxonomicName id="A6B64D78FFA8FF8DDA5A8EC2FC0EFF43" box="[791,989,189,215]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="tucumanensis">
|
||
<emphasis id="53C2EAE9FFA8FF8DDA5A8EC2FCFAFF43" box="[791,809,189,214]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDA708EC2FC0EFF43" box="[829,989,189,214]" italics="true" pageId="43" pageNumber="44">tucumanensis</emphasis>
|
||
</taxonomicName>
|
||
from low altitude Yungas forest of
|
||
<collectingRegion id="A372F819FFA8FF8DD9DA8E9DFEDAFF69" box="[151,265,226,252]" country="Argentina" name="Tucuman" pageId="43" pageNumber="44">Tucumán</collectingRegion>
|
||
. In words of
|
||
<bibRefCitation id="05274B0AFFA8FF8DD8E48E9DFD7DFF69" author="Blaustein, S. A. & Liascovich, R. C. & Apfelbaum, L. I. & Daleffe, L. Barquez, R. M. & Reig, O. A." box="[425,686,226,252]" journalOrPublisher="Zeitschrift fur Saugetierkunde" pageId="43" pageNumber="44" pagination="1 - 13" part="57" refString="Blaustein, S. A., Liascovich, R. C., Apfelbaum, L. I., Daleffe, L. Barquez, R. M. & Reig, O. A. (1992) Correlates of systematic differentiation between two closely related allopatric populations of the Akodon boliviensis group from NW Argentina (Rodentia, Cricetidae). Zeitschrift fur Saugetierkunde, 57, 1 - 13." title="Correlates of systematic differentiation between two closely related allopatric populations of the Akodon boliviensis group from NW Argentina (Rodentia, Cricetidae)" type="journal article" year="1992">
|
||
Blaustein
|
||
<emphasis id="53C2EAE9FFA8FF8DDB6C8E9DFD86FF6E" box="[545,597,226,251]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
|
||
. (1992
|
||
</bibRefCitation>
|
||
, pag. 11) “... we believe that any conclusion on the taxonomic status of the two forms is untimely.” The status of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDBAE8F77FC7EFEB4" box="[739,941,264,290]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="leucolimnaeus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDBAE8F77FD26FEB4" box="[739,757,264,289]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDA4E8F77FC7EFEB4" box="[771,941,264,289]" italics="true" pageId="43" pageNumber="44">leucolimnaeus</emphasis>
|
||
</taxonomicName>
|
||
was even more dubious and, until the preliminary observations of
|
||
<bibRefCitation id="05274B0AFFA8FF8DD8F08F52FD62FED2" author="Galliari, C. A. & Pardinas, U. F. J. & Goin, F. J." box="[445,689,301,327]" journalOrPublisher="Mastozoologia Neotropical" pageId="43" pageNumber="44" pagination="39 - 61" part="3" refString="Galliari, C. A., Pardinas, U. F. J. & Goin, F. J. (1996) Lista comentada de los mamiferos argentinos. Mastozoologia Neotropical, 3, 39 - 61." title="Lista comentada de los mamiferos argentinos" type="journal article" year="1996">
|
||
Galliari
|
||
<emphasis id="53C2EAE9FFA8FF8DDB538F52FD81FED3" box="[542,594,301,326]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
|
||
. (1996)
|
||
</bibRefCitation>
|
||
, it was considered as a synonym of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDD158F52FAF4FED3" box="[1112,1319,301,326]" class="Mammalia" family="Muridae" genus="Necromys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="lactens">
|
||
<emphasis id="53C2EAE9FFA8FF8DDD158F52FAF4FED3" box="[1112,1319,301,326]" italics="true" pageId="43" pageNumber="44">Necromys lactens</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="05274B0AFFA8FF8DDC7B8F52FF06FEF9" author="Cabrera, A." journalOrPublisher="Revista del Museo Argentino de Ciencias Naturales " Bernardino Rivadavia ", Zoologia" pageId="43" pageNumber="44" pagination="309 - 732" part="4" refString="Cabrera, A. (1961) Catalogo de los mamiferos de America del Sur. Parte II. Revista del Museo Argentino de Ciencias Naturales " Bernardino Rivadavia ", Zoologia, 4 (2), 309 - 732." title="Catalogo de los mamiferos de America del Sur. Parte II" type="journal article" year="1961">Cabrera, 1961</bibRefCitation>
|
||
; Reig, 1978, 1987;
|
||
<bibRefCitation id="05274B0AFFA8FF8DD8F08F2DFD4BFEF9" author="Diaz, M. M. & Braun, J. K. & Mares, M. A. & Barquez, R. M." box="[445,664,338,364]" journalOrPublisher="Occasional Papers of the Oklahoma Museum of Natural History" pageId="43" pageNumber="44" pagination="1 - 10" part="2" refString="Diaz, M. M., Braun, J. K., Mares, M. A. & Barquez, R. M. (1997) Key of Mammals of Salta Province, Argentina. Occasional Papers of the Oklahoma Museum of Natural History, 2, 1 - 10." title="Key of Mammals of Salta Province, Argentina" type="journal article" year="1997">
|
||
Mares
|
||
<emphasis id="53C2EAE9FFA8FF8DDB428F2DFD90FEFE" box="[527,579,338,363]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
|
||
., 1997
|
||
</bibRefCitation>
|
||
). However,
|
||
<bibRefCitation id="05274B0AFFA8FF8DDA6A8F2DFBC9FEF9" author="Galliari, C. A. & Pardinas, U. F. J. & Goin, F. J." box="[807,1050,338,364]" journalOrPublisher="Mastozoologia Neotropical" pageId="43" pageNumber="44" pagination="39 - 61" part="3" refString="Galliari, C. A., Pardinas, U. F. J. & Goin, F. J. (1996) Lista comentada de los mamiferos argentinos. Mastozoologia Neotropical, 3, 39 - 61." title="Lista comentada de los mamiferos argentinos" type="journal article" year="1996">
|
||
Galliari
|
||
<emphasis id="53C2EAE9FFA8FF8DDAC48F2DFC6EFEFE" box="[905,957,338,363]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
|
||
. (1996)
|
||
</bibRefCitation>
|
||
ranked
|
||
<taxonomicName id="A6B64D78FFA8FF8DDD378F2DFAF0FEFE" box="[1146,1315,338,363]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="leucolimnaeus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDD378F2DFAF0FEFE" box="[1146,1315,338,363]" italics="true" pageId="43" pageNumber="44">leucolimnaeus</emphasis>
|
||
</taxonomicName>
|
||
as a valid species of
|
||
<taxonomicName id="A6B64D78FFA8FF8DD85B8F07FEA3FE04" box="[278,368,376,401]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="53C2EAE9FFA8FF8DD85B8F07FEA3FE04" box="[278,368,376,401]" italics="true" pageId="43" pageNumber="44">Akodon</emphasis>
|
||
</taxonomicName>
|
||
(see also
|
||
<bibRefCitation id="05274B0AFFA8FF8DD8A78F07FCF5FE07" author="Musser, G. M. & Carleton, M. D." box="[490,806,376,402]" editor="Wilson, D. E." journalOrPublisher="Baltimore: John Hopkins University Press" pageId="43" pageNumber="44" pagination="894 - 1531" refString="Musser, G. M. & Carleton, M. D. (2005) Superfamily Muroidea .. In: Wilson, D. E. & Reeder, D. M. (Eds.) Mammal species of the world: A taxonomic and geographic reference. Third ed. Baltimore: John Hopkins University Press, pp. 894 - 1531." title="Superfamily Muroidea .." type="book chapter" volumeTitle="Mammal species of the world: A taxonomic and geographic reference. Third ed" year="2005">Musser & Carleton [2005]</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="05274B0AFFA8FF8DDA2E8F07FBB4FE07" author="Jayat, J. P. & Ortiz, P. E. & Teta, P. & Pardinas, U. F. J. & D'Elia, G." box="[867,1127,376,402]" journalOrPublisher="Mastozoologia Neotropical" pageId="43" pageNumber="44" pagination="51 - 67" part="13" refString="Jayat, J. P., Ortiz, P. E., Teta, P., Pardinas, U. F. J. & D'Elia, G. (2006) Nuevas localidades argentinas para algunos roedores sigmodontinos (Rodentia: Cricetidae). Mastozoologia Neotropical, 13, 51 - 67." title="Nuevas localidades argentinas para algunos roedores sigmodontinos (Rodentia: Cricetidae)" type="journal article" year="2006">
|
||
Pardiñas
|
||
<emphasis id="53C2EAE9FFA8FF8DDA9C8F07FBD5FE04" box="[977,1030,376,401]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
|
||
. [2006]
|
||
</bibRefCitation>
|
||
), view that contrasts with our results. All the available evidence indicates that the nominal forms
|
||
<taxonomicName id="A6B64D78FFA8FF8DDAAF8FE2FBE0FE23" box="[994,1075,413,438]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="alterus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDAAF8FE2FBE0FE23" box="[994,1075,413,438]" italics="true" pageId="43" pageNumber="44">alterus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="A6B64D78FFA8FF8DDD0F8FE2FB38FE23" box="[1090,1259,413,438]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="leucolimnaeus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDD0F8FE2FB38FE23" box="[1090,1259,413,438]" italics="true" pageId="43" pageNumber="44">leucolimnaeus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="A6B64D78FFA8FF8DDC608FE2FF01FE4E" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="tucumanensis">
|
||
<emphasis id="53C2EAE9FFA8FF8DDC608FE2FF01FE4E" italics="true" pageId="43" pageNumber="44">tucumanensis</emphasis>
|
||
</taxonomicName>
|
||
must be considered junior synonyms of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDB8E8FBDFCBAFE4E" box="[707,873,450,476]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="spegazzinii">
|
||
<emphasis id="53C2EAE9FFA8FF8DDB8E8FBDFD06FE4E" box="[707,725,450,475]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDBAA8FBDFCBAFE4E" box="[743,873,450,475]" italics="true" pageId="43" pageNumber="44">spegazzinii</emphasis>
|
||
</taxonomicName>
|
||
. Haplotypes from specimens coming from the
|
||
<typeStatus id="BE0D8859FFA8FF8DD9DA8F97FF19FD97" box="[151,202,488,514]" pageId="43" pageNumber="44">type</typeStatus>
|
||
localities of these fall within the
|
||
<taxonomicName id="A6B64D78FFA8FF8DDB198F97FD04FD94" box="[596,727,488,513]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="spegazzinii">
|
||
<emphasis id="53C2EAE9FFA8FF8DDB198F97FD04FD94" box="[596,727,488,513]" italics="true" pageId="43" pageNumber="44">spegazzinii</emphasis>
|
||
</taxonomicName>
|
||
clade (
|
||
<figureCitation id="F98D2A7EFFA8FF8DDA608F97FCA1FD97" box="[813,882,488,514]" captionStart="FIGURE 1" captionStartId="7.[151,255,1731,1755]" captionTargetBox="[278,1283,182,1711]" captionTargetId="figure@7.[249,1323,152,1743]" captionTargetPageId="7" captionText="FIGURE 1. Results of the phylogenetic analyses based on the first 801 bases of the cyt b gene of 71 sequences of Akodon. Topotype specimens included in the analyses are marked as follows: (a) A. alterus; (b) A. puer caenosus; (c) A. leucolimnaeus; (d) A. polopi new species (including the holotype); (e) A. spegazzinii; (f) A. sylvanus; and (g) A. tucumanensis. For those sequences retrieved from Genbank, accession numbers are provided next to species labels. For those sequences gathered by us, specimen collection numbers are provided next to species label; see Appendix I for locality data and Genbank accession numbers. A) Strict consensus tree of the 220 most parsimonious trees (length 1563, CI = 0.287, RI = 0.675). Numbers indicate parsimony jackknife values of the nodes at their right. Only jackknife values above 50 % are shown. B) Majority-rule consensus resulting from the Bayesian analysis. Numbers indicate posterior probability values of the nodes at their right." httpUri="https://zenodo.org/record/293462/files/figure.png" pageId="43" pageNumber="44">Fig. 1</figureCitation>
|
||
). The genetic divergence among haplotypes of this clade is low (1.2% in average) and we found no clear morphological or morphometric differences among these forms. Most of the observed differences in coloration among populations may relate to the great environmental heterogeneity inhabited by
|
||
<taxonomicName id="A6B64D78FFA8FF8DDB188C27FD2BFDE4" box="[597,760,600,626]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="spegazzinii">
|
||
<emphasis id="53C2EAE9FFA8FF8DDB188C27FDB4FDE4" box="[597,615,600,625]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDB388C27FD2BFDE4" box="[629,760,600,625]" italics="true" pageId="43" pageNumber="44">spegazzinii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="F98D2A7EFFA8FF8DDA4A8C27FC8CFDE7" box="[775,863,600,626]" captionStart="FIGURE 10" captionStartId="30.[151,255,1323,1347]" captionTargetBox="[159,1427,381,1302]" captionTargetId="figure@30.[159,1427,381,1302]" captionTargetPageId="30" captionText="FIGURE 10. Habitats of the type localities of the four nominal forms associated with A. spegazzinii: A) High altitudinal grasslands at 2400 m, near Otro Cerro, type locality of A. alterus Thomas. B) Puna environments in Laguna Blanca, 3200 m, type locality of A. leucolimnaeus Cabrera. C) Monte environments near Cachi, 2500 m, type locality of A. spegazzinii Thomas. D) Rain forest at Horco Molle, about 700 m, near the type locality of A. tucumanensis J. A. Allen." httpUri="https://zenodo.org/record/293860/files/figure.png" pageId="43" pageNumber="44">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="610936FBFFA8FF8DD98B8C02FF03FC09" blockId="43.[151,1437,152,2007]" pageId="43" pageNumber="44">
|
||
In the
|
||
<taxonomicName id="A6B64D78FFA8FF8DD8548C02FE68FD03" box="[281,443,637,662]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="boliviensis">
|
||
<emphasis id="53C2EAE9FFA8FF8DD8548C02FE68FD03" box="[281,443,637,662]" italics="true" pageId="43" pageNumber="44">A. boliviensis</emphasis>
|
||
</taxonomicName>
|
||
clade, two haplotypes recovered from specimens from northern
|
||
<collectingRegion id="A372F819FFA8FF8DDD988C02FAC3FD02" box="[1237,1296,637,663]" country="Argentina" name="Salta" pageId="43" pageNumber="44">Salta</collectingRegion>
|
||
were more closely related to the haplotype of a Peruvian specimen (MVZ 171607) than to those of other three Argentinean specimens (
|
||
<figureCitation id="F98D2A7EFFA8FF8DD8188CB7FE4EFD77" box="[341,413,712,738]" captionStart="FIGURE 1" captionStartId="7.[151,255,1731,1755]" captionTargetBox="[278,1283,182,1711]" captionTargetId="figure@7.[249,1323,152,1743]" captionTargetPageId="7" captionText="FIGURE 1. Results of the phylogenetic analyses based on the first 801 bases of the cyt b gene of 71 sequences of Akodon. Topotype specimens included in the analyses are marked as follows: (a) A. alterus; (b) A. puer caenosus; (c) A. leucolimnaeus; (d) A. polopi new species (including the holotype); (e) A. spegazzinii; (f) A. sylvanus; and (g) A. tucumanensis. For those sequences retrieved from Genbank, accession numbers are provided next to species labels. For those sequences gathered by us, specimen collection numbers are provided next to species label; see Appendix I for locality data and Genbank accession numbers. A) Strict consensus tree of the 220 most parsimonious trees (length 1563, CI = 0.287, RI = 0.675). Numbers indicate parsimony jackknife values of the nodes at their right. Only jackknife values above 50 % are shown. B) Majority-rule consensus resulting from the Bayesian analysis. Numbers indicate posterior probability values of the nodes at their right." httpUri="https://zenodo.org/record/293462/files/figure.png" pageId="43" pageNumber="44">Fig. 1</figureCitation>
|
||
). This phylogeographic break, together with some morphological, morphometric and genetic differences between these two groups also suggest the possiblity of unrecognized biological diversity within this group. This scenario is similar to that observed in
|
||
<taxonomicName id="A6B64D78FFA8FF8DDA2E8D6DFC3EFCBE" box="[867,1005,786,811]" class="Mammalia" family="Muridae" genus="Necromys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="amoenus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDA2E8D6DFC3EFCBE" box="[867,1005,786,811]" italics="true" pageId="43" pageNumber="44">N. amoenus</emphasis>
|
||
</taxonomicName>
|
||
, another highland inhabitant (D’Elía et al., 2008). Notwithstanding, the studied material of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDA5E8D47FC63FCC4" box="[787,944,824,849]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="boliviensis">
|
||
<emphasis id="53C2EAE9FFA8FF8DDA5E8D47FC63FCC4" box="[787,944,824,849]" italics="true" pageId="43" pageNumber="44">A. boliviensis</emphasis>
|
||
</taxonomicName>
|
||
is scarce and the difference between both clades is moderate (2.1% of average genetic divergence), so we point that further research is needed on this area.
|
||
</paragraph>
|
||
<paragraph id="610936FBFFA8FF8DD98B8DD7FEE3FA32" blockId="43.[151,1437,152,2007]" pageId="43" pageNumber="44">
|
||
<bibRefCitation id="05274B0AFFA8FF8DD98B8DD7FE7DFC57" author="Myers, P. & Patton, J. L. & Smith, M. F." box="[198,430,936,962]" journalOrPublisher="Miscellaneous Publications of the Museum of Zoology, University of Michigan" pageId="43" pageNumber="44" pagination="1 - 89" part="177" refString="Myers, P., Patton, J. L. & Smith, M. F. (1990) A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 177, 1 - 89." title="A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia" type="journal article" year="1990">
|
||
Myers
|
||
<emphasis id="53C2EAE9FFA8FF8DD8578DD7FE84FC54" box="[282,343,936,961]" italics="true" pageId="43" pageNumber="44">et al.</emphasis>
|
||
(1990)
|
||
</bibRefCitation>
|
||
considered
|
||
<taxonomicName id="A6B64D78FFA8FF8DDB0D8DD7FD1EFC54" box="[576,717,936,962]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="caenosus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDB0D8DD7FD81FC54" box="[576,594,936,961]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDB2F8DD7FD1EFC54" box="[610,717,936,961]" italics="true" pageId="43" pageNumber="44">caenosus</emphasis>
|
||
</taxonomicName>
|
||
as a subspecies of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDAFA8DD7FB96FC54" box="[951,1093,936,962]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="lutescens">
|
||
<emphasis id="53C2EAE9FFA8FF8DDAFA8DD7FC1AFC54" box="[951,969,936,961]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDA978DD7FB96FC54" box="[986,1093,936,961]" italics="true" pageId="43" pageNumber="44">lutescens</emphasis>
|
||
</taxonomicName>
|
||
(as
|
||
<taxonomicName id="A6B64D78FFA8FF8DDD378DD7FA95FC54" box="[1146,1350,936,962]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="subSpecies" species="puer" subSpecies="caenosus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDD378DD7FB5FFC54" box="[1146,1164,936,961]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDDD18DD7FA95FC54" box="[1180,1350,936,961]" italics="true" pageId="43" pageNumber="44">puer caenosus</emphasis>
|
||
</taxonomicName>
|
||
in that publication). They highlighted subtle morphometric, morphologic, and cytogenetic characteristics that distinguished specimens from northwestern
|
||
<collectingCountry id="19A1766BFFA8FF8DDB2F8D8DFD04FB99" box="[610,727,1010,1036]" name="Argentina" pageId="43" pageNumber="44">Argentina</collectingCountry>
|
||
from those of
|
||
<collectingCountry id="19A1766BFFA8FF8DDAC48D8DFC6CFB99" box="[905,959,1010,1036]" name="Peru" pageId="43" pageNumber="44">Peru</collectingCountry>
|
||
and
|
||
<collectingCountry id="19A1766BFFA8FF8DDAB68D8DFB86FB99" box="[1019,1109,1010,1036]" name="Bolivia" pageId="43" pageNumber="44">Bolivia</collectingCountry>
|
||
. Our phylogenetic analyses include haplotypes recovered from Argentinean specimens assignable to
|
||
<taxonomicName id="A6B64D78FFA8FF8DDAB08A67FBBAFBA4" box="[1021,1129,1048,1073]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="caenosus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDAB08A67FBBAFBA4" box="[1021,1129,1048,1073]" italics="true" pageId="43" pageNumber="44">caenosus</emphasis>
|
||
</taxonomicName>
|
||
and a Peruvian specimen belonging to
|
||
<taxonomicName id="A6B64D78FFA8FF8DD8748A42FE77FBC3" box="[313,420,1085,1110]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="lutescens">
|
||
<emphasis id="53C2EAE9FFA8FF8DD8748A42FE77FBC3" box="[313,420,1085,1110]" italics="true" pageId="43" pageNumber="44">lutescens</emphasis>
|
||
</taxonomicName>
|
||
. These haplotypes form a paraphyletic group respect to
|
||
<taxonomicName id="A6B64D78FFA8FF8DDD148A42FB3CFBC3" box="[1113,1263,1085,1111]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="subfuscus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDD148A42FBB8FBC3" box="[1113,1131,1085,1110]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDD308A42FB3CFBC3" box="[1149,1263,1085,1110]" italics="true" pageId="43" pageNumber="44">subfuscus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="F98D2A7EFFA8FF8DDC4E8A42FA98FBC2" box="[1283,1355,1085,1111]" captionStart="FIGURE 1" captionStartId="7.[151,255,1731,1755]" captionTargetBox="[278,1283,182,1711]" captionTargetId="figure@7.[249,1323,152,1743]" captionTargetPageId="7" captionText="FIGURE 1. Results of the phylogenetic analyses based on the first 801 bases of the cyt b gene of 71 sequences of Akodon. Topotype specimens included in the analyses are marked as follows: (a) A. alterus; (b) A. puer caenosus; (c) A. leucolimnaeus; (d) A. polopi new species (including the holotype); (e) A. spegazzinii; (f) A. sylvanus; and (g) A. tucumanensis. For those sequences retrieved from Genbank, accession numbers are provided next to species labels. For those sequences gathered by us, specimen collection numbers are provided next to species label; see Appendix I for locality data and Genbank accession numbers. A) Strict consensus tree of the 220 most parsimonious trees (length 1563, CI = 0.287, RI = 0.675). Numbers indicate parsimony jackknife values of the nodes at their right. Only jackknife values above 50 % are shown. B) Majority-rule consensus resulting from the Bayesian analysis. Numbers indicate posterior probability values of the nodes at their right." httpUri="https://zenodo.org/record/293462/files/figure.png" pageId="43" pageNumber="44">Fig. 1</figureCitation>
|
||
). This topology, if
|
||
<taxonomicName id="A6B64D78FFA8FF8DD8668A1CFE45FBE9" box="[299,406,1123,1148]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="lutescens">
|
||
<emphasis id="53C2EAE9FFA8FF8DD8668A1CFE45FBE9" box="[299,406,1123,1148]" italics="true" pageId="43" pageNumber="44">lutescens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A6B64D78FFA8FF8DD8998A1CFD95FBE9" box="[468,582,1123,1148]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="subfuscus">
|
||
<emphasis id="53C2EAE9FFA8FF8DD8998A1CFD95FBE9" box="[468,582,1123,1148]" italics="true" pageId="43" pageNumber="44">subfuscus</emphasis>
|
||
</taxonomicName>
|
||
are not to be considered conspecifics, suggests a specific status for
|
||
<taxonomicName id="A6B64D78FFA8FF8DDCCE8A1CFED1FB34" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="caenosus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDCCE8A1CFA46FBE9" box="[1411,1429,1123,1148]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DD9DA8AF7FED1FB34" box="[151,258,1160,1185]" italics="true" pageId="43" pageNumber="44">caenosus</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, the average genetic distance between the Peruvian haplotype of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDD9B8AF7FA91FB34" box="[1238,1346,1160,1185]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="lutescens">
|
||
<emphasis id="53C2EAE9FFA8FF8DDD9B8AF7FA91FB34" box="[1238,1346,1160,1185]" italics="true" pageId="43" pageNumber="44">lutescens</emphasis>
|
||
</taxonomicName>
|
||
(MVZ 171612) and those from northwestern Argentinean specimens of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDAEA8AD2FBE5FB53" box="[935,1078,1197,1222]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="caenosus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDAEA8AD2FBE5FB53" box="[935,1078,1197,1222]" italics="true" pageId="43" pageNumber="44">A. caenosus</emphasis>
|
||
</taxonomicName>
|
||
is relatively high (3.5%); the average divergence value among specimens of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDB8D8AACFC9FFB79" box="[704,844,1234,1260]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="caenosus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDB8D8AACFD01FB79" box="[704,722,1235,1260]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDBAC8AACFC9FFB79" box="[737,844,1235,1260]" italics="true" pageId="43" pageNumber="44">caenosus</emphasis>
|
||
</taxonomicName>
|
||
is much lower (1%). Although, there are not clear morphological and morphometric differences among our material of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDA9F8A87FBB3FA84" box="[978,1120,1272,1297]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="caenosus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDA9F8A87FBB3FA84" box="[978,1120,1272,1297]" italics="true" pageId="43" pageNumber="44">A. caenosus</emphasis>
|
||
</taxonomicName>
|
||
and the values offered by
|
||
<bibRefCitation id="05274B0AFFA8FF8DD9DA8B62FEAAFAA2" author="Myers, P. & Patton, J. L. & Smith, M. F." box="[151,377,1309,1335]" journalOrPublisher="Miscellaneous Publications of the Museum of Zoology, University of Michigan" pageId="43" pageNumber="44" pagination="1 - 89" part="177" refString="Myers, P., Patton, J. L. & Smith, M. F. (1990) A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 177, 1 - 89." title="A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia" type="journal article" year="1990">
|
||
Myers
|
||
<emphasis id="53C2EAE9FFA8FF8DD9A48B62FECFFAA3" box="[233,284,1309,1334]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
|
||
. (1990)
|
||
</bibRefCitation>
|
||
for
|
||
<taxonomicName id="A6B64D78FFA8FF8DD8E78B62FDE5FAA3" box="[426,566,1309,1335]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="lutescens">
|
||
<emphasis id="53C2EAE9FFA8FF8DD8E78B62FE6FFAA3" box="[426,444,1309,1334]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DD8868B62FDE5FAA3" box="[459,566,1309,1334]" italics="true" pageId="43" pageNumber="44">lutescens</emphasis>
|
||
</taxonomicName>
|
||
, the alternative position of considering
|
||
<taxonomicName id="A6B64D78FFA8FF8DDD448B62FB48FAA3" box="[1033,1179,1309,1335]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="subfuscus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDD448B62FBC8FAA3" box="[1033,1051,1309,1334]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDD648B62FB48FAA3" box="[1065,1179,1309,1334]" italics="true" pageId="43" pageNumber="44">subfuscus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A6B64D78FFA8FF8DDD998B62FAB3FAA3" box="[1236,1376,1309,1335]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="lutescens">
|
||
<emphasis id="53C2EAE9FFA8FF8DDD998B62FB35FAA3" box="[1236,1254,1309,1334]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDDB88B62FAB3FAA3" box="[1269,1376,1309,1334]" italics="true" pageId="43" pageNumber="44">lutescens</emphasis>
|
||
</taxonomicName>
|
||
conspecific seems less supported. These species differ by a substantially high (5.1%) average genetic distance. Furthermore
|
||
<bibRefCitation id="05274B0AFFA8FF8DD87F8B17FDC5FA17" author="Myers, P. & Patton, J. L. & Smith, M. F." box="[306,534,1384,1410]" journalOrPublisher="Miscellaneous Publications of the Museum of Zoology, University of Michigan" pageId="43" pageNumber="44" pagination="1 - 89" part="177" refString="Myers, P., Patton, J. L. & Smith, M. F. (1990) A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia. Miscellaneous Publications of the Museum of Zoology, University of Michigan, 177, 1 - 89." title="A review of the boliviensis group of Akodon (Muridae: Sigmodontinae) with emphasis on Peru and Bolivia" type="journal article" year="1990">
|
||
Myers
|
||
<emphasis id="53C2EAE9FFA8FF8DD8C98B17FE6AFA14" box="[388,441,1384,1409]" italics="true" pageId="43" pageNumber="44">et al</emphasis>
|
||
. (1990)
|
||
</bibRefCitation>
|
||
detailed several morphological characters that distinguish
|
||
<taxonomicName id="A6B64D78FFA8FF8DDD838B17FA88FA14" box="[1230,1371,1384,1410]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="lutescens">
|
||
<emphasis id="53C2EAE9FFA8FF8DDD838B17FB33FA14" box="[1230,1248,1384,1409]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDDA28B17FA88FA14" box="[1263,1371,1384,1409]" italics="true" pageId="43" pageNumber="44">lutescens</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="A6B64D78FFA8FF8DD9DA8BF2FEFAFA33" box="[151,297,1421,1447]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="subfuscus">
|
||
<emphasis id="53C2EAE9FFA8FF8DD9DA8BF2FF7AFA33" box="[151,169,1421,1446]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DD9FA8BF2FEFAFA33" box="[183,297,1421,1446]" italics="true" pageId="43" pageNumber="44">subfuscus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="610936FBFFA8FF8DD98B8BCDFEB5F889" blockId="43.[151,1437,152,2007]" pageId="43" pageNumber="44">
|
||
Although we failed to obtain small specimens of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDA518BCCFCA5FA59" box="[796,886,1459,1484]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDA518BCCFCA5FA59" box="[796,886,1459,1484]" italics="true" pageId="43" pageNumber="44">Akodon</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<typeStatus id="BE0D8859FFA8FF8DDAA28BCDFBF1FA59" box="[1007,1058,1458,1484]" pageId="43" pageNumber="44">type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDDFD8BCCFAB4FA59" box="[1200,1383,1458,1484]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="aliquantulus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDDFD8BCCFB11FA59" box="[1200,1218,1459,1484]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDD998BCCFAB4FA59" box="[1236,1383,1459,1484]" italics="true" pageId="43" pageNumber="44">aliquantulus</emphasis>
|
||
</taxonomicName>
|
||
(we studied ca. 100 individuals of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDB5B8BA7FDA3FA64" box="[534,624,1496,1521]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDB5B8BA7FDA3FA64" box="[534,624,1496,1521]" italics="true" pageId="43" pageNumber="44">Akodon</emphasis>
|
||
</taxonomicName>
|
||
from this locality and its surroundings and more than 390 from
|
||
<collectingRegion id="A372F819FFA8FF8DD9DA8B82FEA4F982" box="[151,375,1533,1559]" country="Argentina" name="Tucuman" pageId="43" pageNumber="44">Tucumán province</collectingRegion>
|
||
), its diagnosis does not permit us to distinguish the two
|
||
<typeStatus id="BE0D8859FFA8FF8DDD418B82FBECF982" box="[1036,1087,1533,1559]" pageId="43" pageNumber="44">type</typeStatus>
|
||
specimens from
|
||
<taxonomicName id="A6B64D78FFA8FF8DDC458B82FA47F983" box="[1288,1428,1533,1559]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="caenosus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDC458B82FAC9F983" box="[1288,1306,1533,1558]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDC648B82FA47F983" box="[1321,1428,1533,1558]" italics="true" pageId="43" pageNumber="44">caenosus</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, the haplotypes of three small
|
||
<taxonomicName id="A6B64D78FFA8FF8DDB31885CFD05F9A9" box="[636,726,1571,1596]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="genus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDB31885CFD05F9A9" box="[636,726,1571,1596]" italics="true" pageId="43" pageNumber="44">Akodon</emphasis>
|
||
</taxonomicName>
|
||
sequenced from
|
||
<collectingRegion id="A372F819FFA8FF8DDAE9885DFBF1F9A9" box="[932,1058,1570,1596]" country="Argentina" name="Catamarca" pageId="43" pageNumber="44">Catamarca</collectingRegion>
|
||
and
|
||
<collectingRegion id="A372F819FFA8FF8DDD13885DFB1FF9A9" box="[1118,1228,1570,1596]" country="Argentina" name="Tucuman" pageId="43" pageNumber="44">Tucumán</collectingRegion>
|
||
were remarkably similar to haplotypes of
|
||
<taxonomicName id="A6B64D78FFA8FF8DD8F98837FDECF9F4" box="[436,575,1608,1634]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="caenosus">
|
||
<emphasis id="53C2EAE9FFA8FF8DD8F98837FE15F9F4" box="[436,454,1608,1633]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DD8998837FDECF9F4" box="[468,575,1608,1633]" italics="true" pageId="43" pageNumber="44">caenosus</emphasis>
|
||
</taxonomicName>
|
||
topotypes (with average genetic distance lower than 1%). On these bases (see also our Species Account discussion above) we place
|
||
<taxonomicName id="A6B64D78FFA8FF8DDA098812FC04F913" box="[836,983,1645,1670]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="aliquantulus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDA098812FC04F913" box="[836,983,1645,1670]" italics="true" pageId="43" pageNumber="44">aliquantulus</emphasis>
|
||
</taxonomicName>
|
||
under the synonymy of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDDB88812FA53F913" box="[1269,1408,1645,1671]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="caenosus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDDB88812FAD4F913" box="[1269,1287,1645,1670]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDC588812FA53F913" box="[1301,1408,1645,1670]" italics="true" pageId="43" pageNumber="44">caenosus</emphasis>
|
||
</taxonomicName>
|
||
. This study corroborated the proposition of
|
||
<bibRefCitation id="05274B0AFFA8FF8DDBDD88EDFCA2F939" author="Jayat, J. P. & Ortiz, P. E. & Pardinas, U. F. J. & D'Elia, G." box="[656,881,1682,1708]" journalOrPublisher="Mastozoologia Neotropical" pageId="43" pageNumber="44" pagination="201 - 225" part="14" refString="Jayat, J. P., Ortiz, P. E., Pardinas, U. F. J. & D'Elia, G. (2007 a) Redescripcion y posicion filogenetica del raton selvatico (Akodon sylvanus: Rodentia: Cricetidae). Mastozoologia Neotropical, 14, 201 - 225." title="Redescripcion y posicion filogenetica del raton selvatico (Akodon sylvanus: Rodentia: Cricetidae)" type="journal article" year="2007">
|
||
Jayat
|
||
<emphasis id="53C2EAE9FFA8FF8DDB9988ECFCDDF939" box="[724,782,1683,1708]" italics="true" pageId="43" pageNumber="44">et al.</emphasis>
|
||
(2007a)
|
||
</bibRefCitation>
|
||
of placing
|
||
<taxonomicName id="A6B64D78FFA8FF8DDABA88ECFBAEF939" box="[1015,1149,1682,1708]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="sylvanus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDABA88ECFBDAF939" box="[1015,1033,1683,1708]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDD5588ECFBAEF939" box="[1048,1149,1683,1708]" italics="true" pageId="43" pageNumber="44">sylvanus</emphasis>
|
||
</taxonomicName>
|
||
as a valid species of the
|
||
<taxonomicName id="A6B64D78FFA8FF8DD9DA88C7FEE7F944" box="[151,308,1720,1745]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="boliviensis">
|
||
<emphasis id="53C2EAE9FFA8FF8DD9DA88C7FEE7F944" box="[151,308,1720,1745]" italics="true" pageId="43" pageNumber="44">A. boliviensis</emphasis>
|
||
</taxonomicName>
|
||
group. In the MP analysis
|
||
<taxonomicName id="A6B64D78FFA8FF8DDB2388C7FD20F944" box="[622,755,1720,1745]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="sylvanus">
|
||
<emphasis id="53C2EAE9FFA8FF8DDB2388C7FD20F944" box="[622,755,1720,1745]" italics="true" pageId="43" pageNumber="44">A. sylvanus</emphasis>
|
||
</taxonomicName>
|
||
appears as the sister species of
|
||
<taxonomicName id="A6B64D78FFA8FF8DDD2E88C7FB1DF944" box="[1123,1230,1720,1746]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="polopi">
|
||
<emphasis id="53C2EAE9FFA8FF8DDD2E88C7FBA6F944" box="[1123,1141,1720,1745]" italics="true" pageId="43" pageNumber="44">A</emphasis>
|
||
.
|
||
<emphasis id="53C2EAE9FFA8FF8DDDCE88C7FB1DF944" box="[1155,1230,1720,1745]" italics="true" pageId="43" pageNumber="44">polopi</emphasis>
|
||
</taxonomicName>
|
||
; in accordance to low support of this clade (JK<50) it is not recovered in the BA where
|
||
<taxonomicName id="A6B64D78FFA8FF8DDA8E88A2FBFDF963" box="[963,1070,1757,1782]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="polopi">
|
||
<emphasis id="53C2EAE9FFA8FF8DDA8E88A2FBFDF963" box="[963,1070,1757,1782]" italics="true" pageId="43" pageNumber="44">A. polopi</emphasis>
|
||
</taxonomicName>
|
||
is sister to the
|
||
<emphasis id="53C2EAE9FFA8FF8DDD9188A2FEC9F889" italics="true" pageId="43" pageNumber="44">
|
||
<taxonomicName id="A6B64D78FFA8FF8DDD9188A2FAAAF963" box="[1244,1401,1757,1782]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="boliviensis">A. boliviensis</taxonomicName>
|
||
-
|
||
<taxonomicName id="A6B64D78FFA8FF8DDCCE88A2FEC9F889" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="spegazzinii">A. spegazzinii</taxonomicName>
|
||
</emphasis>
|
||
clade.
|
||
</paragraph>
|
||
<paragraph id="610936FBFFA8FF8AD98B8957FCFBFEB7" blockId="43.[151,1437,152,2007]" lastBlockId="44.[151,1436,152,551]" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
|
||
<taxonomicName id="A6B64D78FFA8FF8DD98B8957FEA0F8D4" box="[198,371,1832,1857]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="polopi">
|
||
<emphasis id="53C2EAE9FFA8FF8DD98B8957FEA0F8D4" box="[198,371,1832,1857]" italics="true" pageId="43" pageNumber="44">Akodon polopi</emphasis>
|
||
</taxonomicName>
|
||
, the new species here described, is a clearly distinct member of the
|
||
<taxonomicName id="A6B64D78FFA8FF8DDDEA8957FA9BF8D4" box="[1191,1352,1832,1857]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="boliviensis">
|
||
<emphasis id="53C2EAE9FFA8FF8DDDEA8957FA9BF8D4" box="[1191,1352,1832,1857]" italics="true" pageId="43" pageNumber="44">A. boliviensis</emphasis>
|
||
</taxonomicName>
|
||
group, with several diagnostic characters coming from different sources of evidence; not surprinsingly, it was early mentioned as a possible new subspecies by Elio Massoia or a new species (see
|
||
<bibRefCitation id="05274B0AFFA8FF8DDD1A890DFB3FF819" author="Polop, J." box="[1111,1260,1906,1932]" journalOrPublisher="Studies on Neotropical Fauna and Environment" pageId="43" pageNumber="44" pagination="53 - 59" part="24" refString="Polop, J. (1989) Distribution and ecological observations of wild rodents in Pampa de Achala, Cordoba, Argentina. Studies on Neotropical Fauna and Environment, 24, 53 - 59." title="Distribution and ecological observations of wild rodents in Pampa de Achala, Cordoba, Argentina" type="journal article" year="1989">Polop, 1989</bibRefCitation>
|
||
). This species, known only from high altitude grasslands of central
|
||
<collectingCountry id="19A1766BFFA8FF8DDA4C89E7FCAAF827" box="[769,889,1944,1970]" name="Argentina" pageId="43" pageNumber="44">Argentina</collectingCountry>
|
||
, is the only species of the
|
||
<taxonomicName id="A6B64D78FFA8FF8DDDFC89E7FA9CF824" box="[1201,1359,1944,1969]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="43" pageNumber="44" phylum="Chordata" rank="species" species="boliviensis">
|
||
<emphasis id="53C2EAE9FFA8FF8DDDFC89E7FA9CF824" box="[1201,1359,1944,1969]" italics="true" pageId="43" pageNumber="44">A. boliviensis</emphasis>
|
||
</taxonomicName>
|
||
group inhabiting the Sierras Grandes range, a medium altitude mountain system isolated (ca.,
|
||
<quantity id="A64E9B1EFFA8FF8DDDC289C2FB3AF842" box="[1167,1257,1981,2007]" metricMagnitude="5" metricUnit="m" metricValue="6.0" pageId="43" pageNumber="44" unit="km" value="600.0">600 km</quantity>
|
||
) from the main Andean chain by low elevation arid and semiarid environments. Data at hand are insufficient to pose a biogeographic scenario accounting for the diversification of
|
||
<taxonomicName id="A6B64D78FFAFFF8ADA468EC2FCA5FF43" box="[779,886,189,214]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="polopi">
|
||
<emphasis id="53C2EAE9FFAFFF8ADA468EC2FCA5FF43" box="[779,886,189,214]" italics="true" pageId="44" pageNumber="45">A. polopi</emphasis>
|
||
</taxonomicName>
|
||
and the remainder species of the
|
||
<taxonomicName id="A6B64D78FFAFFF8ADDB38EC2FA4FFF43" box="[1278,1436,189,214]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="boliviensis">
|
||
<emphasis id="53C2EAE9FFAFFF8ADDB38EC2FA4FFF43" box="[1278,1436,189,214]" italics="true" pageId="44" pageNumber="45">A. boliviensis</emphasis>
|
||
</taxonomicName>
|
||
group. Notwithstanding, Pampa de Achala has been mentioned as a biogeographic refuge and an area of endemism for other taxa (
|
||
<bibRefCitation id="05274B0AFFAFFF8AD8C08F77FCCFFEB7" author="Polop, J." box="[397,796,264,290]" journalOrPublisher="Studies on Neotropical Fauna and Environment" pageId="44" pageNumber="45" pagination="53 - 59" part="24" refString="Polop, J. (1989) Distribution and ecological observations of wild rodents in Pampa de Achala, Cordoba, Argentina. Studies on Neotropical Fauna and Environment, 24, 53 - 59." title="Distribution and ecological observations of wild rodents in Pampa de Achala, Cordoba, Argentina" type="journal article" year="1989">Polop, 1989, and references there</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="610936FBFFAFFF8AD98B8F52FBE4FEF9" blockId="44.[151,1436,152,551]" pageId="44" pageNumber="45">
|
||
Relationships among species of the
|
||
<taxonomicName id="A6B64D78FFAFFF8ADB338F52FCB2FED3" box="[638,865,301,326]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="boliviensis">
|
||
<emphasis id="53C2EAE9FFAFFF8ADB338F52FCB2FED3" box="[638,865,301,326]" italics="true" pageId="44" pageNumber="45">Akodon boliviensis</emphasis>
|
||
</taxonomicName>
|
||
group are well resolved and mostly coungrent among analyses. Differences relate to the position of
|
||
<taxonomicName id="A6B64D78FFAFFF8ADA4B8F2DFCA1FEFE" box="[774,882,338,363]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="polopi">
|
||
<emphasis id="53C2EAE9FFAFFF8ADA4B8F2DFCA1FEFE" box="[774,882,338,363]" italics="true" pageId="44" pageNumber="45">A. polopi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="A6B64D78FFAFFF8ADAE68F2DFBE3FEFE" box="[939,1072,338,363]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="sylvanus">
|
||
<emphasis id="53C2EAE9FFAFFF8ADAE68F2DFBE3FEFE" box="[939,1072,338,363]" italics="true" pageId="44" pageNumber="45">A. sylvanus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="610936FBFFAFFF8AD98B8F07FE8CFDB2" blockId="44.[151,1436,152,551]" pageId="44" pageNumber="45">
|
||
The present study, as several previous ones, reaffirms the phylogenetic legitimacy of both the
|
||
<taxonomicName id="A6B64D78FFAFFF8ADC538F07FA4FFE04" box="[1310,1436,376,401]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="44" pageNumber="45" phylum="Chordata" rank="species" species="boliviensis">
|
||
<emphasis id="53C2EAE9FFAFFF8ADC538F07FA4FFE04" box="[1310,1436,376,401]" italics="true" pageId="44" pageNumber="45">boliviensis</emphasis>
|
||
</taxonomicName>
|
||
group and extends our knowledge of its contents, limits, and geographic ranges of its member species. However, we assert that additional research is still needed before an adequate picture of Andean
|
||
<taxonomicName id="A6B64D78FFAFFF8ADDA68FBDFA96FE4E" box="[1259,1349,450,475]" class="Mammalia" family="Muridae" genus="Akodon" kingdom="Animalia" order="Rodentia" pageId="44" pageNumber="45" phylum="Chordata" rank="genus">
|
||
<emphasis id="53C2EAE9FFAFFF8ADDA68FBDFA96FE4E" box="[1259,1349,450,475]" italics="true" pageId="44" pageNumber="45">Akodon</emphasis>
|
||
</taxonomicName>
|
||
can be acquired. In this line, more field work coupled with the study of specimen series housed in research institutions is essential.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |